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<document id="2CF4423BBB11E4350BC60E5ED0D1C266" ID-CLB-Dataset="25550" ID-DOI="10.11646/zootaxa.4647.1.18" ID-GBIF-Dataset="6446fee7-99f6-43cd-b37a-bb6ce3fade53" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353104" ID-ZooBank="0A0CA1A2-046F-4D23-A7A9-02D2DB974D4D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564250056233" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2019" docId="03A7BA3FC92BFFDBFF53C513FAD4300D" docLanguage="en" docName="zootaxa.4647.1.18.pdf" docOrigin="Zootaxa 4647 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Topalia corinnensis Colloff 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="297" masterDocId="FF9EC247C92DFFDCFFC4C036FFE0375B" masterDocTitle="The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera" masterLastPageNumber="321" masterPageNumber="290" pageNumber="296" updateTime="1698747565907" updateUser="ExternalLinkService">
<mods:mods id="F77F19475D84130819B57FA558CAB886" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="546CFA5F4A58C176553B1A09DCDD0F3F">
<mods:title id="23B52F87775906D711F743FE5FB91CC5">The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera</mods:title>
</mods:titleInfo>
<mods:name id="FDAFD41309EAD0E44644FBFA14441DBD" type="personal">
<mods:role id="5B3C2453679DD4AD39A98EDCCEA399D4">
<mods:roleTerm id="19C7E6179FD993544416F8523F57ACA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F0635D7BF478F32BEEE803B6F305DB95">Colloff, Matthew J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FC9B99635A88A0112DE4E01618A9B7FB">text</mods:typeOfResource>
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<mods:title id="7F205BEC3F59A83D36D2D64B2365D57B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="44E3D2F16EF1DC72EFE74D1B5B388D26">
<mods:date id="C9291858A9C38AE8659C018A3266FFAA">2019</mods:date>
<mods:detail id="F91D7763F4D07275024680B284E84C70" type="pubDate">
<mods:number id="1B84D7FFED80E07852F61ED7A961BB6C">2019-07-26</mods:number>
</mods:detail>
<mods:detail id="AEFD98182B5E5130268F53B1A13159DB" type="volume">
<mods:number id="CBB24FFF66A31E8C012BB163AD52AB5C">4647</mods:number>
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<mods:number id="C4E0D631FBB91EBC5B399653D4999592">1</mods:number>
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<mods:classification id="EFD97CE93A4378431760ED45508B094B">journal article</mods:classification>
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<mods:identifier id="3D59D9F61D949A887D05DDAAE6476715" type="DOI">10.11646/zootaxa.4647.1.18</mods:identifier>
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<mods:identifier id="1CFD900B15F8F780254C2E751664C831" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="123C5E89817750CE7A1B9BEE23694FFE" type="Zenodo-Dep">3353104</mods:identifier>
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</mods:mods>
<treatment id="03A7BA3FC92BFFDBFF53C513FAD4300D" ID-GBIF-Taxon="159396553" LSID="urn:lsid:plazi:treatment:03A7BA3FC92BFFDBFF53C513FAD4300D" httpUri="http://treatment.plazi.org/id/03A7BA3FC92BFFDBFF53C513FAD4300D" lastPageId="7" lastPageNumber="297" pageId="6" pageNumber="296">
<subSubSection id="C31458A2C92BFFDAFF53C513FE0A3264" box="[151,490,1317,1344]" pageId="6" pageNumber="296" type="nomenclature">
<paragraph id="8BB10B29C92BFFDAFF53C513FE0A3264" blockId="6.[151,490,1317,1379]" box="[151,490,1317,1344]" pageId="6" pageNumber="296">
<heading id="D0F9BC45C92BFFDAFF53C513FE0A3264" bold="true" box="[151,490,1317,1344]" fontSize="11" level="1" pageId="6" pageNumber="296" reason="1">
<emphasis id="B97AD73BC92BFFDAFF53C513FE0A3264" bold="true" box="[151,490,1317,1344]" pageId="6" pageNumber="296">
<taxonomicName id="4C0E70AAC92BFFDAFF53C513FE653264" ID-CoL="8TDQV" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[151,389,1317,1344]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="296" phylum="Arthropoda" rank="species" species="corinnensis" status="sp. nov.">
<emphasis id="B97AD73BC92BFFDAFF53C513FE653264" bold="true" box="[151,389,1317,1344]" italics="true" pageId="6" pageNumber="296">Topalia corinnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92BFFDAFE48C513FE0A3264" box="[396,490,1317,1343]" pageId="6" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31458A2C92BFFDAFF53C57FFF0B3238" box="[151,235,1353,1379]" pageId="6" pageNumber="296" type="description">
<paragraph id="8BB10B29C92BFFDAFF53C57FFF0B3238" blockId="6.[151,490,1317,1379]" box="[151,235,1353,1379]" pageId="6" pageNumber="296">
(
<figureCitation id="133517ACC92BFFDAFF64C57FFF033238" box="[160,227,1353,1379]" captionStart="FIGURE 4" captionStartId="7.[151,250,1349,1374]" captionTargetBox="[237,1358,283,1312]" captionTargetId="figure@7.[227,1361,273,1324]" captionTargetPageId="7" captionText="FIGURE 4. Topalia corinnensis sp. nov. (a) dorsal aspect; (b) ventral aspect; (c) cerotegument. Scale bar = 100 µm." httpUri="https://zenodo.org/record/3353112/files/figure.png" pageId="6" pageNumber="296">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31458A2C92BFFDBFF53C5A6FAD4300D" lastPageId="7" lastPageNumber="297" pageId="6" pageNumber="296" type="materials_examined">
<paragraph id="8BB10B29C92BFFDAFF53C5A6FDCB328B" blockId="6.[151,1437,1424,2028]" pageId="6" pageNumber="296">
<emphasis id="B97AD73BC92BFFDAFF53C5A6FEC932F0" bold="true" box="[151,297,1424,1451]" pageId="6" pageNumber="296">Dimensions.</emphasis>
<typeStatus id="54B5B58BC92BFFDAFEF4C5A7FE7832F0" box="[304,408,1425,1451]" pageId="6" pageNumber="296" type="holotype">Holotype</typeStatus>
length 416 μm, breadth 252 μm.
<typeStatus id="54B5B58BC92BFFDAFCCAC5A4FC9032F7" box="[782,880,1426,1452]" pageId="6" pageNumber="296" type="paratype">Paratype</typeStatus>
length 397 μm, breadth 237 μm. Ratio of prodor- sum to total length: 0.35 (
<typeStatus id="54B5B58BC92BFFDAFE73C583FDC0328B" box="[439,544,1461,1488]" pageId="6" pageNumber="296" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BB10B29C92BFFDAFF03C5EEFF113160" blockId="6.[151,1437,1424,2028]" pageId="6" pageNumber="296">
<emphasis id="B97AD73BC92BFFDAFF03C5EEFEEB32A8" bold="true" box="[199,267,1496,1523]" pageId="6" pageNumber="296">Adult</emphasis>
.
<emphasis id="B97AD73BC92BFFDAFEDBC5ECFE7D32A8" box="[287,413,1498,1523]" italics="true" pageId="6" pageNumber="296">Integument</emphasis>
: prodorsum with sparsely-distributed tubercles in interlamellar region. Notogaster with small granules; angular foveolae lacking. Cerotegument sparing or absent in centrodorsal region. Underlying cuticle smooth.
</paragraph>
<paragraph id="8BB10B29C92BFFDAFF03C673FEC53190" blockId="6.[151,1437,1424,2028]" pageId="6" pageNumber="296">
<emphasis id="B97AD73BC92BFFDAFF03C673FEA43104" box="[199,324,1605,1631]" italics="true" pageId="6" pageNumber="296">Prodorsum</emphasis>
: rostrum incised, with two lateral teeth but no median tooth. Lamellae meeting along anterior midline and with pattern of fine, transverse striae. Interlamellar region slightly convex, smooth; setae
<emphasis id="B97AD73BC92BFFDAFB32C65CFAEC31D8" box="[1270,1292,1642,1667]" italics="true" pageId="6" pageNumber="296">in</emphasis>
on squat tubercles on posteromedian margins of lamellae. Bothridial seta with elongated, ovoid, club-shaped apices bearing blunt spines.
</paragraph>
<paragraph id="8BB10B29C92BFFDAFF03C6E0FD5D3007" blockId="6.[151,1437,1424,2028]" pageId="6" pageNumber="296">
<emphasis id="B97AD73BC92BFFDAFF03C6E0FEA331B4" box="[199,323,1750,1775]" italics="true" pageId="6" pageNumber="296">Notogaster</emphasis>
: ratio of length to breadth: 1.13. Anterior margin of notogaster wavy, indented medially. Humeral projections (
<emphasis id="B97AD73BC92BFFDAFEE0C6CFFED23048" box="[292,306,1785,1811]" italics="true" pageId="6" pageNumber="296">h</emphasis>
) rounded, well-developed, bearing seta
<emphasis id="B97AD73BC92BFFDAFD36C6CCFD1E3048" box="[754,766,1786,1811]" italics="true" pageId="6" pageNumber="296">c</emphasis>
<subScript id="178A096CC92BFFDAFD3AC73DFCE63042" attach="left" box="[766,774,1803,1817]" fontSize="6" pageId="6" pageNumber="296">2</subScript>
, incurved posteriorly at level of lyrifissurae
<emphasis id="B97AD73BC92BFFDAFB33C6CCFAED3048" box="[1271,1293,1786,1811]" italics="true" pageId="6" pageNumber="296">ia</emphasis>
. Margins of centrodorsal ridge well-developed, converging anterior of setae
<emphasis id="B97AD73BC92BFFDAFCA4C72BFC62306C" box="[864,898,1821,1847]" italics="true" pageId="6" pageNumber="296">dm</emphasis>
, then diverging; posterior part of ridge rounded, extending posteriorly to point level with setae
<emphasis id="B97AD73BC92BFFDAFD5BC777FD4D3000" box="[671,685,1857,1883]" italics="true" pageId="6" pageNumber="296">h</emphasis>
<subScript id="178A096CC92BFFDAFD69C765FD55303A" attach="left" box="[685,693,1875,1889]" fontSize="6" pageId="6" pageNumber="296">3</subScript>
.
</paragraph>
<paragraph id="8BB10B29C92BFFDBFF03C753FD4837A0" blockId="6.[151,1437,1424,2028]" lastBlockId="7.[151,1437,153,251]" lastPageId="7" lastPageNumber="297" pageId="6" pageNumber="296">
<emphasis id="B97AD73BC92BFFDAFF03C753FE893024" box="[199,361,1893,1919]" italics="true" pageId="6" pageNumber="296">Ventral aspect</emphasis>
: epimeral plates I trapezoid, twice as long as broad; epimera
<collectionCode id="ED1F93ECC92BFFDAFBF4C750FBAC30DB" box="[1072,1100,1894,1920]" pageId="6" pageNumber="296">III</collectionCode>
and
<collectionCode id="ED1F93ECC92BFFDAFB43C750FB4430DB" box="[1159,1188,1894,1920]" pageId="6" pageNumber="296">IV</collectionCode>
separate, rectangular. Enantiophysis V present; epimeral setation 2-1-2-2. Pedotectum
<collectionCode id="ED1F93ECC92BFFDAFCBBC7BCFC7230FF" box="[895,914,1930,1956]" pageId="6" pageNumber="296">II</collectionCode>
not extending laterally beyond circumgastric scissure. Perigenital carina more-or-less straight, ca.
<quantity id="4CF6A6CCC92BFFDAFD2DC79BFCD33093" box="[745,819,1965,1992]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="296" unit="mm" value="38.0">38 µm</quantity>
long, extending posteriorly to point level with seta
<emphasis id="B97AD73BC92BFFDAFABDC798FA75309C" box="[1401,1429,1966,1991]" italics="true" pageId="6" pageNumber="296">ag</emphasis>
. Genital plates
<quantity id="4CF6A6CCC92BFFDAFEFEC7E7FE6330B7" box="[314,387,2001,2028]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="296" unit="mm" value="44.0">44 µm</quantity>
long, same length as anal plates and almost three times distance between them. Cuticle ante- rior to genital plate granulate. Pedotectum I long, thin, sparingly granulate, extending anteriorly to point almost half way along length of lamellae in dorsal view. Lateral margins of pedotecta
<collectionCode id="ED1F93ECC92AFFDBFC1DC08BFC0C378C" box="[985,1004,189,215]" pageId="7" pageNumber="297">II</collectionCode>
extending to apices of bothridia setae.
<taxonomicName id="4C0E70AAC92AFFDBFF53C0D7FEC537A0" authorityName="Krefft" authorityYear="1871" box="[151,293,225,251]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="311" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
the same as in
<taxonomicName id="4C0E70AAC92AFFDBFE15C0D7FD4137A0" authorityName="Colloff" authorityYear="2019" box="[465,673,225,251]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="caliginosa">
<emphasis id="B97AD73BC92AFFDBFE15C0D7FD4137A0" box="[465,673,225,251]" italics="true" pageId="7" pageNumber="297">Topalia caliginosa</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF715BA1C92AFFDBFF53C573FAD63205" box="[151,1334,1349,1374]" httpUri="https://zenodo.org/record/3353112/files/figure.png" pageId="7" pageNumber="297" startId="7.[151,250,1349,1374]" targetBox="[237,1358,283,1312]" targetPageId="7">
<paragraph id="8BB10B29C92AFFDBFF53C573FAD63205" blockId="7.[151,1334,1349,1374]" box="[151,1334,1349,1374]" pageId="7" pageNumber="297">
<emphasis id="B97AD73BC92AFFDBFF53C573FEF33205" bold="true" box="[151,275,1349,1374]" pageId="7" pageNumber="297">FIGURE 4.</emphasis>
<taxonomicName id="4C0E70AAC92AFFDBFEDDC573FE013206" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[281,481,1349,1373]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="corinnensis" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFEDDC573FE013206" box="[281,481,1349,1373]" italics="true" pageId="7" pageNumber="297">Topalia corinnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFE23C570FDD83205" box="[487,568,1350,1374]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFE23C570FDD83205" bold="true" box="[487,568,1350,1374]" pageId="7" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
(a) dorsal aspect; (b) ventral aspect; (c) cerotegument. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph id="8BB10B29C92AFFDBFF03C599FDF83149" blockId="7.[151,1437,1455,1878]" pageId="7" pageNumber="297">
<emphasis id="B97AD73BC92AFFDBFF03C599FD7B3291" bold="true" box="[199,667,1455,1482]" pageId="7" pageNumber="297">Material Examined and Locality Data</emphasis>
.
<materialsCitation id="3B660174C92AFFDBFD68C586FDF43149" ID-GBIF-Occurrence="2413160445" collectingDate="2008-03-14" collectionCode="ANIC" collectorName="T. Weir &amp; C. Lemann" elevation="45" latitude="-41.64528" location="State Forest" longLatPrecision="19" longitude="145.09721" pageId="7" pageNumber="297" specimenCode="ANIC 3757" specimenCount="45" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="54B5B58BC92AFFDBFD68C586FCF43291" box="[684,788,1456,1482]" pageId="7" pageNumber="297" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54B5B58BC92AFFDBFC95C586FC563291" box="[849,950,1456,1482]" pageId="7" pageNumber="297" type="paratype">paratype</typeStatus>
:
<specimenCode id="DBA8A352C92AFFDBFC06C586FBAC3291" box="[962,1100,1455,1482]" collectionCode="ANIC" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="297">ANIC 3757</specimenCode>
, sifted litter and mossy logs, rainforest with
<taxonomicName id="4C0E70AAC92AFFDBFE86C5E2FE5E32B5" box="[322,446,1492,1518]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="297" phylum="Tracheophyta" rank="genus">
<emphasis id="B97AD73BC92AFFDBFE86C5E2FE5E32B5" box="[322,446,1492,1518]" italics="true" pageId="7" pageNumber="297">Eucalyptus</emphasis>
</taxonomicName>
sp.,
<location id="8ED15DF2C92AFFDBFE30C5E2FD9C32B5" LSID="urn:lsid:plazi:treatment:03A7BA3FC92BFFDBFF53C513FAD4300D:8ED15DF2C92AFFDBFE30C5E2FD9C32B5" box="[500,636,1492,1518]" latitude="-41.64528" longLatPrecision="19" longitude="145.09721" name="State Forest" pageId="7" pageNumber="297" stateProvince="Tasmania">State Forest</location>
,
<locationDeviation id="9A03120BC92AFFDBFD43C5E2FC8532B5" box="[647,869,1492,1518]" location="Corinna" pageId="7" pageNumber="297">
<quantity id="4CF6A6CCC92AFFDBFD43C5E2FD3532B5" box="[647,725,1492,1518]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="297" unit="km" value="2.2">2.2 km</quantity>
NE Corinna
</locationDeviation>
,
<collectingRegion id="49CAC5CBC92AFFDBFCB4C5E2FC3E32B5" box="[880,990,1492,1518]" country="Australia" name="Tasmania" pageId="7" pageNumber="297">Tasmania</collectingRegion>
,
<geoCoordinate id="EE3A6DEEC92AFFDBFC2DC5E2FB8D32B5" box="[1001,1133,1492,1518]" degrees="41" direction="south" minutes="38" orientation="latitude" pageId="7" pageNumber="297" precision="15" seconds="43" value="-41.64528">41°38’43”S</geoCoordinate>
<geoCoordinate id="EE3A6DEEC92AFFDBFBB0C5E5FAEE32B5" box="[1140,1294,1491,1518]" degrees="145" direction="east" minutes="05" orientation="longitude" pageId="7" pageNumber="297" precision="15" seconds="50" value="145.09721">145°05’50”E</geoCoordinate>
.,
<specimenCount id="9D08C0A0C92AFFDBFAD9C5E5FABF32B5" box="[1309,1375,1491,1518]" pageId="7" pageNumber="297" type="generic">
<elevation id="0023EC1AC92AFFDBFAD9C5E5FABB32B5" box="[1309,1371,1491,1518]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="297" unit="m" value="45.0">
<quantity id="4CF6A6CCC92AFFDBFAD9C5E5FABB32B5" box="[1309,1371,1491,1518]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="297" unit="m" value="45.0">45 m</quantity>
</elevation>
.
</specimenCount>
, coll.
<collectorName id="26FB6EFFC92AFFDBFF53C5CEFF0A3149" box="[151,234,1528,1554]" pageId="7" pageNumber="297">T. Weir</collectorName>
&amp;
<collectorName id="26FB6EFFC92AFFDBFEC9C5CEFE6D3149" box="[269,397,1528,1554]" pageId="7" pageNumber="297">C. Lemann</collectorName>
,
<collectingDate id="EFF4D401C92AFFDBFE5CC5CEFDF43149" box="[408,532,1528,1554]" pageId="7" pageNumber="297" value="2008-03-14">
<date id="FFB02DE9C92AFFDBFE5CC5CEFDF43149" box="[408,532,1528,1554]" pageId="7" pageNumber="297" value="2008-03-14">14.iii.2008</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB10B29C92AFFDBFF03C62AFBEF316D" blockId="7.[151,1437,1455,1878]" box="[199,1039,1564,1590]" pageId="7" pageNumber="297">
<emphasis id="B97AD73BC92AFFDBFF03C62AFEA8316D" bold="true" box="[199,328,1564,1590]" pageId="7" pageNumber="297">Etymology</emphasis>
. This species is named for its
<typeStatus id="54B5B58BC92AFFDBFD5FC62AFD2B316D" box="[667,715,1564,1590]" pageId="7" pageNumber="297">type</typeStatus>
locality; Corinna,
<collectingRegion id="49CAC5CBC92AFFDBFC58C62AFBEB316D" box="[924,1035,1564,1590]" country="Australia" name="Tasmania" pageId="7" pageNumber="297">Tasmania</collectingRegion>
.
</paragraph>
<paragraph id="8BB10B29C92AFFDBFF03C609FCBD319D" blockId="7.[151,1437,1455,1878]" pageId="7" pageNumber="297">
<emphasis id="B97AD73BC92AFFDBFF03C609FEA23101" bold="true" box="[199,322,1599,1626]" pageId="7" pageNumber="297">Diagnosis.</emphasis>
<taxonomicName id="4C0E70AAC92AFFDBFE8DC676FDC63101" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[329,550,1600,1626]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="corinnensis" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFE8DC676FDC63101" box="[329,550,1600,1626]" italics="true" pageId="7" pageNumber="297">Topalia corinnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFDE9C676FD683101" box="[557,648,1600,1626]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFDE9C676FD623101" bold="true" box="[557,642,1600,1626]" pageId="7" pageNumber="297">sp. nov</emphasis>
.
</taxonomicNameLabel>
can be distinguished from other species in the genus by the following combination of characters: (1) rostrum incised medially; (2) anteromedial edges of lamellae meeting; (3) setae
<emphasis id="B97AD73BC92AFFDBFA42C653FA7C3125" box="[1414,1436,1637,1662]" italics="true" pageId="7" pageNumber="297">in</emphasis>
on squat tubercles; (4) anterior margin of notogaster wavy, indented medially; (5) epimera
<collectionCode id="ED1F93ECC92AFFDBFB60C6BEFB2031F9" box="[1188,1216,1672,1698]" pageId="7" pageNumber="297">III</collectionCode>
and
<collectionCode id="ED1F93ECC92AFFDBFB3EC6BEFAF731F9" box="[1274,1303,1672,1698]" pageId="7" pageNumber="297">IV</collectionCode>
completely separated; (5) with enantiophysis V between epimera
<collectionCode id="ED1F93ECC92AFFDBFD35C69AFCE4319D" box="[753,772,1708,1734]" pageId="7" pageNumber="297">II</collectionCode>
and
<collectionCode id="ED1F93ECC92AFFDBFCFEC69AFCB9319D" box="[826,857,1708,1734]" pageId="7" pageNumber="297">III</collectionCode>
.
</paragraph>
<paragraph id="8BB10B29C92AFFDBFF03C6E6FAD4300D" blockId="7.[151,1437,1455,1878]" pageId="7" pageNumber="297">
<emphasis id="B97AD73BC92AFFDBFF03C6E6FED431B1" bold="true" box="[199,308,1744,1770]" pageId="7" pageNumber="297">Remarks</emphasis>
.
<taxonomicName id="4C0E70AAC92AFFDBFE85C6E6FDFD31B1" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[321,541,1744,1770]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="corinnensis" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFE85C6E6FDFD31B1" box="[321,541,1744,1770]" italics="true" pageId="7" pageNumber="297">Topalia corinnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFDE0C6E6FD9D31B1" box="[548,637,1744,1770]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFDE0C6E6FD9D31B1" bold="true" box="[548,637,1744,1770]" pageId="7" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
is morphologically most similar to
<taxonomicName id="4C0E70AAC92AFFDBFBCCC6E7FBAF31B1" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[1032,1103,1745,1770]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="royi" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFBCCC6E7FBAF31B1" box="[1032,1103,1745,1770]" italics="true" pageId="7" pageNumber="297">T. royi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFB92C6E6FB4F31B1" box="[1110,1199,1744,1770]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFB92C6E6FB4A31B1" bold="true" box="[1110,1194,1744,1770]" pageId="7" pageNumber="297">sp. nov</emphasis>
.
</taxonomicNameLabel>
, with which it shares the wavy anterior notogastral margin with a median indentation and the lamellae with fine striae.
<taxonomicName id="4C0E70AAC92AFFDBFB10C6C2FF553069" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="corinnensis" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFB10C6C2FF553069" italics="true" pageId="7" pageNumber="297">Topalia corinnensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFF79C72EFEF73069" box="[189,279,1816,1842]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFF79C72EFEF73069" bold="true" box="[189,279,1816,1842]" pageId="7" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C0E70AAC92AFFDBFE6AC72FFE173069" authority="Colloff, 2019" authorityName="Colloff" authorityYear="2019" box="[430,503,1817,1842]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="royi" status="sp. nov.">
<emphasis id="B97AD73BC92AFFDBFE6AC72FFE173069" box="[430,503,1817,1842]" italics="true" pageId="7" pageNumber="297">T. royi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2496A40C92AFFDBFE3BC72EFDBA3069" box="[511,602,1816,1842]" pageId="7" pageNumber="297" rank="species">
<emphasis id="B97AD73BC92AFFDBFE3BC72EFDBA3069" bold="true" box="[511,602,1816,1842]" pageId="7" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
in the separated epimera
<collectionCode id="ED1F93ECC92AFFDBFCBAC72EFC713069" box="[894,913,1816,1842]" pageId="7" pageNumber="297">II</collectionCode>
and
<collectionCode id="ED1F93ECC92AFFDBFC0DC72EFC063069" box="[969,998,1816,1842]" pageId="7" pageNumber="297">IV</collectionCode>
, the granular, non-polygonal cerotegument and the presence of enantiophysis V, a character state it shares with
<taxonomicName id="4C0E70AAC92AFFDBFC09C70BFBA7300D" authorityName="Mahunka" authorityYear="1985" box="[973,1095,1852,1878]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B97AD73BC92AFFDBFC09C70BFBA7300D" box="[973,1095,1852,1878]" italics="true" pageId="7" pageNumber="297">T. africana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E70AAC92AFFDBFBBAC70BFACD300D" authorityName="Balogh &amp; Csiszar" authorityYear="1963" box="[1150,1325,1852,1878]" class="Arachnida" family="Nosybeidae" genus="Topalia" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="297" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B97AD73BC92AFFDBFBBAC70BFACD300D" box="[1150,1325,1852,1878]" italics="true" pageId="7" pageNumber="297">T. problematica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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