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<document id="CAF89409E64DFB12E5F870D009AFE367" ID-CLB-Dataset="21836" ID-DOI="10.5852/ejt.2012.17" ID-GBIF-Dataset="ac80d266-8f8e-4fc5-a172-599a331449a4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3857844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590517236165" checkinUser="carolina" docAuthor="Karner, Michael" docDate="2012" docId="03BB8D13E113FF9DFD8E7DEE815FFBAC" docLanguage="en" docName="ejt-2012-17.pdf.imf" docOrigin="European Journal of Taxonomy 17" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Psammoecus hacquardi Grouvelle 1889" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="FF82F56BE117FF9BFFA87F6B8405FFDA" masterDocTitle="A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de l’Afrique centrale" masterLastPageNumber="31" masterPageNumber="1" pageNumber="5" updateTime="1698841482259" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D8683016183F79B293C4157932DACDC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="75971EE40287CBBB06822D862ADB9142">
<mods:title id="E4FF400FA0DC3BCF1DCDF6CD12445B62">A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de l’Afrique centrale</mods:title>
</mods:titleInfo>
<mods:name id="A0B7544CF5A6B929C0B642DD24F8252F" type="personal">
<mods:role id="C9D06F3483779FA0E8A53BD4F25956DF">
<mods:roleTerm id="96840F721F0482E1DCC1CB1EB67F4D35">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="617172CB8DF1633BDAC7CDA3F465FA2C">Karner, Michael</mods:namePart>
</mods:name>
<mods:typeOfResource id="3B7529D4D8166A835D7E5360B654D6B4">text</mods:typeOfResource>
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<mods:titleInfo id="C281F55D6905CA04A19FD8044ADEE1DB">
<mods:title id="A859F94541B3859C8527ABD99F6036F5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="1708CC338C17F22F61CD0D1049251D9D">
<mods:date id="B333D24B98E0D2884522C99589BDD416">2012</mods:date>
<mods:detail id="D16F6722DCE8EA6CB0B4B36BE9123536" type="pubDate">
<mods:number id="D67C5522B96E5150154A81831CA1958C">2012-06-04</mods:number>
</mods:detail>
<mods:detail id="CB71EBB1D080EAC6620EFC023BBBD824" type="issue">
<mods:number id="C89B7FFE8E8FC5606D08BE1C5D4876BB">17</mods:number>
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<mods:start id="F0AA9BEA0C756F42ECD61451E166D035">1</mods:start>
<mods:end id="EBA58952695AD33BA30563C837D1850B">31</mods:end>
</mods:extent>
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<mods:classification id="D9E3D855015551B53481D1C57B86916B">journal article</mods:classification>
<mods:identifier id="62A22AAB6FE84CA383D627A04B1CB2AA" type="CLB-Dataset">21836</mods:identifier>
<mods:identifier id="A84821C56CBC207E19E35471FEFD7C58" type="DOI">10.5852/ejt.2012.17</mods:identifier>
<mods:identifier id="115A77D5D9BF0CAFD8388E8E70060C8C" type="GBIF-Dataset">ac80d266-8f8e-4fc5-a172-599a331449a4</mods:identifier>
<mods:identifier id="437E44A69A3CD2094519CCA52871EC35" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="DC840A7006EDB26873CB74D6BACD25B5" type="Zenodo-Dep">3857844</mods:identifier>
<mods:identifier id="2CD9AD447C6D0CC3771FCA73F7F8F033" type="ZooBank">1238951A-FE17-40B2-9DCD-9D7939BECD09</mods:identifier>
</mods:mods>
<treatment id="03BB8D13E113FF9DFD8E7DEE815FFBAC" ID-DOI="http://doi.org/10.5281/zenodo.3858722" ID-GBIF-Taxon="164302229" ID-Zenodo-Dep="3858722" LSID="urn:lsid:plazi:treatment:03BB8D13E113FF9DFD8E7DEE815FFBAC" httpUri="http://treatment.plazi.org/id/03BB8D13E113FF9DFD8E7DEE815FFBAC" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
<subSubSection id="C3086F8EE113FF9FFD8E7DEE800BFD45" box="[550,1038,644,671]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BAD3C05E113FF9FFD8E7DEE800BFD45" blockId="4.[550,1038,644,707]" box="[550,1038,644,671]" pageId="4" pageNumber="5">
<heading id="D0E58B69E113FF9FFD8E7DEE800BFD45" box="[550,1038,644,671]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C124786E113FF9FFD8E7DEE800BFD45" authority="Grouvelle, 1889" authorityName="Grouvelle" authorityYear="1889" box="[550,1038,644,671]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hacquardi">
<emphasis id="B966E017E113FF9FFD8E7DEE8742FD45" bold="true" box="[550,839,644,671]" italics="true" pageId="4" pageNumber="5">Psammoecus hacquardi</emphasis>
Grouvelle, 1889
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE113FF9FFD467DC38743FD19" box="[750,838,680,707]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BAD3C05E113FF9FFD467DC38743FD19" blockId="4.[550,1038,644,707]" box="[750,838,680,707]" pageId="4" pageNumber="5">
(
<figureCitation id="13292080E113FF9FFD5E7DC38738FD19" box="[758,829,680,707]" captionStart="Fig" captionStartId="5.[189,232,1965,1991]" captionTargetBox="[267,1327,760,1846]" captionTargetId="figure@5.[189,1398,705,1925]" captionTargetPageId="5" captionText="Fig. 2. Psammoecus hacquardi Grouvelle, 1889. A. Habitus of lectotype. B. Left antenna of lectotype. C. Parameres of specimen from Tanzanie: Uluguru Mountains. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857850" httpUri="https://zenodo.org/record/3857850/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE113FF9FFF157D80872AFCDF" box="[189,815,746,773]" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8BAD3C05E113FF9FFF157D80872AFCDF" blockId="4.[189,815,746,773]" box="[189,815,746,773]" pageId="4" pageNumber="5">
<treatmentCitationGroup id="AB021B2BE113FF9FFF157D80872AFCDF" box="[189,815,746,773]" pageId="4" pageNumber="5">
<treatmentCitation id="0AB31A14E113FF9FFF157D80872EFCDF" box="[189,811,746,773]" pageId="4" pageNumber="5">
<taxonomicName id="4C124786E113FF9FFF157D8085D0FCDE" box="[189,469,746,772]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="haequardi">
<emphasis id="B966E017E113FF9FFF157D8085D0FCDE" box="[189,469,746,772]" italics="true" pageId="4" pageNumber="5">Psammoecus haequardi</emphasis>
</taxonomicName>
[sic.]
<treatmentCitation id="0AB31A14E113FF9FFDB77D81872EFCDF" author="Grouvelle A." box="[543,811,746,773]" page="185" pageId="4" pageNumber="5" year="1908">
<bibRefCitation id="EF8341F4E113FF9FFDB77D81872EFCDF" author="Grouvelle A." box="[543,811,746,773]" pageId="4" pageNumber="5" pagination="127 - 206" refId="ref15107" refString="Grouvelle A. 1908 b. Coleopteres clavicornes de l'Afrique Australe et Orientale. Annales de la Societe Entomologique de France 77: 127 - 206." type="journal article" year="1908">Grouvelle, 1908b: 185</bibRefCitation>
</treatmentCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE113FF9FFF157C478025F8D4" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BAD3C05E113FF9FFF157C4785AFFC9C" blockId="4.[189,426,812,838]" box="[189,426,812,838]" pageId="4" pageNumber="5">
<heading id="D0E58B69E113FF9FFF157C4785AFFC9C" bold="true" box="[189,426,812,838]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B966E017E113FF9FFF157C4785AFFC9C" bold="true" box="[189,426,812,838]" pageId="4" pageNumber="5">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF427C378563FCAC" blockId="4.[189,1399,860,994]" box="[234,358,860,886]" pageId="4" pageNumber="5">
<emphasis id="B966E017E113FF9FFF427C378563FCAC" bold="true" box="[234,358,860,886]" pageId="4" pageNumber="5">
<typeStatus id="54A982A7E113FF9FFF427C378563FCAC" box="[234,358,860,886]" pageId="4" pageNumber="5" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF157CEB8579FC3B" blockId="4.[189,1399,860,994]" pageId="4" pageNumber="5">
<specimenCount id="9D14F78CE113FF9FFF157CEB84D6FC40" box="[189,211,896,922]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
, by present designation: ‘
<collectingCountry id="F3057C95E113FF9FFDBE7C148681FC43" box="[534,644,895,921]" name="Tanzania" pageId="4" pageNumber="5">Zanzibar</collectingCountry>
| Raffray’ [blue label, Grouvelle’s hand], ‘Type’ [red label], ‘MUSEUM PARIS | 1917 | Coll. GROUVELLE’ [blue label], ‘Hacquardi |
<collectorName id="26E759D3E113FF9FFBEB7CCF80A9FC67" box="[1091,1196,931,958]" pageId="4" pageNumber="5">A. Grou’</collectorName>
[not Grouvelle’s hand] (
<collectionCode id="ED03A4C0E113FF9FFEA77CAC8575FC3B" box="[271,368,967,993]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF427B628592FBF9" blockId="4.[189,1002,1033,1096]" box="[234,407,1033,1059]" pageId="4" pageNumber="5">
<emphasis id="B966E017E113FF9FFF427B628592FBF9" bold="true" box="[234,407,1033,1059]" pageId="4" pageNumber="5">
<typeStatus id="54A982A7E113FF9FFF427B628592FBF9" box="[234,407,1033,1059]" pageId="4" pageNumber="5" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF157B4587EFFB9D" blockId="4.[189,1002,1033,1096]" box="[189,1002,1069,1096]" pageId="4" pageNumber="5">
<specimenCount id="9D14F78CE113FF9FFF157B4584D6FB92" box="[189,211,1070,1096]" pageId="4" pageNumber="5" type="female">♀</specimenCount>
, by present designation, with identical data as
<typeStatus id="54A982A7E113FF9FFD507B458768FB92" box="[760,877,1070,1096]" pageId="4" pageNumber="5" type="lectotype">Lectotype</typeStatus>
(
<collectionCode id="ED03A4C0E113FF9FFCD57B4687DBFB9D" box="[893,990,1069,1095]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF427B0485A3FB53" blockId="4.[189,1399,1135,1807]" box="[234,422,1135,1161]" pageId="4" pageNumber="5">
<emphasis id="B966E017E113FF9FFF427B0485A3FB53" bold="true" box="[234,422,1135,1161]" pageId="4" pageNumber="5">Other material</emphasis>
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF157BF88025F8D4" blockId="4.[189,1399,1135,1807]" pageId="4" pageNumber="5">
2 spms ‘
<collectingCountry id="F3057C95E113FF9FFEB47BF8858AFB77" box="[284,399,1171,1197]" name="Tanzania" pageId="4" pageNumber="5">Tanzanie</collectingCountry>
: Mts. Uluguru | Kiroka, for. héliophile | alt.
<quantity id="4CEA91E0E113FF9FFC3F7BF887E4FB74" box="[919,993,1171,1198]" metricMagnitude="2" metricUnit="m" metricValue="7.25" pageId="4" pageNumber="5" unit="m" value="725.0">725 m</quantity>
sous écorces | 27-31/V/71’, ‘Coll. Mus. Tervuren | Mission Mts. Uluguru | L. Berger, N. Leleup | J. Debecker V/VIII/71’ (
<collectionCode id="ED03A4C0E113FF9FFB1D7BDC8117FB0B" box="[1205,1298,1207,1233]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 3 spms ‘
<collectingCountry id="F3057C95E113FF9FFF697BB08531FB2F" box="[193,308,1243,1269]" name="Tanzania" pageId="4" pageNumber="5">Tanzanie</collectingCountry>
: Mts. Uluguru | Kiroka, for. héliophile | alt.
<quantity id="4CEA91E0E113FF9FFCEA7BB1878BFB2F" box="[834,910,1242,1269]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="m" value="600.0">600 m</quantity>
24-30/VII/71 | dans arbres morts’, ‘Coll. Mus. Tervuren | Mission Mts. Uluguru | L. Berger, N. Leleup | J. Debecker V/VIII/71’ (
<collectionCode id="ED03A4C0E113FF9FFB6F7B948127FAC3" box="[1223,1314,1279,1305]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
, 1 spm
<collectionCode id="ED03A4C0E113FF9FFF157A488507FAE7" box="[189,258,1315,1341]" pageId="4" pageNumber="5">MKF</collectionCode>
). 1 spm ‘
<collectingCountry id="F3057C95E113FF9FFEC27A4985D8FAE6" box="[362,477,1314,1340]" name="Tanzania" pageId="4" pageNumber="5">Tanzanie</collectingCountry>
: Mts. Uluguru | Kiroka, for. héliophile | alt.
<quantity id="4CEA91E0E113FF9FFC407A498036FAE7" box="[1000,1075,1314,1341]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="m" value="600.0">600 m</quantity>
24-30/VII/71’, ‘Coll. Mus. Tervuren | Mission Mts. Uluguru | L. Berger, N. Leleup | J. Debecker V/VIII/71’ (
<collectionCode id="ED03A4C0E113FF9FFBDC7A2D80D4FABA" box="[1140,1233,1350,1376]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 2 spms ‘A la lumière’ [blue label], ‘I.R.S.A.C.-MUS.
<collectingCountry id="F3057C95E113FF9FFD287A0186F2FA5E" box="[640,759,1386,1412]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
|
<collectingRegion id="49D6F2E7E113FF9FFCA37A01874FFA5E" box="[779,842,1386,1412]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="4" pageNumber="5">Kivu</collectingRegion>
: Terr Masisi,
<specimenCount id="9D14F78CE113FF9FFC417A01803FFA5E" box="[1001,1082,1386,1412]" pageId="4" pageNumber="5" type="generic">
<quantity id="4CEA91E0E113FF9FFC417A018033FA5E" box="[1001,1078,1386,1412]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="5" unit="m" value="720.0">720 m</quantity>
.
</specimenCount>
| Walikale (Rives Loa) IX- 53 | N. Leleup’ (
<collectionCode id="ED03A4C0E113FF9FFED67AE585DCFA72" box="[382,473,1422,1448]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
,
<collectionCode id="ED03A4C0E113FF9FFE4E7AE5862EFA72" box="[486,555,1422,1448]" pageId="4" pageNumber="5">MKF</collectionCode>
). 3 spms ‘Dans | humus | en forêt’ [blue rectangular label], ‘I.R.S.A.C.- MUS.
<collectingCountry id="F3057C95E113FF9FFEA87AD98569FA16" box="[256,364,1458,1484]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
|
<collectingRegion id="49D6F2E7E113FF9FFE2A7ADA85C4FA11" box="[386,449,1457,1483]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="4" pageNumber="5">Kivu</collectingRegion>
: Terr Masisi,
<specimenCount id="9D14F78CE113FF9FFDCB7AD986B3FA16" box="[611,694,1458,1484]" pageId="4" pageNumber="5" type="generic">
<quantity id="4CEA91E0E113FF9FFDCB7AD986B7FA16" box="[611,690,1458,1484]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="5" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
| Mutakato IX-53 | N. Leleup’, (
<collectionCode id="ED03A4C0E113FF9FFB917AD98093FA16" box="[1081,1174,1458,1484]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
).
<specimenCount id="9D14F78CE113FF9FFB027AD980CBFA16" box="[1194,1230,1458,1484]" pageId="4" pageNumber="5" type="female">1♀</specimenCount>
‘MUSÉE DU
<collectingCountry id="F3057C95E113FF9FFF157ABD8520FA2A" box="[189,293,1494,1520]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
| Eala |
<date id="FFAC1AC5E113FF9FFED67ABD85E7FA2A" box="[382,482,1494,1520]" pageId="4" pageNumber="5" value="1934-11">XI-1934</date>
| J. Ghesquiére’ (
<collectionCode id="ED03A4C0E113FF9FFD017ABD8703FA2A" box="[681,774,1494,1520]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 3 spms ‘MUSÉE DU
<collectingCountry id="F3057C95E113FF9FFBBC7ABD8079FA2A" box="[1044,1148,1494,1520]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
| Eala |
<date id="FFAC1AC5E113FF9FFB7E7ABE816FFA35" box="[1238,1386,1493,1519]" pageId="4" pageNumber="5" value="1936-07-10">10-VII-1936</date>
| J. Ghesquiére | 2873’ (
<collectionCode id="ED03A4C0E113FF9FFE657A92862FF9C9" box="[461,554,1529,1555]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
).
<specimenCount id="9D14F78CE113FF9FFDE87A928661F9C9" box="[576,612,1529,1555]" pageId="4" pageNumber="5" type="male">1♂</specimenCount>
‘MUSÉE DU
<collectingCountry id="F3057C95E113FF9FFCB17A928784F9C9" box="[793,897,1529,1555]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
| Eala |
<date id="FFAC1AC5E113FF9FFC407A928050F9C9" box="[1000,1109,1529,1555]" pageId="4" pageNumber="5" value="1936-07">VII-1936</date>
| J. Ghesquiére | 2875’, (
<collectionCode id="ED03A4C0E113FF9FFF6D79768527F9ED" box="[197,290,1565,1591]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
).
<specimenCount id="9D14F78CE113FF9FFE9F7976855EF9ED" box="[311,347,1565,1591]" pageId="4" pageNumber="5" type="female">1♀</specimenCount>
‘I.R.S.A.C.-MUS.
<collectingCountry id="F3057C95E113FF9FFD82797686A4F9ED" box="[554,673,1565,1591]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">CONGO</collectingCountry>
| Bolobo (fleuve
<collectingCountry id="F3057C95E113FF9FFCD9797687BAF9ED" box="[881,959,1565,1591]" name="Democratic Republic of the Congo" pageId="4" pageNumber="5">Congo</collectingCountry>
|
<date id="FFAC1AC5E113FF9FFC7E79768042F9E2" box="[982,1095,1565,1592]" pageId="4" pageNumber="5" value="1949-10-05">5-X-1949</date>
| N. Leleup’, (
<collectionCode id="ED03A4C0E113FF9FFB5E79768156F9ED" box="[1270,1363,1565,1591]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 1 spm ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFDF0792A86F2F981" box="[600,759,1601,1627]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d‘ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFC34792A8000F981" box="[924,1029,1601,1627]" pageId="4" pageNumber="5" value="1961-11">XI. 1961</date>
| J. Decelle’, (
<collectionCode id="ED03A4C0E113FF9FFB1B792A8115F981" box="[1203,1296,1601,1627]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 4 spms ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFDB3790E86BFF9A5" box="[539,698,1637,1663]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d‘ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFCF4790E87D2F9A5" box="[860,983,1637,1663]" pageId="4" pageNumber="5" value="1962-08">VIII. 1962</date>
| J. Decelle’ (
<collectionCode id="ED03A4C0E113FF9FFBDB790E80CBF9A5" box="[1139,1230,1637,1663]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
, 1 spm
<collectionCode id="ED03A4C0E113FF9FFA8E790E816EF9A5" box="[1318,1387,1637,1663]" pageId="4" pageNumber="5">MKF</collectionCode>
). 2 spms ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFDDB79E28714F978" box="[627,785,1672,1699]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d’ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFC1A79E2801FF978" box="[946,1050,1672,1699]" pageId="4" pageNumber="5" value="1962-09">IX. 1962</date>
| J. Decelle’ (
<collectionCode id="ED03A4C0E113FF9FFB1E79E28116F979" box="[1206,1299,1673,1699]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 1 spms ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFDB379C786BCF91C" box="[539,697,1708,1734]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d’ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFCF279C687BDF91C" box="[858,952,1708,1735]" pageId="4" pageNumber="5" value="1962-10">X. 1962</date>
| J. Decelle’, (
<collectionCode id="ED03A4C0E113FF9FFBF579C780BFF91C" box="[1117,1210,1708,1734]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 1 spm ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFE1379BB865EF930" box="[443,603,1744,1770]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d’ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFCA879BA876CF930" box="[768,873,1744,1771]" pageId="4" pageNumber="5" value="1962-11">XI. 1962</date>
| J. Decelle’ (
<collectionCode id="ED03A4C0E113FF9FFBA279BB8062F930" box="[1034,1127,1744,1770]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
). 1 spm ‘COLL. MUS. TERVUREN |
<collectingCountry id="F3057C95E113FF9FFEC4799F860EF8D4" box="[364,523,1780,1806]" name="Ivory Coast" pageId="4" pageNumber="5">Côte d’ivoire</collectingCountry>
: Bingerville |
<date id="FFAC1AC5E113FF9FFD07799E871DF8D4" box="[687,792,1780,1807]" pageId="4" pageNumber="5" value="1963-09">IX. 1963</date>
| J. Decelle’ (
<collectionCode id="ED03A4C0E113FF9FFC1F799F8011F8D4" box="[951,1044,1780,1806]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE113FF9EFF15785D8647FE92" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BAD3C05E113FF9FFF15785D85CCF88A" blockId="4.[189,457,1846,1872]" box="[189,457,1846,1872]" pageId="4" pageNumber="5">
<heading id="D0E58B69E113FF9FFF15785D85CCF88A" bold="true" box="[189,457,1846,1872]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B966E017E113FF9FFF15785D85CCF88A" bold="true" box="[189,457,1846,1872]" pageId="4" pageNumber="5">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E113FF9FFF15780D8172F831" blockId="4.[189,1399,1893,2027]" pageId="4" pageNumber="5">
<taxonomicName id="4C124786E113FF9FFF15780D855AF8A5" authorityName="Grouvelle" authorityYear="1889" box="[189,351,1893,1919]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hacquardi">
<emphasis id="B966E017E113FF9FFF15780D855AF8A5" box="[189,351,1893,1919]" italics="true" pageId="4" pageNumber="5">Ps. hacquardi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C124786E113FF9FFE5C780D86BCF8A5" authorityName="Grouvelle" authorityYear="1908" box="[500,697,1893,1919]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B966E017E113FF9FFE5C780D86BCF8A5" box="[500,697,1893,1919]" italics="true" pageId="4" pageNumber="5">Ps. marginicollis</emphasis>
</taxonomicName>
by the shape of the parameres and shorter antennae; from
<taxonomicName id="4C124786E113FF9FFAFA780D853BF879" authorityName="Grouvelle" authorityYear="1919" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B966E017E113FF9FFAFA780D853BF879" italics="true" pageId="4" pageNumber="5">Ps. personatus</emphasis>
</taxonomicName>
by the 1
<superScript id="7C67914DE113FF9FFE0B78E385ABF84C" attach="left" box="[419,430,1928,1942]" fontSize="6" pageId="4" pageNumber="5">st</superScript>
antennomere being more than two times longer as 2
<superScript id="7C67914DE113FF9FFBBC78EC8020F84F" attach="none" box="[1044,1061,1927,1941]" fontSize="6" pageId="4" pageNumber="5">nd</superScript>
; from
<taxonomicName id="4C124786E113FF9FFBDD78E180F4F879" authority="Karner, 2012" authorityName="Karner" authorityYear="2012" box="[1141,1265,1929,1955]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="leleupi" status="sp. nov.">
<emphasis id="B966E017E113FF9FFBDD78E180F4F879" box="[1141,1265,1929,1955]" italics="true" pageId="4" pageNumber="5">Ps. leleupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555D6CE113FF9FFB5078E18157F87E" box="[1272,1362,1930,1956]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
by the narrower elytral striae; from
<taxonomicName id="4C124786E113FF9FFD9E78C586BBF81D" authorityName="Grouvelle" authorityYear="1914" box="[566,702,1965,1991]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="laetulus">
<emphasis id="B966E017E113FF9FFD9E78C586BBF81D" box="[566,702,1965,1991]" italics="true" pageId="4" pageNumber="5">Ps. laetulus</emphasis>
</taxonomicName>
by the maximum pronotal width being closer to the middle; from
<taxonomicName id="4C124786E113FF9FFF5578B985A5F831" authorityName="Grouvelle" authorityYear="1919" box="[253,416,2001,2027]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B966E017E113FF9FFF5578B985A5F831" box="[253,416,2001,2027]" italics="true" pageId="4" pageNumber="5">Ps. parallelus</emphasis>
</taxonomicName>
by more extensive dark maculae on elytra, darkened elytral basis and slender shape
</paragraph>
<paragraph id="8BAD3C05E112FF9EFF157E608647FE92" blockId="5.[189,1399,267,328]" pageId="5" pageNumber="6">
of parameres; from
<taxonomicName id="4C124786E112FF9EFE047E67861BFEFF" authority="Karner, 2012" authorityName="Karner" authorityYear="2012" box="[428,542,267,293]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="luchti" status="sp. nov.">
<emphasis id="B966E017E112FF9EFE047E67861BFEFF" box="[428,542,267,293]" italics="true" pageId="5" pageNumber="6">Ps. luchti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555D6CE112FF9EFD8F7E678681FEFC" box="[551,644,268,294]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
by narrow bases of lateral teeth of pronotum, narrower elytral striae and darkened elytral bases.
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE112FF9DFF157E1B866AFC3B" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph id="8BAD3C05E112FF9EFF157E1B856EFE50" blockId="5.[189,363,368,394]" box="[189,363,368,394]" pageId="5" pageNumber="6">
<heading id="D0E58B69E112FF9EFF157E1B856EFE50" bold="true" box="[189,363,368,394]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B966E017E112FF9EFF157E1B856EFE50" bold="true" box="[189,363,368,394]" pageId="5" pageNumber="6">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E112FF9EFF157EF487E6FD56" blockId="5.[189,1399,414,652]" pageId="5" pageNumber="6">
BODY. Elongate-oval, total length
<quantity id="4CEA91E0E112FF9EFDF87EF586F2FE63" box="[592,759,414,441]" metricMagnitude="-3" metricUnit="m" metricValue="2.7249999999999996" metricValueMax="3.05" metricValueMin="2.4" pageId="5" pageNumber="6" unit="mm" value="2.7249999999999996" valueMax="3.05" valueMin="2.4">2.40-3.05 mm</quantity>
(
<figureCitation id="13292080E112FF9EFCA07EF5876DFE63" box="[776,872,414,441]" captionStart="Fig" captionStartId="5.[189,232,1965,1991]" captionTargetBox="[267,1327,760,1846]" captionTargetId="figure@5.[189,1398,705,1925]" captionTargetPageId="5" captionText="Fig. 2. Psammoecus hacquardi Grouvelle, 1889. A. Habitus of lectotype. B. Left antenna of lectotype. C. Parameres of specimen from Tanzanie: Uluguru Mountains. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857850" httpUri="https://zenodo.org/record/3857850/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
). Surface yellowish to reddish brown, elytra with variable patterns of maculae; the basic pattern consists of brown or blackish brown maculae at the elytral bases, sutures, lateral margins and apices. Specimens of lighter colour show no darker coloration on anterior half of elytral suture and posterior half of lateral margins; very dark specimens have dark elytra with light brown maculae on the anterior and posterior aspects of the disk. Antennae as in
<figureCitation id="13292080E112FF9EFAB07D408177FD9F" box="[1304,1394,555,581]" captionStart="Fig" captionStartId="5.[189,232,1965,1991]" captionTargetBox="[267,1327,760,1846]" captionTargetId="figure@5.[189,1398,705,1925]" captionTargetPageId="5" captionText="Fig. 2. Psammoecus hacquardi Grouvelle, 1889. A. Habitus of lectotype. B. Left antenna of lectotype. C. Parameres of specimen from Tanzanie: Uluguru Mountains. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857850" httpUri="https://zenodo.org/record/3857850/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
, yellowish brown; 6
<superScript id="7C67914DE112FF9EFE337D2685ADFD81" attach="left" box="[411,424,589,603]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
to 10
<superScript id="7C67914DE112FF9EFE4F7D2685F1FD81" attach="left" box="[487,500,589,603]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
and apical half of 5
<superScript id="7C67914DE112FF9EFD7C7D2686E4FD81" attach="left" box="[724,737,589,603]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
antennomere brown to blackish-brown, 11
<superScript id="7C67914DE112FF9EFB677D2680D9FD81" attach="right" box="[1231,1244,589,603]" fontSize="6" pageId="5" pageNumber="6">th</superScript>
antennomere yellowish brown, sometimes a little lighter than basal antennomeres.
</paragraph>
<caption id="DF6D6C8DE112FF9EFF1578C68138F831" ID-DOI="http://doi.org/10.5281/zenodo.3857850" ID-Zenodo-Dep="3857850" httpUri="https://zenodo.org/record/3857850/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1965,1991]" targetBox="[267,1327,760,1846]" targetPageId="5">
<paragraph id="8BAD3C05E112FF9EFF1578C68138F831" blockId="5.[189,1399,1965,2027]" pageId="5" pageNumber="6">
<emphasis id="B966E017E112FF9EFF1578C6850EF81D" bold="true" box="[189,267,1965,1991]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
<taxonomicName id="4C124786E112FF9EFEBB78C586F3F812" authority="Grouvelle, 1889" authorityName="Grouvelle" authorityYear="1889" box="[275,758,1965,1992]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="hacquardi">
<emphasis id="B966E017E112FF9EFEBB78C58629F81D" box="[275,556,1965,1991]" italics="true" pageId="5" pageNumber="6">Psammoecus hacquardi</emphasis>
Grouvelle, 1889
</taxonomicName>
.
<emphasis id="B966E017E112FF9EFCAA78C68712F81D" bold="true" box="[770,791,1965,1991]" pageId="5" pageNumber="6">A</emphasis>
. Habitus of lectotype.
<emphasis id="B966E017E112FF9EFB8D78C5803CF812" bold="true" box="[1061,1081,1966,1992]" pageId="5" pageNumber="6">B</emphasis>
. Left antenna of lectotype.
<emphasis id="B966E017E112FF9EFF1578BB84D7F830" bold="true" box="[189,210,2000,2026]" pageId="5" pageNumber="6">C</emphasis>
. Parameres of specimen from Tanzanie: Uluguru Mountains. Scale line A, B: 1 mm; C: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BAD3C05E111FF9DFF157E6780D8FE26" blockId="6.[189,1399,267,509]" pageId="6" pageNumber="7">
HEAD. Broad, temples immediately narrowed behind eyes; width
<quantity id="4CEA91E0E111FF9DFC697E60806EFEFC" box="[961,1131,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="6.95" metricValueMax="7.4" metricValueMin="6.5" pageId="6" pageNumber="7" unit="mm" value="0.695" valueMax="0.74" valueMin="0.65">0.65-0.74 mm</quantity>
, length
<quantity id="4CEA91E0E111FF9DFB617E608177FEFC" box="[1225,1394,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="6" pageNumber="7" unit="mm" value="0.41" valueMax="0.44" valueMin="0.38">0.38-0.44 mm</quantity>
, 1.61-1.80 times as wide as long. Eyes moderately protuberant,
<quantity id="4CEA91E0E111FF9DFC1F7E44805AFE90" box="[951,1119,303,330]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="6" pageNumber="7" unit="mm" value="0.21" valueMax="0.23" valueMin="0.19">0.19-0.23 mm</quantity>
long, distance of inner margins
<quantity id="4CEA91E0E111FF9DFE807E3885D3FEB7" box="[296,470,339,365]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" metricValueMax="4.8" metricValueMin="4.0" pageId="6" pageNumber="7" unit="mm" value="0.44" valueMax="0.48" valueMin="0.4">0.40-0.48 mm</quantity>
. Puncturation on vertex moderate, density of punctures in some specimens irregular, varying from sparse to very dense. Pubescence composed of long, semierect setae, directed anteriorly; microsculpture distinct, reticulate. Longitudinal impressions on vertex distinct, almost attaining the middle of the eyes. Antennae comparatively short,
<quantity id="4CEA91E0E111FF9DFC767ED4808CFE03" box="[990,1161,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.43" metricValueMin="1.3" pageId="6" pageNumber="7" unit="mm" value="1.365" valueMax="1.43" valueMin="1.3">1.30-1.43 mm</quantity>
long; antennomere proportions of
<typeStatus id="54A982A7E111FF9DFEC37E8985D2FE26" box="[363,471,482,508]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
as follows: 2.8: 1.0: 1.7: 1.1: 1.5: 1.3: 1.2: 1.2:1.2: 1.2: 2.7.
</paragraph>
<paragraph id="8BAD3C05E111FF9DFF157D4E814FFD2B" blockId="6.[189,1399,548,754]" pageId="6" pageNumber="7">
PRONOTUM. Broad, some specimens with shallow impressions close to the apical margin; width
<quantity id="4CEA91E0E111FF9DFA917D4F852FFDB9" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="7.8" metricValueMin="6.5" pageId="6" pageNumber="7" unit="mm" value="0.715" valueMax="0.78" valueMin="0.65">0.65- 0.78 mm</quantity>
, length
<quantity id="4CEA91E0E111FF9DFE217D238631FDB9" box="[393,564,584,611]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="7" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50-0.60 mm</quantity>
, 1.27-1.33 times as wide as long. Anterior angles with distinct groups of small, short teeth; lateral margins with five teeth, tooth I very short and slender, tooth II longer and wider than tooth I, tooth III longer than tooth II, tooth IV a little shorter than tooth III, tooth V very small; posterior angle with a very small, almost obtuse tooth. Puncturation on disc denser than on vertex, punctures sometimes hardly separated. Pubescence as on vertex; microsculpture distinct, reticulate.
</paragraph>
<paragraph id="8BAD3C05E111FF9DFF157C71868CFCA1" blockId="6.[189,1399,793,892]" pageId="6" pageNumber="7">
ELYTRA. Elongate-oval, length
<quantity id="4CEA91E0E111FF9DFD807C7286D6FCEE" box="[552,723,793,820]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="6" pageNumber="7" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.50-1.90 mm</quantity>
, combined width
<quantity id="4CEA91E0E111FF9DFC007C718057FCEE" box="[936,1106,794,820]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.3" metricValueMin="0.98" pageId="6" pageNumber="7" unit="mm" value="1.14" valueMax="1.3" valueMin="0.98">0.98-1.30 mm</quantity>
, 1.46-1.67 times as long as combined width. Rows of punctures on disc narrower than interstices. Pubescence composed of long, semierect setae; microsculpture absent.
</paragraph>
<paragraph id="8BAD3C05E111FF9DFF157CCF866AFC3B" blockId="6.[189,1398,931,994]" pageId="6" pageNumber="7">
PARAMERES. Slender, short, with dense rows of small setae at the inner margins of their bases and two larger setae at their apices. (
<figureCitation id="13292080E111FF9DFDAC7CAC8664FC3B" box="[516,609,967,993]" captionStart="Fig" captionStartId="5.[189,232,1965,1991]" captionTargetBox="[267,1327,760,1846]" captionTargetId="figure@5.[189,1398,705,1925]" captionTargetPageId="5" captionText="Fig. 2. Psammoecus hacquardi Grouvelle, 1889. A. Habitus of lectotype. B. Left antenna of lectotype. C. Parameres of specimen from Tanzanie: Uluguru Mountains. Scale line A, B: 1 mm; C: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3857850" httpUri="https://zenodo.org/record/3857850/files/figure.png" pageId="6" pageNumber="7">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3086F8EE111FF9DFF157B62815FFBAC" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BAD3C05E111FF9DFF157B62852AFBF9" blockId="6.[189,303,1033,1059]" box="[189,303,1033,1059]" pageId="6" pageNumber="7">
<heading id="D0E58B69E111FF9DFF157B62852AFBF9" bold="true" box="[189,303,1033,1059]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="B966E017E111FF9DFF157B62852AFBF9" bold="true" box="[189,303,1033,1059]" pageId="6" pageNumber="7">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD3C05E111FF9DFF157B53815FFBAC" blockId="6.[189,1399,1080,1143]" pageId="6" pageNumber="7">
The determination of single female specimens, especially if they are lightly colored or immature, can be difficult.
<bibRefCitation id="EF8341F4E111FF9DFEE47B378620FBAD" author="Grouvelle A." box="[332,549,1116,1143]" pageId="6" pageNumber="7" pagination="127 - 206" refId="ref15107" refString="Grouvelle A. 1908 b. Coleopteres clavicornes de l'Afrique Australe et Orientale. Annales de la Societe Entomologique de France 77: 127 - 206." type="journal article" year="1908">Grouvelle (1908b)</bibRefCitation>
spells the name ‘Haequardi’, which is to be considered a misspelling.
</paragraph>
</subSubSection>
</treatment>
</document>
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