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<document id="930173824D1D6BDFE5E32729C30B6C79" ID-CLB-Dataset="22092" ID-DOI="10.5852/ejt.2020.646" ID-GBIF-Dataset="c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b" ID-Zenodo-Dep="3829682" ID-ZooBank="4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589570996260" checkinUser="valdenar" docAuthor="Moret, Pierre &amp; Murienne, Jérôme" docDate="2020" docId="03B7726A13445338FDAFFC1FFDC7FC7B" docLanguage="en" docName="ejt-646_moret_murienne.pdf.imf" docOrigin="European Journal of Taxonomy 646" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dyscolus giselae Moret 2020, sp. nov." docType="treatment" docUuid="235B5F1A-B526-4F29-BB49-0C16A08A5218" docUuidSource="ZooBank" docVersion="8" lastPageNumber="14" masterDocId="FF8E0A1213485335FFF1FFFAFF86FF83" masterDocTitle="Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes" masterLastPageNumber="55" masterPageNumber="1" pageNumber="13" updateTime="1698839917149" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="0652B178B1D0B71C3424991B93F94BEB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5B591D738D7174522E008EAEB7972C99">
<mods:title id="844D9D28F36FD5CD2158C7B3A52C0780">Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes</mods:title>
</mods:titleInfo>
<mods:name id="A4BFC4C1CE22BB0D9995EA74D73C7704" type="personal">
<mods:role id="6855D47EE7E4015D57F815EE90A81E42">
<mods:roleTerm id="9843CF4AA79F301690F3D0A5936F1FC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4F8322B9A64C321F2C255F9A23F3D74">Moret, Pierre</mods:namePart>
<mods:nameIdentifier id="40D44A5FBCFA568594A5DE12D357FF5F" type="LSID">2E194645-D255-438B-819E-A2B6C39AD699</mods:nameIdentifier>
<mods:affiliation id="37F3269B3969BE24B1651EDD507B4C10">Laboratoire TRACES (UMR 5608), CNRS, Université Toulouse 2 Jean Jaurès, Toulouse 31058, France.</mods:affiliation>
<mods:nameIdentifier id="684BA5BEF5E5A4DFFB16EE991CE33064" type="email">pierre.moret@univ-tlse2.fr&amp;pierre.moret@univ-tlse2.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="9CB933E1E3BBA6CA6938A2FCDF6931DE" type="personal">
<mods:role id="14B25CAD7603FA399DA481F47C7A6A51">
<mods:roleTerm id="31B57844094DA3B15FF15B71806CD815">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1AB85E3B4E8EE1213A312DE8F845C23D">Murienne, Jérôme</mods:namePart>
<mods:nameIdentifier id="E530386FFCEEF0DCF546894F5C2FCC7A" type="LSID">3A6964D4-A7D6-46DB-8129-B762D14BC8F9</mods:nameIdentifier>
<mods:affiliation id="7A5B45A5275699458E1C3EDC9BFF5AA1">Laboratoire Evolution et Diversité Biologique (UMR 5174), CNRS, Université Toulouse 3 Paul Sabatier, IRD, Toulouse 31062, France.</mods:affiliation>
<mods:nameIdentifier id="BEFF9A4C93CEDE1E408BCDCA33A63007" type="email">jerome.murienne@univ-tlse3.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7AC6B3DA5C0E20E66FBAE7D36A67CB1F">text</mods:typeOfResource>
<mods:relatedItem id="1BD6017BEFD781CC873B5A458525DA15" type="host">
<mods:titleInfo id="BA665B2C4AEDB7076B3AA2C6C13A494D">
<mods:title id="AF57BE6ED1D6B8B0B17A793B6287D906">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A82C8E1DD1A68F63A800A265FCCDDB83">
<mods:date id="61D0012F10824C44218F7119656EB97B">2020</mods:date>
<mods:detail id="1B0D625535B2B12D1B767640682F1EB4" type="pubDate">
<mods:number id="5EB140C6114E582B934501EF6E943D09">2020-05-15</mods:number>
</mods:detail>
<mods:detail id="FCAAB8B7F4F439864E52424CC6EB6840" type="volume">
<mods:number id="E8B2883E601147F5E84FFF0F1D31696F">646</mods:number>
</mods:detail>
<mods:extent id="3C38BBA13BA27AC939442AD405514CD2" unit="page">
<mods:start id="C8B35A59C2810622069AC58B2B751957">1</mods:start>
<mods:end id="7E90A448FC96B189FE2DD5C4DC60D53F">55</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6C4B1CE0648B356B2D8A90971D157796">journal article</mods:classification>
<mods:identifier id="4082D841E8E1486282B522667638EA69" type="CLB-Dataset">22092</mods:identifier>
<mods:identifier id="09093C96FA708D4C673CECD0ECEE0DF2" type="DOI">10.5852/ejt.2020.646</mods:identifier>
<mods:identifier id="6049B024C3BEE38431657874602CC129" type="GBIF-Dataset">c5cf0da1-13e3-4bf4-844c-9b5a9d4f6d5b</mods:identifier>
<mods:identifier id="59717AD9F5C0B2DAFAB2CB6AA6BA506D" type="Zenodo-Dep">3829682</mods:identifier>
<mods:identifier id="4348785BCFD6AD0866274F99575C4F38" type="ZooBank">4C9F63B2-DB17-4EDB-ADEE-13AC9EFB921B</mods:identifier>
</mods:mods>
<treatment id="03B7726A13445338FDAFFC1FFDC7FC7B" ID-DOI="http://doi.org/10.5281/zenodo.3848393" ID-GBIF-Taxon="164160415" ID-Zenodo-Dep="3848393" LSID="urn:lsid:zoobank.org:act:235B5F1A-B526-4F29-BB49-0C16A08A5218" httpUri="http://treatment.plazi.org/id/03B7726A13445338FDAFFC1FFDC7FC7B" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C30490F713445339FDAFFC1FFB44FBA0" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BA1C37C13445339FDAFFC1FFC50FB83" blockId="12.[369,1218,997,1095]" box="[606,982,997,1024]" pageId="12" pageNumber="13">
<heading id="D0E9741013445339FDAFFC1FFC50FB83" box="[606,982,997,1024]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C1EB8FF13445339FDAFFC1FFCF3FB83" ID-CoL="8MHVR" authority="Moret" authorityName="Moret" authorityYear="2020" box="[606,885,997,1024]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="giselae" status="sp. nov.">
<emphasis id="B96A1F6E13445339FDAFFC1FFCA3FB83" bold="true" box="[606,805,997,1024]" italics="true" pageId="12" pageNumber="13">Dyscolus giselae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513445339FC8DFC1CFC50FB83" box="[892,982,998,1024]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BA1C37C13445339FE80FBF2FB44FBA0" blockId="12.[369,1218,997,1095]" box="[369,1218,1032,1059]" pageId="12" pageNumber="13">
<uri id="FF8FCF7E13445339FE80FBF2FB44FBA0" box="[369,1218,1032,1059]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="FFB8F9A913445339FD65FBF2FB44FBA0" box="[660,1218,1032,1059]" pageId="12" pageNumber="13">235B5F1A-B526-4F29-BB49-0C16A08A5218</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713445339FD16FBD6FCCAFBC4" box="[743,844,1068,1095]" pageId="12" pageNumber="13" type="description">
<paragraph id="8BA1C37C13445339FD16FBD6FCCAFBC4" blockId="12.[369,1218,997,1095]" box="[743,844,1068,1095]" pageId="12" pageNumber="13">
<figureCitation id="1325DFF913445339FD16FBD6FCCAFBC4" box="[743,844,1068,1095]" captionStart="Figs 3–6. 3–4" captionStartId="11.[189,241,1919,1945]" captionTargetBox="[220,1347,329,1804]" captionTargetId="figure@11.[195,1393,311,1821]" captionTargetPageId="11" captionText="Figs 3–6. 3–4. Dyscolus eleonorae Moret sp. nov. 3. Habitus of the male holotype (QCAZ). 4. Aedeagus, median lobe in lateral view. 5–6. D. giselae Moret sp. nov. 5. Habitus of the male holotype (QCAZ). 6. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829686" httpUri="https://zenodo.org/record/3829686/files/figure.png" pageId="12" pageNumber="13">Figs 5–6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30490F713445339FF4CFB97FC1AFB35" box="[189,924,1133,1206]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="8BA1C37C13445339FF4CFB97FEC2FB04" blockId="12.[189,324,1133,1159]" box="[189,324,1133,1159]" pageId="12" pageNumber="13">
<heading id="D0E9741013445339FF4CFB97FEC2FB04" bold="true" box="[189,324,1133,1159]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96A1F6E13445339FF4CFB97FEC2FB04" bold="true" box="[189,324,1133,1159]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13445339FF4CFB66FC1AFB35" blockId="12.[189,924,1180,1206]" box="[189,924,1180,1206]" pageId="12" pageNumber="13">Noun in the genitive case, dedicated to Gisèle Moret-Deysson.</paragraph>
</subSubSection>
<subSubSection id="C30490F713445338FF4CFB27FDC7FC7B" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BA1C37C13445339FF4CFB27FEEAFB74" blockId="12.[189,364,1245,1271]" box="[189,364,1245,1271]" pageId="12" pageNumber="13">
<heading id="D0E9741013445339FF4CFB27FEEAFB74" bold="true" box="[189,364,1245,1271]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96A1F6E13445339FF4CFB27FEEAFB74" bold="true" box="[189,364,1245,1271]" pageId="12" pageNumber="13">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13445339FF1BFAF6FEDAFAA5" blockId="12.[189,1399,1292,1388]" box="[234,348,1292,1318]" pageId="12" pageNumber="13">
<emphasis id="B96A1F6E13445339FF1BFAF6FEDAFAA5" bold="true" box="[234,348,1292,1318]" pageId="12" pageNumber="13">
<typeStatus id="54A57DDE13445339FF1BFAF6FEDAFAA5" box="[234,348,1292,1318]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA1C37C13445339FF4CFAD5FEC9FAEF" blockId="12.[189,1399,1292,1388]" pageId="12" pageNumber="13">
<materialsCitation id="3B76C92113445339FF4CFAD5FEC9FAEF" ID-GBIF-Occurrence="2620303351" collectingDate="2001-07-03" collectingDateMax="2001-07-05" collectingDateMin="2001-07-03" collectionCode="QCAZ" collectorName="P. Moret" country="Ecuador" elevation="1950" location="Otonga" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Cotopaxi Province" typeStatus="holotype">
<collectingCountry id="F30983EC13445339FF4CFAD5FECDFACA" box="[189,331,1327,1353]" name="Ecuador" pageId="12" pageNumber="13">ECUADOR</collectingCountry>
•
<specimenCount id="9D1808F513445339FE94FAD7FEFAFACA" box="[357,380,1325,1353]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
;
<collectingRegion id="49DA0D9E13445339FE7AFAD5FDECFACA" box="[395,618,1327,1353]" country="Ecuador" name="Cotopaxi" pageId="12" pageNumber="13">Cotopaxi Province</collectingRegion>
,
<location id="8EC195A713445339FD86FAD5FD56FACA" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FD86FAD5FD56FACA" box="[631,720,1327,1353]" country="Ecuador" name="Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi Province">Otonga</location>
, berge de ruisseau;
<elevation id="0033244F13445339FC48FAD5FBD2FACA" box="[953,1108,1327,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="12" pageNumber="13" unit="m" value="1950.0">
<quantity id="4CE66E9913445339FC48FAD5FB95FACA" box="[953,1043,1327,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="12" pageNumber="13" unit="m" value="1950.0">1950 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FB93FAD5FA83FACA" box="[1122,1285,1327,1354]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">
<collectingDate id="EFE41C5413445339FB93FAD5FA83FACA" box="[1122,1285,1327,1354]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">3–5 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FAE3FAD5FAF1FACA" box="[1298,1399,1327,1353]" pageId="12" pageNumber="13">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FF09FAA8FECAFAEF" box="[248,332,1362,1388]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="12" pageNumber="13" type="Museum">QCAZ</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA1C37C13445339FF4CFA6EFE71FA2D" blockId="12.[189,503,1426,1454]" box="[189,503,1426,1454]" pageId="12" pageNumber="13">
<emphasis id="B96A1F6E13445339FF4CFA6EFEBCFA2D" bold="true" box="[189,314,1428,1454]" pageId="12" pageNumber="13">
<typeStatus id="54A57DDE13445339FF4CFA6EFEBCFA2D" box="[189,314,1428,1454]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
</emphasis>
(9 ♂♂,
<specimenCount id="9D1808F513445339FE6DFA6EFE6DFA2D" box="[412,491,1428,1454]" pageId="12" pageNumber="13" type="female">13 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BA1C37C13445339FF4CFA2FFE5CF88A" blockId="12.[189,1399,1491,1801]" pageId="12" pageNumber="13">
<materialsCitation id="3B76C92113445339FF4CFA2FFA96FA6C" ID-GBIF-Occurrence="2620303309" box="[189,1296,1491,1520]" collectingDate="2001-07-03" collectingDateMax="2001-07-05" collectingDateMin="2001-07-03" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="1950" location="Cotopaxi Province" pageId="12" pageNumber="13" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Cotopaxi Province">
<collectingCountry id="F30983EC13445339FF4CFA2FFECDFA6C" box="[189,331,1493,1519]" name="Ecuador" pageId="12" pageNumber="13">ECUADOR</collectingCountry>
–
<emphasis id="B96A1F6E13445339FE9BFA2FFDD5FA6C" bold="true" box="[362,595,1493,1519]" pageId="12" pageNumber="13">
<collectingRegion id="49DA0D9E13445339FE9BFA2FFDD5FA6C" box="[362,595,1493,1519]" country="Ecuador" name="Cotopaxi" pageId="12" pageNumber="13">Cotopaxi Province</collectingRegion>
</emphasis>
•
<specimenCount id="9D1808F513445339FD9FFA2FFD34FA6C" box="[622,690,1491,1519]" pageId="12" pageNumber="13" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1808F513445339FD30FA2FFC83FA73" box="[705,773,1493,1520]" pageId="12" pageNumber="13" type="female">6 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED0F5BB913445339FB23FA2FFA96FA6C" box="[1234,1296,1493,1519]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="12" pageNumber="13">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FADAFA2FFBBEF991" ID-GBIF-Occurrence="2620303368" collectingDate="2001-07-03" collectingDateMax="2001-07-05" collectingDateMin="2001-07-03" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="1800" location="rio Las Damas" municipality="Otonga" pageId="12" pageNumber="13" specimenCount="3" specimenCount-male="3" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FADAFA2FFAE9FA6C" box="[1323,1391,1491,1519]" pageId="12" pageNumber="13" type="male">3 ♂♂</specimenCount>
;
<collectingMunicipality id="6BC5590613445339FF4CFA02FE90F991" box="[189,278,1528,1554]" pageId="12" pageNumber="13">Otonga</collectingMunicipality>
,
<location id="8EC195A713445339FED4FA02FE5FF990" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FED4FA02FE5FF990" box="[293,473,1528,1555]" country="Ecuador" municipality="Otonga" name="rio Las Damas" pageId="12" pageNumber="13" stateProvince="Cotopaxi">río Las Damas</location>
;
<elevation id="0033244F13445339FE19FA02FD00F991" box="[488,646,1528,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">
<quantity id="4CE66E9913445339FE19FA02FDC5F990" box="[488,579,1528,1555]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FD67FA02FCB8F991" box="[662,830,1528,1555]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">
<collectingDate id="EFE41C5413445339FD67FA02FCB8F991" box="[662,830,1528,1555]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">3–5 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FCBCFA03FC35F990" box="[845,947,1529,1555]" pageId="12" pageNumber="13">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FC0BFA02FBBEF991" box="[1018,1080,1528,1554]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="12" pageNumber="13">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FBA6FA02FC00F9B5" ID-GBIF-Occurrence="2620303317" collectingDate="2001-07-03" collectingDateMax="2001-07-05" collectingDateMin="2001-07-03" collectionCode="CPM" collectorName="P. Moret" country="Ecuador" elevation="1800" location="Otonga" pageId="12" pageNumber="13" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Esmeraldas">
<specimenCount id="9D1808F513445339FBA6FA02FB1BF991" box="[1111,1181,1526,1554]" pageId="12" pageNumber="13" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1808F513445339FB5CFA02FB5BF990" box="[1197,1245,1528,1555]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
;
<location id="8EC195A713445339FB1FFA02FAC1F991" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FB1FFA02FAC1F991" box="[1262,1351,1528,1554]" country="Ecuador" name="Otonga" pageId="12" pageNumber="13" stateProvince="Esmeraldas">Otonga</location>
, río
<collectingRegion id="49DA0D9E13445339FF4CF9E1FECEF9B6" box="[189,328,1563,1589]" country="Ecuador" name="Esmeraldas" pageId="12" pageNumber="13">Esmeraldas</collectingRegion>
;
<elevation id="0033244F13445339FEA2F9E6FE6DF9B6" box="[339,491,1563,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">
<quantity id="4CE66E9913445339FEA2F9E6FE2AF9B5" box="[339,428,1564,1590]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="13" unit="m" value="1800.0">1800 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FE09F9E1FD1FF9B5" box="[504,665,1563,1590]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">
<collectingDate id="EFE41C5413445339FE09F9E1FD1FF9B5" box="[504,665,1563,1590]" pageId="12" pageNumber="13" value="2001-07-03" valueMax="2001-07-05" valueMin="2001-07-03">3–5 Jul. 2001</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FD55F9E6FC81F9B5" box="[676,775,1564,1590]" pageId="12" pageNumber="13">P. Moret</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FCB9F9E6FC00F9B5" box="[840,902,1564,1590]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="12" pageNumber="13">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FC6FF9E6FD5EF9DA" ID-GBIF-Occurrence="2620303349" collectedFrom="debajo de piedras en rio" collectingDate="1997-05-01" collectionCode="QCAZ" collectorName="J. Gil" country="Ecuador" elevation="2000" location="Otonga" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FC6FF9E6FC4CF9B5" box="[926,970,1564,1590]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
;
<location id="8EC195A713445339FC29F9E6FBB4F9B5" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FC29F9E6FBB4F9B5" box="[984,1074,1564,1590]" country="Ecuador" name="Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi">Otonga</location>
;
<elevation id="0033244F13445339FBCCF9E6FB53F9B6" box="[1085,1237,1563,1590]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="2000.0">
<quantity id="4CE66E9913445339FBCCF9E6FB10F9B5" box="[1085,1174,1564,1590]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FB13F9E6FAF4F9B5" box="[1250,1394,1564,1590]" pageId="12" pageNumber="13" value="1997-05-01">
<collectingDate id="EFE41C5413445339FB13F9E6FAF4F9B5" box="[1250,1394,1564,1590]" pageId="12" pageNumber="13" value="1997-05-01">1 May 1997</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FF4CF9C5FF7BF9DA" box="[189,253,1599,1625]" pageId="12" pageNumber="13">J. Gil</collectorName>
leg.; “
<collectedFrom id="3802808113445339FEBAF9C5FDF6F9DA" box="[331,624,1599,1625]" pageId="12" pageNumber="13">debajo de piedras en río</collectedFrom>
”;
<collectionCode id="ED0F5BB913445339FD79F9C5FD5EF9DA" box="[648,728,1599,1625]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="12" pageNumber="13" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FD03F9C5FE8BF9FF" ID-GBIF-Occurrence="2620303331" collectingDate="1997-01-01" collectionCode="QCAZ" collectorName="G. Onore" country="Ecuador" location="Otonga" municipality="Las Pampas" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FD03F9C5FCA6F9DA" box="[754,800,1597,1625]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BC5590613445339FCC1F9C5FC46F9DA" box="[816,960,1599,1625]" pageId="12" pageNumber="13">Las Pampas</collectingMunicipality>
,
<location id="8EC195A713445339FC3CF9C5FBA1F9DA" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FC3CF9C5FBA1F9DA" box="[973,1063,1599,1625]" country="Ecuador" municipality="Las Pampas" name="Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi">Otonga</location>
;
<date id="FFA0E5BC13445339FBC5F9C5FB46F9DA" box="[1076,1216,1599,1625]" pageId="12" pageNumber="13" value="1997-01-01">
<collectingDate id="EFE41C5413445339FBC5F9C5FB46F9DA" box="[1076,1216,1599,1625]" pageId="12" pageNumber="13" value="1997-01-01">1 Jan. 1997</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FB3CF9C5FABDF9DA" box="[1229,1339,1599,1625]" pageId="12" pageNumber="13">G. Onore</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FF4CF998FE8BF9FF" box="[189,269,1634,1660]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="12" pageNumber="13" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FED6F998FB84F9FF" ID-GBIF-Occurrence="2620303362" box="[295,1026,1634,1660]" collectingDate="1998-08-07" collectionCode="QCAZ" collectorName="I. Tapia" country="Ecuador" elevation="1900" location="Otonga" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FED6F998FED3F9FF" box="[295,341,1634,1660]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
;
<location id="8EC195A713445339FE95F998FE38F9FF" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FE95F998FE38F9FF" box="[356,446,1634,1660]" country="Ecuador" name="Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi">Otonga</location>
;
<elevation id="0033244F13445339FE3AF998FDE3F9FF" box="[459,613,1634,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="13" unit="m" value="1900.0">
<quantity id="4CE66E9913445339FE3AF998FDA2F9FF" box="[459,548,1634,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="12" pageNumber="13" unit="m" value="1900.0">1900 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FD82F998FC8EF9FF" box="[627,776,1634,1660]" pageId="12" pageNumber="13" value="1998-08-07">
<collectingDate id="EFE41C5413445339FD82F998FC8EF9FF" box="[627,776,1634,1660]" pageId="12" pageNumber="13" value="1998-08-07">7 Aug. 1998</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FCE4F998FCE8F9FF" box="[789,878,1634,1660]" pageId="12" pageNumber="13">I. Tapia</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FC43F998FB84F9FF" box="[946,1026,1634,1660]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="12" pageNumber="13" type="Museum">QCAZ</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FBEDF998FD89F941" ID-GBIF-Occurrence="2620303344" collectingDate="2016-08-13" collectionCode="CPM" collectorName="V. Crespo" country="Ecuador" elevation="1883" latitude="-0.42425" location="Otonga" longLatPrecision="1" longitude="-79.006" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Esmeraldas">
<specimenCount id="9D1808F513445339FBEDF998FBCCF9FF" box="[1052,1098,1632,1660]" pageId="12" pageNumber="13" type="male">1 ♂</specimenCount>
;
<location id="8EC195A713445339FBA8F998FB34F9FF" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FBA8F998FB34F9FF" box="[1113,1202,1634,1660]" country="Ecuador" latitude="-0.42425" longLatPrecision="1" longitude="-79.006" name="Otonga" pageId="12" pageNumber="13" stateProvince="Esmeraldas">Otonga</location>
, río
<collectingRegion id="49DA0D9E13445339FB16F998FAF4F9FF" box="[1255,1394,1634,1660]" country="Ecuador" name="Esmeraldas" pageId="12" pageNumber="13">Esmeraldas</collectingRegion>
;
<geoCoordinate id="EE2AA5BB13445339FF4CF979FECFF91C" box="[189,329,1667,1695]" degrees="0" direction="south" minutes="25.455" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="-0.42425">0°25.455′ S</geoCoordinate>
,
<geoCoordinate id="EE2AA5BB13445339FEA6F979FE7BF923" box="[343,509,1667,1696]" degrees="79" direction="west" minutes="00.360" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="-79.006">79°00.360′ W</geoCoordinate>
;
<elevation id="0033244F13445339FDFAF97FFD20F91C" box="[523,678,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.883" pageId="12" pageNumber="13" unit="m" value="1883.0">
<quantity id="4CE66E9913445339FDFAF97FFDE3F923" box="[523,613,1669,1696]" metricMagnitude="3" metricUnit="m" metricValue="1.883" pageId="12" pageNumber="13" unit="m" value="1883.0">1883 m</quantity>
a.s.l.
</elevation>
;
<date id="FFA0E5BC13445339FD45F97FFCDFF91C" box="[692,857,1669,1695]" pageId="12" pageNumber="13" value="2016-08-13">
<collectingDate id="EFE41C5413445339FD45F97FFCDFF91C" box="[692,857,1669,1695]" pageId="12" pageNumber="13" value="2016-08-13">13 Aug. 2016</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FC97F97CFC5CF91C" box="[870,986,1669,1696]" pageId="12" pageNumber="13">V. Crespo</collectorName>
leg.; COI voucher PM513, BOLD sequence SUM313-18;
<collectionCode id="ED0F5BB913445339FE20F952FD89F941" box="[465,527,1704,1730]" country="0" httpUri="http://grbio.org/cool/qy31-gen1" name="Christoffel Park Museum" pageId="12" pageNumber="13">CPM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FDD6F952FD82F965" ID-GBIF-Occurrence="2620303381" collectingDate="2007-07-11" collectingDateMax="2007-07-12" collectingDateMin="2007-07-11" collectionCode="CPMG" collectorName="W. Rossi" country="Ecuador" county="Canton Sigchos" location="Bosque Integral Otonga" municipality="Las Pampas" pageId="12" pageNumber="13" specimenCount="3" specimenCount-female="3" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FDD6F952FDEFF940" box="[551,617,1704,1731]" pageId="12" pageNumber="13" type="female">3 ♀♀</specimenCount>
;
<collectingCounty id="62C0BBF013445339FD86F952FCB4F941" box="[631,818,1704,1730]" pageId="12" pageNumber="13">Cantón Sigchos</collectingCounty>
,
<collectingMunicipality id="6BC5590613445339FCCCF953FC4BF940" box="[829,973,1705,1731]" pageId="12" pageNumber="13">Las Pampas</collectingMunicipality>
,
<location id="8EC195A713445339FC29F953FB75F941" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FC29F953FB75F941" box="[984,1267,1704,1731]" country="Ecuador" county="Canton Sigchos" municipality="Las Pampas" name="Bosque Integral Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi">Bosque Integral Otonga</location>
;
<date id="FFA0E5BC13445339FB0FF952FF7DF965" pageId="12" pageNumber="13" value="2007-07-11" valueMax="2007-07-12" valueMin="2007-07-11">
<collectingDate id="EFE41C5413445339FB0FF952FF7DF965" pageId="12" pageNumber="13" value="2007-07-11" valueMax="2007-07-12" valueMin="2007-07-11">11–12 Jul. 2007</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FEF7F936FEE9F966" box="[262,367,1739,1766]" pageId="12" pageNumber="13">W. Rossi</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FE40F936FD82F965" box="[433,516,1740,1766]" pageId="12" pageNumber="13">CPMG</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B76C92113445339FDEDF936FE50F88A" ID-GBIF-Occurrence="2620303420" collectingDate="2004-07-30" collectionCode="CPMG" collectorName="G. Caoduro &amp; G. Avesani" country="Ecuador" county="San Francisco de las Pampas" location="Otonga" municipality="Pampas" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Cotopaxi">
<specimenCount id="9D1808F513445339FDEDF936FDCFF965" box="[540,585,1740,1766]" pageId="12" pageNumber="13" type="female">1 ♀</specimenCount>
;
<collectingCounty id="62C0BBF013445339FDA6F936FC2AF965" box="[599,940,1739,1766]" pageId="12" pageNumber="13">San Francisco de las Pampas</collectingCounty>
,
<location id="8EC195A713445339FC49F936FB94F965" LSID="urn:lsid:plazi:treatment:03B7726A13445338FDAFFC1FFDC7FC7B:8EC195A713445339FC49F936FB94F965" box="[952,1042,1740,1766]" country="Ecuador" county="San Francisco de las Pampas" municipality="Pampas" name="Otonga" pageId="12" pageNumber="13" stateProvince="Cotopaxi">Otonga</location>
;
<date id="FFA0E5BC13445339FBECF936FB36F965" box="[1053,1200,1739,1766]" pageId="12" pageNumber="13" value="2004-07-30">
<collectingDate id="EFE41C5413445339FBECF936FB36F965" box="[1053,1200,1739,1766]" pageId="12" pageNumber="13" value="2004-07-30">30 Jul. 2004</collectingDate>
</date>
;
<collectorName id="26EBA6AA13445339FB4DF936FAC3F966" box="[1212,1349,1739,1766]" pageId="12" pageNumber="13">G. Caoduro</collectorName>
and
<collectorName id="26EBA6AA13445339FF4CF915FEBBF88A" box="[189,317,1775,1801]" pageId="12" pageNumber="13">G. Avesani</collectorName>
leg.;
<collectionCode id="ED0F5BB913445339FE8EF915FE50F88A" box="[383,470,1775,1801]" pageId="12" pageNumber="13">CPMG</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA1C37C13445339FF4CF8CAFE52F8C9" blockId="12.[189,468,1840,1866]" box="[189,468,1840,1866]" pageId="12" pageNumber="13">
<heading id="D0E9741013445339FF4CF8CAFE52F8C9" bold="true" box="[189,468,1840,1866]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B96A1F6E13445339FF4CF8CAFE52F8C9" bold="true" box="[189,468,1840,1866]" pageId="12" pageNumber="13">Diagnostic description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13445338FF4CF8A5FCEDFDBC" blockId="12.[189,1398,1886,2019]" lastBlockId="13.[189,1399,267,575]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Habitus:
<figureCitation id="1325DFF913445339FED6F8A5FEE8F8FA" box="[295,366,1886,1913]" captionStart="Figs 3–6. 3–4" captionStartId="11.[189,241,1919,1945]" captionTargetBox="[220,1347,329,1804]" captionTargetId="figure@11.[195,1393,311,1821]" captionTargetPageId="11" captionText="Figs 3–6. 3–4. Dyscolus eleonorae Moret sp. nov. 3. Habitus of the male holotype (QCAZ). 4. Aedeagus, median lobe in lateral view. 5–6. D. giselae Moret sp. nov. 5. Habitus of the male holotype (QCAZ). 6. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829686" httpUri="https://zenodo.org/record/3829686/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
. Wingless. Body length:
<quantity id="4CE66E9913445339FD62F8A5FCC6F8FA" box="[659,832,1887,1913]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.22" metricValueMin="1.02" pageId="12" pageNumber="13" unit="mm" value="11.2" valueMax="12.2" valueMin="10.2">10.2–12.2 mm</quantity>
. Shiny, piceous black with a faint metallic lustre on the elytra, varying from dark green to copper or pink; legs dark brown, antennae and mouthparts reddish brown to brownish. Elytral microsculpture transverse, shallowly impressed. Head broad, convex, eyes bulging, mandibles long and acute. Pronotum convex, rounded, much narrower at base than at apex, not sinuate, hind angles very obtuse, almost rounded; two pairs of lateral setae. Elytra elongate, convex, base narrow, humeri effaced; striae shallowly impressed, finely punctate, intervals flat to subconvex; third interval with three setae, except one male
<typeStatus id="54A57DDE13455338FC33FEA8FBA1FEEF" box="[962,1063,338,364]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
that bears five setae on each side; subapical sinuation deep; apical extension triangular, recurved dorsally, more or less acute at apex. Last visible abdominal ventrite with one pair (♂) or two pairs (
<specimenCount id="9D1808F513455338FC6FFE62FC37FE31" box="[926,945,408,434]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
) of setae along its apical margin. Legs moderately slender; fourth metatarsomere without subapical dorsolateral setae, apical lobes parallel, the outer lobe 1.9 times longer than the inner lobe.
<emphasis id="B96A1F6E13455338FD10FE25FC0DFE7A" box="[737,907,479,505]" italics="true" pageId="13" pageNumber="14">Male genitalia</emphasis>
:
<figureCitation id="1325DFF913455338FC6BFE24FC67FE7B" box="[922,993,478,504]" captionStart="Figs 3–6. 3–4" captionStartId="11.[189,241,1919,1945]" captionTargetBox="[220,1347,329,1804]" captionTargetId="figure@11.[195,1393,311,1821]" captionTargetPageId="11" captionText="Figs 3–6. 3–4. Dyscolus eleonorae Moret sp. nov. 3. Habitus of the male holotype (QCAZ). 4. Aedeagus, median lobe in lateral view. 5–6. D. giselae Moret sp. nov. 5. Habitus of the male holotype (QCAZ). 6. Aedeagus, median lobe in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3829686" httpUri="https://zenodo.org/record/3829686/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
. Median lobe swollen in its middle part, abruptly bent ventrally in distal fourth; apical blade long and sinuate in lateral view; endophallus without sclerotized structure.
<emphasis id="B96A1F6E13455338FDEBFDDFFD67FDBC" box="[538,737,549,575]" italics="true" pageId="13" pageNumber="14">Female genitalia</emphasis>
: unstudied.
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFD9CFEE5FD03" blockId="13.[189,355,614,640]" box="[189,355,614,640]" pageId="13" pageNumber="14">
<heading id="D0E9741013455338FF4CFD9CFEE5FD03" bold="true" box="[189,355,614,640]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B96A1F6E13455338FF4CFD9CFEE5FD03" bold="true" box="[189,355,614,640]" pageId="13" pageNumber="14">Comparisons</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFD6FFD9BFD75" blockId="13.[189,1399,661,758]" pageId="13" pageNumber="14">
Metallic coloration and apically subspinose elytra separate
<taxonomicName id="4C1EB8FF13455338FC9DFD6CFBB6FD2C" authority="Moret" authorityName="Moret" authorityYear="2020" box="[876,1072,661,687]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="giselae" status="sp. nov.">
<emphasis id="B96A1F6E13455338FC9DFD6CFC64FD2C" box="[876,994,661,687]" italics="true" pageId="13" pageNumber="14">D. giselae</emphasis>
Moret
</taxonomicName>
<taxonomicNameLabel id="A259A21513455338FBC7FD6FFB09FD2C" box="[1078,1167,661,687]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
from all the species (previously placed in the subgenus
<taxonomicName id="4C1EB8FF13455338FD80FD42FCA6FD51" authorityName="Moret" authorityYear="1996" box="[625,800,696,722]" class="Insecta" family="Carabidae" genus="Dyscolus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subGenus" subGenus="Hydrodyscolus">
<emphasis id="B96A1F6E13455338FD80FD42FCA6FD51" box="[625,800,696,722]" italics="true" pageId="13" pageNumber="14">Hydrodyscolus</emphasis>
</taxonomicName>
) which share the derived character of a dorsally asetose fourth metatarsomere.
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFCE7FE9BFCB4" blockId="13.[189,285,797,823]" box="[189,285,797,823]" pageId="13" pageNumber="14">
<heading id="D0E9741013455338FF4CFCE7FE9BFCB4" bold="true" box="[189,285,797,823]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B96A1F6E13455338FF4CFCE7FE9BFCB4" bold="true" box="[189,285,797,823]" pageId="13" pageNumber="14">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFCB1FA9BFCE6" blockId="13.[189,1309,843,870]" box="[189,1309,843,870]" pageId="13" pageNumber="14">
Riparian in the montane forest on the Western slope of the
<taxonomicName id="4C1EB8FF13455338FC9FFCB1FC3AFCE6" authorityName="Stal" authorityYear="1866" box="[878,956,843,869]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
, at around
<quantity id="4CE66E9913455338FBCCFCB1FB59FCE5" box="[1085,1247,843,870]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="13" pageNumber="14" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">1800–2000 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFC77FE6DFC24" blockId="13.[189,491,909,935]" box="[189,491,909,935]" pageId="13" pageNumber="14">
<heading id="D0E9741013455338FF4CFC77FE6DFC24" bold="true" box="[189,491,909,935]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B96A1F6E13455338FF4CFC77FE6DFC24" bold="true" box="[189,491,909,935]" pageId="13" pageNumber="14">Geographic distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1C37C13455338FF4CFC41FDC7FC7B" blockId="13.[189,1399,955,1016]" pageId="13" pageNumber="14">
Southern end of the Chocó biogeographic region in northwestern
<collectingCountry id="F30983EC13455338FC3BFC41FBA8FC56" box="[970,1070,955,981]" name="Ecuador" pageId="13" pageNumber="14">Ecuador</collectingCountry>
. Only known from the
<typeStatus id="54A57DDE13455338FAB5FC46FAF1FC55" box="[1348,1399,956,982]" pageId="13" pageNumber="14">type</typeStatus>
locality, probably microendemic.
</paragraph>
</subSubSection>
</treatment>
</document>
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