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<document id="9EA578F084B40F61D1EC3C11C23149C9" ID-DOI="10.11646/phytotaxa.268.2.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="13666769" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725392870972" checkinUser="felipe" docAuthor="Polyiam, Jutarat Kalb Wetchasart, Plata, Eimy Rivas, Bawingan, Paulina A., Kalb, Klaus &amp; Lücking, Robert" docDate="2016" docId="0397B65BFFBE5C30FF75F8B255847135" docLanguage="en" docName="phytotaxa.268.2.2.pdf" docOrigin="Phytotaxa 268 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.268.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Dyplolabia dalywaiana Rivas Plata, Bawingan &amp; Lucking 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="119" masterDocId="FFAECE23FFB95C3AFFFDFFF154697010" masterDocTitle=" Missing links alive? Novel taxa represent morphological transitions between distinctive phenotypes among extant Graphidaceae (lichenized Ascomycota: Ostropales)" masterLastPageNumber="122" masterPageNumber="110" pageNumber="117" updateTime="1725414794177" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1A32EEB860274D664D38C64944FEE1C3"> Missing links alive? Novel taxa represent morphological transitions between distinctive phenotypes among extant Graphidaceae (lichenized Ascomycota: Ostropales)</mods:title>
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<mods:namePart id="D65F2D95A699FAD2ABDB2DA2EA3C0219">Polyiam, Jutarat Kalb Wetchasart</mods:namePart>
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<mods:namePart id="ED852DFF1CA4ABA6AB1F6C75E9F032A1">Plata, Eimy Rivas</mods:namePart>
<mods:affiliation id="7BC71A1A8EB2E54898006708F03195DA">Science &amp; Education, Integrative Research &amp; Collections Center, The Field Museum, 1400 South Lake Shore Drive, Chicago, Illinois 60605 - 2496, U. S. A.</mods:affiliation>
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<mods:namePart id="5B6575605F2909A8E4CCD6E6AF8DF16D">Bawingan, Paulina A.</mods:namePart>
<mods:affiliation id="8277EFB2B80A2177CD5FC60694C2D0E0">School of Natural Sciences, Saint Louis University, Baguio City, Philippines.</mods:affiliation>
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<mods:namePart id="3AF6D735D88BA6CB4C430E6F8C15F310">Kalb, Klaus</mods:namePart>
<mods:affiliation id="F52402CA1A4F52935FAAF0345377FBDA">Lichenologisches Institut Neumarkt, Im Tal 12, D- 92318 Neumarkt, Germany, and University of Regensburg, Institute of Plant Sciences, Universitätsstraβe 31, D- 93040 Regensburg, Germany.</mods:affiliation>
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<mods:namePart id="4994454D7F3B585C5A1C7076404612FE">Lücking, Robert</mods:namePart>
<mods:affiliation id="40C0BE315AEC2D2AFF375926815D0F6F">Botanical Garden and Botanical Museum, Königin-Luise-Strasse 6 - 8, 14195 Berlin, Germany.</mods:affiliation>
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<mods:title id="3C06B6F58D9DC3009DBBCEF5148BEF0D">Phytotaxa</mods:title>
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<mods:date id="4F541774D5B5908D97DD871EAC03D396">2016</mods:date>
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<heading id="D0C9B021FFBE5C3DFF75F8B2506A774E" box="[136,1027,1859,1886]" fontSize="11" level="2" pageId="7" pageNumber="117" reason="3">
<emphasis id="B94ADB5FFFBE5C3DFF75F8B257FA774E" bold="true" box="[136,915,1859,1886]" pageId="7" pageNumber="117">
<taxonomicName id="4C3E7CCEFFBE5C3DFF75F8B25758774E" ID-CoL="386J2" authority="Rivas Plata, Bawingan &amp; Lucking" authorityName="Rivas Plata, Bawingan &amp; Lucking" authorityYear="2016" box="[136,817,1859,1886]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="7" pageNumber="117" phylum="Ascomycota" rank="species" species="dalywaiana" status="sp. nov.">
<emphasis id="B94ADB5FFFBE5C3DFF75F8B255F8774E" bold="true" box="[136,401,1859,1886]" italics="true" pageId="7" pageNumber="117">Dyplolabia dalywaiana</emphasis>
Rivas Plata, Bawingan &amp; Lücking
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,
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<taxonomicNameLabel id="A2796624FFBE5C3DFCC0F8B557FA774E" box="[829,915,1860,1886]" pageId="7" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
(
<figureCitation id="13051BC8FFBE5C3DFC5EF8B45792774E" box="[931,1019,1861,1886]" captionStart="FIGURE 6" captionStartId="9.[136,229,1576,1597]" captionTargetBox="[189,1398,190,1551]" captionTargetId="figure-14@9.[189,1398,190,1551]" captionTargetPageId="9" captionText="FIGURE 6. A. Dyplolabia dalywaiana (holotype; thallus with ascomata). BD. D. oryzoides (B, Brazil, Kalb 31120; C, D, Sri Lanka, Weerakoon 116; thallus with ascomata and ascospores). EF. D. chumphonensis (holotype; thallus with ascomata and ascospores). Scale = 1 mm, in D, F = 10 μm." figureDoi="http://doi.org/10.5281/zenodo.13666795" httpUri="https://zenodo.org/record/13666795/files/figure.png" pageId="7" pageNumber="117">Fig. 6A</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8B81074DFFBE5C3DFF75F89855E27792" blockId="7.[136,395,1897,1922]" box="[136,395,1897,1922]" pageId="7" pageNumber="117">MycoBank MB817437</paragraph>
</subSubSection>
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<paragraph id="8B81074DFFBE5C3DFF75F84550747801" blockId="7.[136,1452,1972,2065]" pageId="7" pageNumber="117">
<materialsCitation id="3B560D10FFBE5C3DFF75F845572277FD" collectorName="Differing &amp; Type" country="Philippines" county="Luzon" location="Solano" municipality="Mt. Palali" pageId="7" pageNumber="117" specimenCount="1" stateProvince="Nueva Vizcaya">
<collectorName id="26CB629BFFBE5C3DFF75F845548877D9" box="[136,225,1972,1993]" pageId="7" pageNumber="117">Differing</collectorName>
from other
<taxonomicName id="4C3E7CCEFFBE5C3DFEAEF845566377D9" box="[339,522,1972,1993]" pageId="7" pageNumber="117">
<emphasis id="B94ADB5FFFBE5C3DFEAEF84555D677D9" box="[339,447,1972,1993]" italics="true" pageId="7" pageNumber="117">Dyplolabia</emphasis>
species
</taxonomicName>
in the angular, erumpent asomata with broadly exposed disc and irregular pseudocolumella.
<emphasis id="B94ADB5FFFBE5C3DFF75F82954AB77FD" bold="true" box="[136,194,2008,2029]" pageId="7" pageNumber="117">
<collectorName id="26CB629BFFBE5C3DFF75F82954D777FD" box="[136,190,2008,2029]" pageId="7" pageNumber="117">Type</collectorName>
:
</emphasis>
<collectingCountry id="F32947DDFFBE5C3DFF27F829550277FD" box="[218,363,2008,2029]" name="Philippines" pageId="7" pageNumber="117">PHILIPPINES</collectingCountry>
.
<collectingRegion id="49FAC9AFFFBE5C3DFE89F829566F77FD" box="[372,518,2008,2029]" country="Philippines" name="Nueva Vizcaya" pageId="7" pageNumber="117">Nueva Vizcaya</collectingRegion>
(
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):
<collectingMunicipality id="6BE59D37FFBE5C3DFD95F82956A377FD" box="[616,714,2008,2029]" pageId="7" pageNumber="117">Mt. Palali</collectingMunicipality>
, near
<location id="8EE15196FFBE5C3DFCF8F829572277FD" LSID="urn:lsid:plazi:treatment:0397B65BFFBE5C30FF75F8B255847135:8EE15196FFBE5C3DFCF8F829572277FD" box="[773,843,2008,2029]" country="Philippines" county="Luzon" municipality="Mt. Palali" name="Solano" pageId="7" pageNumber="117" stateProvince="Nueva Vizcaya">Solano</location>
</materialsCitation>
;
<geoCoordinate id="EE0A618AFFBE5C3DFCA8F82957DC77FD" box="[853,949,2008,2029]" degrees="16" direction="north" minutes="26" orientation="latitude" pageId="7" pageNumber="117" precision="925" value="16.433332">16° 26 N</geoCoordinate>
,
<geoCoordinate id="EE0A618AFFBE5C3DFC3DF829504277FD" box="[960,1067,2008,2029]" degrees="121" direction="east" minutes="13" orientation="longitude" pageId="7" pageNumber="117" precision="925" value="121.21667">121° 13 E</geoCoordinate>
,
<quantity id="4CC6AAA8FFBE5C3DFBC8F82950E877FD" box="[1077,1153,2008,2029]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="117" unit="m" value="1000.0">1000 m</quantity>
; montane rain forest; on lower trunk in semi-exposed situation;
<date id="FF80218DFFBE5C3DFE05F80D56067801" box="[504,623,2044,2065]" pageId="7" pageNumber="117" value="2007-03">March 2007</date>
,
<emphasis id="B94ADB5FFFBE5C3DFD84F80D57F37801" box="[633,922,2044,2065]" italics="true" pageId="7" pageNumber="117">Rivas Plata &amp; Lücking 1194D</emphasis>
(
<typeStatus id="5485B9EFFFBE5C3DFC5BF80D57957801" box="[934,1020,2044,2065]" pageId="7" pageNumber="117" type="holotype">holotype</typeStatus>
F).
</paragraph>
<caption id="DF4157C5FFB15C32FF75F9D9516D7671" ID-DOI="http://doi.org/10.5281/zenodo.13666789" ID-Zenodo-Dep="13666789" httpUri="https://zenodo.org/record/13666789/files/figure.png" pageId="8" pageNumber="118" startId="8.[136,229,1576,1597]" targetBox="[189,1398,190,1551]" targetPageId="8" targetType="figure">
<paragraph id="8B81074DFFB15C32FF75F9D9516D7671" blockId="8.[136,1453,1576,1633]" pageId="8" pageNumber="118">
<emphasis id="B94ADB5FFFB15C32FF75F9D95495762D" bold="true" box="[136,252,1576,1597]" pageId="8" pageNumber="118">FIGURE 5.</emphasis>
AC.
<taxonomicName id="4C3E7CCEFFB15C32FEC5F9D9558D762D" box="[312,484,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="118" phylum="Ascomycota" rank="species" species="afzelii">
<emphasis id="B94ADB5FFFB15C32FEC5F9D9558D762D" box="[312,484,1576,1597]" italics="true" pageId="8" pageNumber="118">Dyplolabia afzelii</emphasis>
</taxonomicName>
(A, C, Brazil,
<emphasis id="B94ADB5FFFB15C32FD91F9D9568A762D" box="[620,739,1576,1597]" italics="true" pageId="8" pageNumber="118">Cáceres 245</emphasis>
; B, Thailand,
<emphasis id="B94ADB5FFFB15C32FC94F9D95059762D" box="[873,1072,1576,1597]" italics="true" pageId="8" pageNumber="118">Sutjaritturakan 2931</emphasis>
; thallus with ascomata and ascospores). DF.
<taxonomicName id="4C3E7CCEFFB15C32FF40F9BD55257671" box="[189,332,1612,1633]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="8" pageNumber="118" phylum="Ascomycota" rank="species" species="ochrocheila">
<emphasis id="B94ADB5FFFB15C32FF40F9BD55257671" box="[189,332,1612,1633]" italics="true" pageId="8" pageNumber="118">D. ochrocheila</emphasis>
</taxonomicName>
(A, holotype; B, C, Sri Lanka,
<emphasis id="B94ADB5FFFB15C32FD87F9BD56927671" box="[634,763,1612,1633]" italics="true" pageId="8" pageNumber="118">Thwaites 150</emphasis>
; thallus with ascomata). Scale = 1 mm, in C = 10 μm.
</paragraph>
</caption>
<paragraph id="8B81074DFFB15C32FF75F960505977DA" blockId="8.[136,1452,1681,2030]" pageId="8" pageNumber="118">
Thallus corticolous, epiperidermal, yellow-olive, smooth to uneven, with dense, prosoplectenchymatous cortex; photobiont layer with scattered clusters of calcium oxalate crystals.Apothecia erumpent, angular,
<quantity id="4CC6AAA8FFB15C32FB40F944514776DE" box="[1213,1326,1717,1742]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="118" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
diam.; disc immersed, thickly white-pruinose, partly covered by margin; proper margin distinct, thick, erect, fissured, black but with a very thick, white cover; thalline margin absent. Excipulum laterally carbonized, jet-black, without discernable structure. Periphysoids absent but hymenium laterally with a broad zone of widely spaced, anastomosing paraphyses embedded in a gelatinous matrix. Pseudocolumella sometimes present, plug-shaped. Hymenium 170200 μm high; paraphyses unbranched except for lateral areas as described above. Ascospores 8/ascus, uniseriate, muriform with (3)5 transverse and 13 longitudinal septa per segment, 2530 × 1520 μm, broadly oval to subglobose, with thick septa and diamond-shaped lumina (
<taxonomicName id="4C3E7CCEFFB15C32FDEBF84056CF77DA" box="[534,678,1969,1994]" class="Dothideomycetes" family="Trypetheliaceae" genus="Astrothelium" kingdom="Fungi" order="Trypetheliales" pageId="8" pageNumber="118" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB15C32FDEBF84056CF77DA" box="[534,678,1969,1994]" italics="true" pageId="8" pageNumber="118">Astrothelium</emphasis>
</taxonomicName>
-
<typeStatus id="5485B9EFFFB15C32FD53F840568A77DA" box="[686,739,1969,1994]" pageId="8" pageNumber="118">type</typeStatus>
), colorless, I (non-amyloid).
</paragraph>
<paragraph id="8B81074DFFB15C32FF40F824502177FE" blockId="8.[136,1452,1681,2030]" box="[189,1096,2005,2030]" pageId="8" pageNumber="118">
<emphasis id="B94ADB5FFFB15C32FF40F82455AB77FE" bold="true" box="[189,450,2005,2030]" pageId="8" pageNumber="118">Secondary chemistry:</emphasis>
—White cover of ascoma margin C+ red (lecanoric acid).
</paragraph>
</subSubSection>
<caption id="DF4157C5FFB05C33FF75F9D955E87695" ID-DOI="http://doi.org/10.5281/zenodo.13666795" ID-Zenodo-Dep="13666795" httpUri="https://zenodo.org/record/13666795/files/figure.png" pageId="9" pageNumber="119" startId="9.[136,229,1576,1597]" targetBox="[189,1398,190,1551]" targetPageId="9" targetType="figure">
<paragraph id="8B81074DFFB05C33FF75F9D955E87695" blockId="9.[136,1452,1576,1669]" pageId="9" pageNumber="119">
<emphasis id="B94ADB5FFFB05C33FF75F9D95496762D" bold="true" box="[136,255,1576,1597]" pageId="9" pageNumber="119">FIGURE 6.</emphasis>
A.
<taxonomicName id="4C3E7CCEFFB05C33FED9F9D9566E762D" box="[292,519,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="species" species="dalywaiana">
<emphasis id="B94ADB5FFFB05C33FED9F9D9566E762D" box="[292,519,1576,1597]" italics="true" pageId="9" pageNumber="119">Dyplolabia dalywaiana</emphasis>
</taxonomicName>
(holotype; thallus with ascomata). BD.
<taxonomicName id="4C3E7CCEFFB05C33FC61F9D9507F762D" box="[924,1046,1576,1597]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="species" species="oryzoides">
<emphasis id="B94ADB5FFFB05C33FC61F9D9507F762D" box="[924,1046,1576,1597]" italics="true" pageId="9" pageNumber="119">D. oryzoides</emphasis>
</taxonomicName>
(B, Brazil,
<emphasis id="B94ADB5FFFB05C33FB76F9D95092762D" box="[1163,1275,1576,1597]" italics="true" pageId="9" pageNumber="119">Kalb 31120</emphasis>
; C, D, Sri Lanka,
<emphasis id="B94ADB5FFFB05C33FF75F9BD55757671" box="[136,284,1612,1633]" italics="true" pageId="9" pageNumber="119">Weerakoon 116</emphasis>
; thallus with ascomata and ascospores). EF.
<taxonomicName id="4C3E7CCEFFB05C33FD24F9BD57E27671" box="[729,907,1612,1633]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="species" species="chumphonensis">
<emphasis id="B94ADB5FFFB05C33FD24F9BD57E27671" box="[729,907,1612,1633]" italics="true" pageId="9" pageNumber="119">D. chumphonensis</emphasis>
</taxonomicName>
(holotype; thallus with ascomata and ascospores). Scale = 1 mm, in D, F = 10 μm.
</paragraph>
</caption>
<subSubSection id="C32454C6FFB05C33FF40F94456F17706" pageId="9" pageNumber="119" type="etymology">
<paragraph id="8B81074DFFB05C33FF40F94456F17706" blockId="9.[136,1452,1717,2066]" pageId="9" pageNumber="119">
<emphasis id="B94ADB5FFFB05C33FF40F944552176DE" bold="true" box="[189,328,1717,1742]" pageId="9" pageNumber="119">Etymology:</emphasis>
―This new species is dedicated to the legacy of Charles R. Darwin and his contemporaneous colleagues, Charles Lyell and Afred Russell Wallace, bringing forward the theory of evolution by natural selection and popularizing the use of the term missing link.
</paragraph>
</subSubSection>
<subSubSection id="C32454C6FFB05C33FF40F8D057FC774E" pageId="9" pageNumber="119" type="distribution">
<paragraph id="8B81074DFFB05C33FF40F8D057FC774E" blockId="9.[136,1452,1717,2066]" pageId="9" pageNumber="119">
<emphasis id="B94ADB5FFFB05C33FF40F8D0559D772A" bold="true" box="[189,500,1825,1850]" pageId="9" pageNumber="119">Distribution and ecology:</emphasis>
<collectingCountry id="F32947DDFFB05C33FDEDF8D056FB772A" box="[528,658,1825,1850]" name="Philippines" pageId="9" pageNumber="119">Philippines</collectingCountry>
; thus far only known a single collection from the
<typeStatus id="5485B9EFFFB05C33FB18F8D0517C772A" box="[1253,1301,1825,1850]" pageId="9" pageNumber="119">type</typeStatus>
locality, Mt. Palali, growing on bark of trees in the montane rain forest understory.
</paragraph>
</subSubSection>
<subSubSection id="C32454C6FFB05C30FF40F89855847135" lastPageId="10" lastPageNumber="120" pageId="9" pageNumber="119" type="discussion">
<paragraph id="8B81074DFFB05C30FF40F89855847135" blockId="9.[136,1452,1717,2066]" lastBlockId="10.[136,1452,160,293]" lastPageId="10" lastPageNumber="120" pageId="9" pageNumber="119">
<emphasis id="B94ADB5FFFB05C33FF40F898555D7792" bold="true" box="[189,308,1897,1922]" pageId="9" pageNumber="119">Remarks:</emphasis>
<taxonomicName id="4C3E7CCEFFB05C33FEADF89856347792" box="[336,605,1897,1922]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="species" species="dalywaiana">
<emphasis id="B94ADB5FFFB05C33FEADF89856347792" box="[336,605,1897,1922]" italics="true" pageId="9" pageNumber="119">Dyplolabia dalywaiana</emphasis>
</taxonomicName>
is a very peculiar species. It was first believed to represent a member of
<taxonomicName id="4C3E7CCEFFB05C33FF56F87C554577B6" box="[171,300,1933,1958]" class="Lecanoromycetes" family="Graphidaceae" genus="Ocellularia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="genus" sensu="lato">
<emphasis id="B94ADB5FFFB05C33FF56F87C554577B6" box="[171,300,1933,1958]" italics="true" pageId="9" pageNumber="119">Ocellularia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2796624FFB05C33FECAF87C550777B6" box="[311,366,1933,1958]" pageId="9" pageNumber="119" sensu="lato">s.lat.</taxonomicNameLabel>
, similar to the genera
<taxonomicName id="4C3E7CCEFFB05C33FD80F87C577377B6" box="[637,794,1933,1958]" class="Lecanoromycetes" family="Graphidaceae" genus="Rhabdodiscus" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB05C33FD80F87C577377B6" box="[637,794,1933,1958]" italics="true" pageId="9" pageNumber="119">Rhabdodiscus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E7CCEFFB05C33FCA7F87C57BC77B6" box="[858,981,1933,1958]" class="Lecanoromycetes" family="Graphidaceae" genus="Stegobolus" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB05C33FCA7F87C57BC77B6" box="[858,981,1933,1958]" italics="true" pageId="9" pageNumber="119">Stegobolus</emphasis>
</taxonomicName>
because of the presence of an irregular columella. The close phylogenetic relationship with
<taxonomicName id="4C3E7CCEFFB05C33FD1AF840573877DA" box="[743,849,1969,1994]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="9" pageNumber="119" phylum="Ascomycota" rank="species" species="afzelii">
<emphasis id="B94ADB5FFFB05C33FD1AF840573877DA" box="[743,849,1969,1994]" italics="true" pageId="9" pageNumber="119">D. afzelii</emphasis>
</taxonomicName>
initially came as a surprise, but closer examination of the material then revealed that this species shares important characters with the latter, namely the ascoma margin forming a carbonized base with a thick white, C+ red cover containing lecanoric acid, and the I-negative, astrothelioid ascospores. Morphologically,
<taxonomicName id="4C3E7CCEFFB35C30FE2BFF51561370A9" box="[470,634,160,185]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="10" pageNumber="120" phylum="Ascomycota" rank="species" species="dalywaiana">
<emphasis id="B94ADB5FFFB35C30FE2BFF51561370A9" box="[470,634,160,185]" italics="true" pageId="10" pageNumber="120">D. dalywaiana</emphasis>
</taxonomicName>
is transitional between
<taxonomicName id="4C3E7CCEFFB35C30FC7DFF51579770A9" box="[896,1022,160,185]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="10" pageNumber="120" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB35C30FC7DFF51579770A9" box="[896,1022,160,185]" italics="true" pageId="10" pageNumber="120">Dyplolabia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3E7CCEFFB35C30FBCEFF5150A670A9" authorityName="Rivas Plata, Papong, Lumbsch &amp; Lücking" authorityYear="2012" box="[1075,1231,160,185]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="10" pageNumber="120" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB35C30FBCEFF5150A670A9" box="[1075,1231,160,185]" italics="true" pageId="10" pageNumber="120">Cruentotrema</emphasis>
</taxonomicName>
and hence provides an important taxon to explain the close relationship between both genera: its ascomata have a partially exposed disc and lobulate margins, as in
<taxonomicName id="4C3E7CCEFFB35C30FE46FF19563E7111" authorityName="Rivas Plata, Papong, Lumbsch &amp; Lücking" authorityYear="2012" box="[443,599,232,257]" class="Lecanoromycetes" family="Graphidaceae" genus="Cruentotrema" kingdom="Fungi" order="Ostropales" pageId="10" pageNumber="120" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB35C30FE46FF19563E7111" box="[443,599,232,257]" italics="true" pageId="10" pageNumber="120">Cruentotrema</emphasis>
</taxonomicName>
, whereas the internal anatomy and the thick white cover with lecanoric acid place the species in
<taxonomicName id="4C3E7CCEFFB35C30FE95FEFD558F7135" box="[360,486,268,293]" class="Lecanoromycetes" family="Graphidaceae" genus="Dyplolabia" kingdom="Fungi" order="Ostropales" pageId="10" pageNumber="120" phylum="Ascomycota" rank="genus">
<emphasis id="B94ADB5FFFB35C30FE95FEFD558F7135" box="[360,486,268,293]" italics="true" pageId="10" pageNumber="120">Dyplolabia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>