171 lines
23 KiB
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171 lines
23 KiB
XML
<document id="032E930701909C41CC5B1B8CE95EB89D" ID-DOI="10.5281/zenodo.5312012" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312012" ID-ZooBank="D72D1DC9-87D1-4209-A01F-8185A4D21F88" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118846529" checkinUser="marcus" docAuthor="Sehnal, Richard, Král, David & Bezděk, Aleš" docDate="2014" docId="03A4923D7D78AD2715D5FEECF6B84B3F" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.139-171.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Canuschiza Lacroix 1999" docType="treatment" docVersion="4" lastPageNumber="142" masterDocId="FF9DEA457D7AAD241413FFF8F470496C" masterDocTitle="Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group" masterLastPageNumber="171" masterPageNumber="139" pageNumber="141" updateTime="1699158890131" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="62AABD40A423F2FC5B54AF3F4D798774">Canuschiza of Socotra Island (Coleoptera: Scarabaeidae: Melolonthinae) Part 1. Canuschiza insularis species group</mods:title>
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<mods:namePart id="19725973A5BF89340A83207346AF58D9">Sehnal, Richard</mods:namePart>
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<mods:affiliation id="6209998F669AF2011EDD7370333C9559">Czech University of Life Sciences Prague, Faculty of Agrobiology, Food and Natural Resources, Department of Zoology and Fisheries, Kamýcká 129, CZ- 165 21 Praha 6, Czech Republic; e-mail: richard. sehnal @ seznam. cz</mods:affiliation>
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<mods:namePart id="0AA135937D5E65E63AA3B5437E7CA19D">Král, David</mods:namePart>
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<mods:affiliation id="C166AD15079CFF324DCF4628ADC76CBE">Charles University in Prague, Faculty of Science, Department of Zoology, Viničná 7, CZ- 128 43 Praha 2, Czech Republic; e-mail: kral @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="36611BA54C9450F40C871546F5050BAA">Bezděk, Aleš</mods:namePart>
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<mods:affiliation id="82436838910134222E4DB4A872EB2B15">Biology Centre ASCR, Institute of Entomology, Branišovská 31, CZ- 370 05 České Budějovice, Czech Republic; e-mail: bezdek @ entu. cas. cz</mods:affiliation>
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<mods:title id="1F26AAF6EA19A6B00963C86A72FAD3AF">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="4831C0E08F2A90F15BB23ABB8946503E">2014</mods:date>
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<paragraph id="8BB2232B7D78AD2615D5FEECF7474847" blockId="2.[454,823,275,299]" box="[454,823,275,299]" pageId="2" pageNumber="141">
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<heading id="D0FA94477D78AD2615D5FEECF7474847" bold="true" box="[454,823,275,299]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="141" reason="3">
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<emphasis id="B979FF397D78AD2615D5FEECF7474847" bold="true" box="[454,823,275,299]" pageId="2" pageNumber="141">
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Genus
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<taxonomicName id="4C0D58A87D78AD261606FEEBF7474847" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" box="[533,823,275,299]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B979FF397D78AD261606FEEBF6E64847" bold="true" box="[533,662,275,299]" italics="true" pageId="2" pageNumber="141">Canuschiza</emphasis>
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<bibRefCitation id="EF9C5EDA7D78AD26168EFEECF7474847" author="LACROIX M." box="[669,823,276,299]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
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<taxonomicName id="4C0D58A87D78AD261484FEC1F5EB4821" authority="Lacroix, 1999: 89" authorityName="Lacroix" authorityPageNumber="89" authorityYear="1999" box="[151,411,313,333]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B979FF397D78AD261484FEC1F48B4820" box="[151,251,313,332]" italics="true" pageId="2" pageNumber="141">Canuschiza</emphasis>
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<bibRefCitation id="EF9C5EDA7D78AD261513FEC2F5EB4821" author="LACROIX M." box="[256,411,314,333]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999: 89</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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,
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<figureCitation id="13363FAE7D78AD2615B7FEC2F5A54821" box="[420,469,314,333]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="2" pageNumber="141">Figs 1</figureCitation>
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–27 (original description);
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<treatmentCitation id="0AAC053A7D78AD2616ADFEC2F7004821" author="LACROIX M." box="[702,880,314,333]" page="404" pageId="2" pageNumber="141" year="2002">
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<bibRefCitation id="EF9C5EDA7D78AD2616ADFEC2F7364821" author="LACROIX M." box="[702,838,314,333]" pageId="2" pageNumber="141" pagination="399 - 414" refId="ref15524" refString="LACROIX M. 2002: Melolonthinae (Coleoptera: Scarabaeidae) of Socotra island, Yemen. Fauna of Arabia 19: 399 - 414." type="journal article" year="2002">LACROIX (2002)</bibRefCitation>
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: 404
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||
</treatmentCitation>
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,
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<figureCitation id="13363FAE7D78AD26176AFEC2F7DA4821" box="[889,938,314,333]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="2" pageNumber="141">Figs 1</figureCitation>
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–25 (review);
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||
<treatmentCitation id="0AAC053A7D78AD261034FEC2F56A4804" author="BEZDEK A." page="199" pageId="2" pageNumber="141" year="2006">
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<bibRefCitation id="EF9C5EDA7D78AD261034FEC2F4814804" author="BEZDEK A." pageId="2" pageNumber="141" pagination="228" refId="ref15172" refString="BEZDEK A. 2006: Tribe Schizonychini Burmeister, 1855. P. 228. In: LOBL I. & SMETANA A. (eds): Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea." type="book chapter" year="2006">
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<subScript id="1789216E7D78AD261034FEC2F0174827" attach="right" box="[1063,1127,314,333]" fontSize="6" pageId="2" pageNumber="141">BEZDĚK</subScript>
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(2006)
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</bibRefCitation>
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: 199
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||
</treatmentCitation>
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(catalogue);
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||
<treatmentCitation id="0AAC053A7D78AD261599FEADF64A4804" author="LACROIX M." box="[394,570,341,360]" page="207" pageId="2" pageNumber="141" year="2010">
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<bibRefCitation id="EF9C5EDA7D78AD261599FEADF6624804" author="LACROIX M." box="[394,530,341,360]" pageId="2" pageNumber="141" refId="ref15550" refString="LACROIX M. 2010: Melolonthinae afrotropicaux (Scarabaeoidea, Melolonthidae). Genera et catalogue commente. Editions Marc Lacroix, Paris, 277 pp + 192 pls." type="book" year="2010">LACROIX (2010)</bibRefCitation>
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: 207
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||
</treatmentCitation>
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(catalogue).
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</paragraph>
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<subSubSection id="C31770A07D78AD261484FE78F7C948FB" box="[151,953,384,408]" pageId="2" pageNumber="141" type="type_taxon">
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<paragraph id="8BB2232B7D78AD261484FE78F7C948FB" blockId="2.[151,1127,384,599]" box="[151,953,384,408]" pageId="2" pageNumber="141">
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<emphasis id="B979FF397D78AD261484FE78F55B48FB" bold="true" box="[151,299,384,407]" pageId="2" pageNumber="141">
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<typeStatus id="54B69D897D78AD261484FE78F4BF48FB" box="[151,207,384,407]" pageId="2" pageNumber="141">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="4C0D58A87D78AD261521FE78F6C148FB" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" box="[306,689,384,408]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="insularis">
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<emphasis id="B979FF397D78AD261521FE78F66248F4" box="[306,530,384,408]" italics="true" pageId="2" pageNumber="141">Canuschiza insularis</emphasis>
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<bibRefCitation id="EF9C5EDA7D78AD26160AFE78F6C148FB" author="LACROIX M." box="[537,689,384,407]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31770A07D78AD271484FE58F6B84B3F" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141" type="diagnosis">
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<paragraph id="8BB2232B7D78AD271484FE58F5974B5F" blockId="2.[151,1127,384,599]" lastBlockId="3.[113,1090,220,595]" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
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<emphasis id="B979FF397D78AD261484FE58F5FE48DB" bold="true" box="[151,398,416,439]" pageId="2" pageNumber="141">Differential diagnosis.</emphasis>
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Genus
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<taxonomicName id="4C0D58A87D78AD2615C8FE58F62748D4" authorityName="Lacroix" authorityYear="1999" box="[475,599,416,440]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
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<emphasis id="B979FF397D78AD2615C8FE58F62748D4" box="[475,599,416,440]" italics="true" pageId="2" pageNumber="141">Canuschiza</emphasis>
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</taxonomicName>
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differs from other schizonychine genera in combination of the following characters. Body elongate (length
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<quantity id="4CF58ECE7D78AD2616FBFE38F70648BB" box="[744,886,448,471]" metricMagnitude="-2" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.72" metricValueMin="0.75" pageId="2" pageNumber="141" unit="mm" value="12.35" valueMax="17.2" valueMin="7.5">7.5–17.2 mm</quantity>
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), almost parallel-sided, weakly to moderately convex. Labrum transverse, deeply bilobed, lobes rounded. Clypeus transverse, anterior angles rounded. Fronto-clypeal suture present, forming continuous shallowly bisinuate line or better visible elevated double fronto-clypeal carina (
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<figureCitation id="13363FAE7D78AD2617AAFDD8F05B4B5B" box="[953,1067,544,567]" captionStart="Figs 1" captionStartId="2.[148,185,1540,1559]" captionTargetBox="[237,1038,641,1515]" captionTargetId="figure-123@2.[232,1044,633,1522]" captionTargetPageId="2" captionText="Figs 1A–D. Canuschiza Lacroix, 1999:A–B – head, dorsal view; C–D – metathoracic wing.A – Canuschiza insularis Lacroix, 1999, holotype; B – C. minuta Lacroix, 1999, holotype; C – C. hagher sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 2.0 mm. Figs A, B not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312014" httpUri="https://zenodo.org/record/5312014/files/figure.png" pageId="2" pageNumber="141">Figs 1A,B</figureCitation>
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). Occipital carina missing. Eyes small (width of both eyes combined lower than maximum width of frons between eyes) to large (width of both eyes combined larger than maximum width of frons between eyes). Antenna with eight to ten antennomeres; club with three antennomeres. Terminal maxillary palpomere elongate, truncate apically. Pronotum transverse, simply convex, without anteromedian tubercle. Basal border complete, anterior border missing. Posterior angles of pronotum broadly rounded to rectangular, in some species even distinctly produced posteriad. Elytra weakly to moderately convex, parallel-sided, rounded apically, apical angle approximately rectangular. Striae missing, except of feebly visible sutural stria. Macropterous, rarely micropterous. Protibia narrow, tridentate, terminal calcar inserted against medial dent. Claws bifid, with ventrobasal teeth (e.g.
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<figureCitation id="13363FAE7D79AD271678FE24F6B0489F" box="[619,704,476,499]" captionStart="Figs 2" captionStartId="5.[113,150,1529,1548]" captionTargetBox="[130,1078,233,1493]" captionTargetId="figure-14@5.[121,1082,219,1495]" captionTargetPageId="5" captionText="Figs 2A–H. Canuschiza adah sp. nov., dorsal view: A–D – male, holotype, body length 13.7 mm; E–H – female, paratype No. 7, body length 13.0 mm. A, E – habitus; B, F – protarsal claw; C, G – detail of laterobasal area of pronotum and elytron; D, H – head. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5312018" httpUri="https://zenodo.org/record/5312018/files/figure.png" pageId="3" pageNumber="142">Fig. 2B</figureCitation>
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). Parameres symmetrical, regularly curved in lateral aspect, rounded apically in dorsal aspect; apex with tuft of long yellowish macrosetae (
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<figureCitation id="13363FAE7D79AD2714E4FDE4F50F4B5F" box="[247,383,540,563]" captionStart="Figs 10" captionStartId="27.[113,150,1513,1532]" captionTargetBox="[121,1075,225,1464]" captionTargetId="figure-14@27.[113,1088,220,1476]" captionTargetPageId="27" captionText="Figs 10A–D.Aedeagus in dorsal (left) and lateral (right) view:A – Canuschiza adah sp. nov., holotype; B – C. croton sp. nov., holotype; C – C. dracaena sp. nov., holotype; D – C. firmihin sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312042" httpUri="https://zenodo.org/record/5312042/files/figure.png" pageId="3" pageNumber="142">Figs 10A–D</figureCitation>
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,
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<figureCitation id="13363FAE7D79AD27159AFDE4F5AB4B5F" box="[393,475,540,563]" captionStart="Figs 11" captionStartId="28.[150,187,1487,1506]" captionTargetBox="[159,1117,227,1393]" captionTargetId="figure-11@28.[151,1126,218,1396]" captionTargetPageId="28" captionText="Figs 11A–D. Aedeagus in dorsal (left) and lateral (right) view: A – Canuschiza hagher sp. nov., holotype; B – C. insularis Lacroix, 1999, holotype; C – C. jatropha sp. nov., holotype; D – C. skand sp. nov., holotype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5312044" httpUri="https://zenodo.org/record/5312044/files/figure.png" pageId="3" pageNumber="142">11A–D</figureCitation>
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||
).
|
||
</paragraph>
|
||
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|
||
<paragraph id="8BB2232B7D78AD261487F9FCF60D4F20" blockId="2.[148,1127,1539,1612]" pageId="2" pageNumber="141">
|
||
Figs 1A–D.
|
||
<taxonomicName id="4C0D58A87D78AD2614E9F9FBF5AA4F7B" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" box="[250,474,1539,1559]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979FF397D78AD2614E9F9FBF52D4F7A" box="[250,349,1539,1558]" italics="true" pageId="2" pageNumber="141">Canuschiza</emphasis>
|
||
<bibRefCitation id="EF9C5EDA7D78AD261572F9FCF5AA4F7B" author="LACROIX M." box="[353,474,1540,1559]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
:A–B – head, dorsal view; C–D – metathoracic wing.A –
|
||
<taxonomicName id="4C0D58A87D78AD2617A7F9FBF57E4F5D" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="insularis">
|
||
<emphasis id="B979FF397D78AD2617A7F9FBF0154F7A" box="[948,1125,1539,1558]" italics="true" pageId="2" pageNumber="141">Canuschiza insularis</emphasis>
|
||
<bibRefCitation id="EF9C5EDA7D78AD261486F9E6F57E4F5D" author="LACROIX M." box="[149,270,1566,1585]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype; B –
|
||
<taxonomicName id="4C0D58A87D78AD26159DF9E6F6104F5D" authority="Lacroix, 1999" authorityName="Lacroix" authorityYear="1999" box="[398,608,1566,1585]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="minuta">
|
||
<emphasis id="B979FF397D78AD26159DF9E6F5914F5D" box="[398,481,1566,1585]" italics="true" pageId="2" pageNumber="141">C. minuta</emphasis>
|
||
<bibRefCitation id="EF9C5EDA7D78AD2615F4F9E6F6104F5D" author="LACROIX M." box="[487,608,1566,1585]" pageId="2" pageNumber="141" pagination="87 - 96" refId="ref15497" refString="LACROIX M. 1999: Melolonthidae de Socotra (Coleoptera, Scarabaeoidea). Revue Francaise d'Entomologie (Nouvelle Serie) 21: 87 - 96." type="journal article" year="1999">Lacroix, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype; C –
|
||
<taxonomicName id="4C0D58A87D78AD2616F3F9E6F7444F5D" authority="Sehnal & Král & Bezděk, 2014" authorityName="Sehnal & Král & Bezděk" authorityYear="2014" box="[736,820,1566,1585]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="hagher" status="sp. nov.">
|
||
<emphasis id="B979FF397D78AD2616F3F9E6F7444F5D" box="[736,820,1566,1585]" italics="true" pageId="2" pageNumber="141">C. hagher</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A42427D78AD261729F9E6F70E4F5D" box="[826,894,1566,1585]" pageId="2" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype; D –
|
||
<taxonomicName id="4C0D58A87D78AD2617EDF9E6F0394F5D" authority="Sehnal & Král & Bezděk, 2014" authorityName="Sehnal & Král & Bezděk" authorityYear="2014" box="[1022,1097,1566,1585]" class="Insecta" family="Melolonthidae" genus="Canuschiza" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="skand" status="sp. nov.">
|
||
<emphasis id="B979FF397D78AD2617EDF9E6F0394F5D" box="[1022,1097,1566,1585]" italics="true" pageId="2" pageNumber="141">C. skand</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A42427D78AD26105EF9E6F4CC4F20" pageId="2" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. Scale bar: 2.0 mm. Figs A, B not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2232B7D79AD271462FDC4F6B84B3F" blockId="3.[113,1090,220,595]" box="[113,712,572,595]" pageId="3" pageNumber="142">
|
||
<emphasis id="B979FF397D79AD271462FDC4F5D04B3F" bold="true" box="[113,416,572,595]" pageId="3" pageNumber="142">Geographical distribution.</emphasis>
|
||
Endemic to
|
||
<collectingCountry id="F31A63BB7D79AD271634FDC4F6B44B3F" box="[551,708,572,595]" name="Socotra Island" pageId="3" pageNumber="142">Socotra Island</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |