treatments-xml/data/03/94/87/039487CEF011FFA9FF5DFAD8FC47F841.xml
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<document id="941818B501E5425AF37B994FF7B52ADA" ID-CLB-Dataset="7950" ID-DOI="10.11646/zootaxa.4878.3.3" ID-GBIF-Dataset="b9c711ba-146d-447b-ac2d-e1a0fcce0053" ID-ISSN="1175-5326" ID-Zenodo-Dep="4425178" ID-ZooBank="B660B3B2-7D4A-44C4-92C9-4BE24A1EA950" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610040207257" checkinUser="plazi" docAuthor="Sirenko, Boris I." docDate="2020" docId="039487CEF011FFA9FF5DFAD8FC47F841" docLanguage="en" docName="zootaxa.4878.3.3.pdf" docOrigin="Zootaxa 4878 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptochiton costatoacus Sirenko 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="487" masterDocId="FFADFFB6F003FFBDFFCAFF86FFD6FF91" masterDocTitle="New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)" masterLastPageNumber="500" masterPageNumber="467" pageNumber="485" updateTime="1698923802988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0F74E0FE624296458DF6339B9AB1ED67">New shelf and bathyal Philippine leptochitons (Mollusca: Polyplacophora Lepidopleurida)</mods:title>
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<mods:namePart id="74511C8704553E6865FA7226347A68BA">Sirenko, Boris I.</mods:namePart>
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<mods:date id="F1F01718954E18F0D2DE799C46C0265A">2020</mods:date>
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<treatment id="039487CEF011FFA9FF5DFAD8FC47F841" ID-DOI="http://doi.org/10.5281/zenodo.4426188" ID-GBIF-Taxon="180324283" ID-Zenodo-Dep="4426188" LSID="urn:lsid:plazi:treatment:039487CEF011FFA9FF5DFAD8FC47F841" httpUri="http://treatment.plazi.org/id/039487CEF011FFA9FF5DFAD8FC47F841" lastPageId="20" lastPageNumber="487" pageId="18" pageNumber="485">
<subSubSection id="C3276553F011FFAFFF5DFAD8FDD3FAE8" box="[151,517,1374,1401]" pageId="18" pageNumber="485" type="nomenclature">
<paragraph id="8B8236D8F011FFAFFF5DFAD8FDD3FAE8" blockId="18.[151,517,1374,1437]" box="[151,517,1374,1401]" pageId="18" pageNumber="485">
<heading id="D0CA81B4F011FFAFFF5DFAD8FDD3FAE8" bold="true" box="[151,517,1374,1401]" fontSize="11" level="1" pageId="18" pageNumber="485" reason="1">
<emphasis id="B949EACAF011FFAFFF5DFAD8FDD3FAE8" bold="true" box="[151,517,1374,1401]" pageId="18" pageNumber="485">
<taxonomicName id="4C3D4D5BF011FFAFFF5DFAD8FE6AFAE8" ID-CoL="4feb86f2-136f-438a-a2d9-a4830297135e" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[151,444,1374,1401]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="18" pageNumber="485" phylum="Mollusca" rank="species" species="costatoacus" status="sp. nov.">
<emphasis id="B949EACAF011FFAFFF5DFAD8FE6AFAE8" bold="true" box="[151,444,1374,1401]" italics="true" pageId="18" pageNumber="485">Leptochiton costatoacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B1F011FFAFFE09FAD9FDD3FAE8" box="[451,517,1375,1401]" pageId="18" pageNumber="485" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3276553F011FFAFFF5DFA05FEF8FA0C" box="[151,302,1411,1437]" pageId="18" pageNumber="485" type="description">
<paragraph id="8B8236D8F011FFAFFF5DFA05FEF8FA0C" blockId="18.[151,517,1374,1437]" box="[151,302,1411,1437]" pageId="18" pageNumber="485">
(
<figureCitation id="13062A5DF011FFAFFF55FA05FEF0FA0C" box="[159,294,1411,1437]" captionStart-0="FIGURE 21" captionStart-1="FIGURE 22" captionStart-2="FIGURE 23" captionStartId-0="19.[151,250,1916,1940]" captionStartId-1="20.[151,250,1291,1315]" captionStartId-2="21.[151,250,1225,1249]" captionTargetBox-0="[204,1381,403,1906]" captionTargetBox-1="[168,1416,186,1262]" captionTargetBox-2="[210,1417,160,1199]" captionTargetId-0="figure-165@19.[197,1386,398,1910]" captionTargetId-1="figure-21@20.[164,1423,181,1268]" captionTargetId-2="figure-291@21.[202,1426,151,1209]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionText-0="FIGURE 21. Leptochiton costatoacus n. sp., Philippines, holotype (MNHN-IM-2007-35506), BL 5.5 mm: A. Valve I, dorsal view; B. Valve III, dorsal view; C. Valve IV, dorsal view; D. Valve VIII, dorsal view; E. Valve IV, detail of tegmentum in central and lateral areas; F. Valve III, rostral view; G. Valve VIII, lateral view; H. valve V, ventral view." captionText-1="FIGURE 22. Leptochiton costatoacus n. sp., Philippines, holotype (MNHN-IM-2007-35506), BL 5.5 mm: A. Dorsal and ventral scales and marginal needles; B. Radula; C. Sutural scales and needles; D. Central and first lateral teeth of radula." captionText-2="FIGURE 23. Leptochiton costatoacus n. sp., Philippines, holotype (MNHN-IM-2007-35506), BL 5.5 mm: A. Sutural needle; B. Dorsal scale; C. Marginal needle; D. Ventral scale near margin; E. Ventral scale from middle hyponotum; F. Heads of major lateral teeth of radula; G. Central and first lateral teeth of radula; H. Aesthete group. Scale bars 100 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4425224" figureDoi-1="http://doi.org/10.5281/zenodo.4425226" figureDoi-2="http://doi.org/10.5281/zenodo.4425228" httpUri-0="https://zenodo.org/record/4425224/files/figure.png" httpUri-1="https://zenodo.org/record/4425226/files/figure.png" httpUri-2="https://zenodo.org/record/4425228/files/figure.png" pageId="18" pageNumber="485">Figs. 2123</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFAFFF5DFA4DFABCF9BC" pageId="18" pageNumber="485" type="materials_examined">
<paragraph id="8B8236D8F011FFAFFF5DFA4DFB57F998" blockId="18.[151,1437,1482,2013]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF5DFA4DFEEAFA75" bold="true" box="[151,316,1482,1509]" pageId="18" pageNumber="485">Type material.</emphasis>
<materialsCitation id="3B553C85F011FFAFFE8BFA4DFB57F998" ID-GBIF-Occurrence="3044017315" collectionCode="III, IV, V, VIII" pageId="18" pageNumber="485" specimenCount="1" typeStatus="holotype">
<typeStatus id="5486887AF011FFAFFE8BFA4DFE72FA74" box="[321,420,1483,1509]" pageId="18" pageNumber="485" type="holotype">Holotype</typeStatus>
(MNHN-IM-2007-35506) now disarticulated consisting of SEM stub of valves I,
<collectionCode id="ED2CAE1DF011FFAFFAC8FA4DFAC8FA74" box="[1282,1310,1483,1509]" pageId="18" pageNumber="485">III</collectionCode>
,
<collectionCode id="ED2CAE1DF011FFAFFAE2FA4DFA95FA74" box="[1320,1347,1483,1509]" pageId="18" pageNumber="485">IV</collectionCode>
,
<collectionCode id="ED2CAE1DF011FFAFFA86FA4DFA88FA74" box="[1356,1374,1483,1509]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="485" type="Museum">V</collectionCode>
,
<collectionCode id="ED2CAE1DF011FFAFFAA2FA4DFA4EFA74" box="[1384,1432,1483,1509]" pageId="18" pageNumber="485">VIII</collectionCode>
, part of perinotum and radula, mount of part of perinotum and radula and vial with other valves.
</materialsCitation>
</paragraph>
<paragraph id="8B8236D8F011FFAFFF0DF995FABCF9BC" blockId="18.[151,1437,1482,2013]" box="[199,1386,1554,1581]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF995FE8BF9BD" bold="true" box="[199,349,1554,1581]" pageId="18" pageNumber="485">Type locality.</emphasis>
<materialsCitation id="3B553C85F011FFAFFEA8F995FAB0F9BC" ID-GBIF-Occurrence="3044017327" box="[354,1382,1554,1581]" country="Philippines" elevation="107" latitude="8.728333" location="Bohol" longLatPrecision="129" longitude="123.23333" pageId="18" pageNumber="485" specimenCount="1" stateProvince="Bohol">
<collectingCountry id="F32A7648F011FFAFFEA8F995FE0DF9BC" box="[354,475,1555,1581]" name="Philippines" pageId="18" pageNumber="485">Philippines</collectingCountry>
,
<collectingRegion id="49F9F83AF011FFAFFE2FF995FDF0F9BC" box="[485,550,1555,1581]" country="Philippines" name="Bohol" pageId="18" pageNumber="485">Bohol</collectingRegion>
/
<collectingRegion id="49F9F83AF011FFAFFDE4F995FD88F9BC" box="[558,606,1555,1581]" country="Philippines" name="Sulu" pageId="18" pageNumber="485">Sulu</collectingRegion>
seas,
<geoCoordinate id="EE09501FF011FFAFFD56F995FCD6F9BC" box="[668,768,1555,1581]" degrees="8" direction="north" minutes="43.7" orientation="latitude" pageId="18" pageNumber="485" precision="92" value="8.728333">8°43.7N</geoCoordinate>
,
<geoCoordinate id="EE09501FF011FFAFFCC1F995FC50F9BC" box="[779,902,1555,1581]" degrees="123" direction="east" minutes="14.0" orientation="longitude" pageId="18" pageNumber="485" precision="92" value="123.23333">123°14.0E</geoCoordinate>
,
<quantity id="4CC59B3DF011FFAFFC5BF994FBDFF9BC" box="[913,1033,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.09" metricValueMin="1.05" pageId="18" pageNumber="485" unit="m" value="107.0" valueMax="109.0" valueMin="105.0">
<elevation id="0010D1EBF011FFAFFC5BF994FBDFF9BC" box="[913,1033,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.09" metricValueMin="1.05" pageId="18" pageNumber="485" unit="m" value="107.0" valueMax="109.0" valueMin="105.0">105109 m</elevation>
</quantity>
(PANGLAO 2005, stn CP2374)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFAFFF0DF9B1FEF8F9E4" pageId="18" pageNumber="485" type="etymology">
<paragraph id="8B8236D8F011FFAFFF0DF9B1FEF8F9E4" blockId="18.[151,1437,1482,2013]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF9B1FE9EF9C0" bold="true" box="[199,328,1591,1617]" pageId="18" pageNumber="485">Etymology.</emphasis>
From the Latin costatus, ribbed, and acus, needle, referring to the armature of the girdle in this species, which has ribbed needles.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFAFFF0DF9F8FD2FF908" box="[199,761,1662,1689]" pageId="18" pageNumber="485" type="materials_examined">
<paragraph id="8B8236D8F011FFAFFF0DF9F8FD2FF908" blockId="18.[151,1437,1482,2013]" box="[199,761,1662,1689]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF9F8FE73F909" bold="true" box="[199,421,1662,1688]" pageId="18" pageNumber="485">Material examined.</emphasis>
Known only from the
<typeStatus id="5486887AF011FFAFFD5FF9F9FD23F908" box="[661,757,1663,1689]" pageId="18" pageNumber="485" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFAFFF0DF924FD7CF92C" box="[199,682,1698,1725]" pageId="18" pageNumber="485" type="distribution">
<paragraph id="8B8236D8F011FFAFFF0DF924FD7CF92C" blockId="18.[151,1437,1482,2013]" box="[199,682,1698,1725]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF924FE8FF92D" bold="true" box="[199,345,1698,1724]" pageId="18" pageNumber="485">Distribution.</emphasis>
Only known from
<typeStatus id="5486887AF011FFAFFDE8F925FD99F92C" box="[546,591,1699,1725]" pageId="18" pageNumber="485">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFAFFF0DF940FDC3F804" pageId="18" pageNumber="485" type="diagnosis">
<paragraph id="8B8236D8F011FFAFFF0DF940FDC3F804" blockId="18.[151,1437,1482,2013]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF940FEEDF971" bold="true" box="[199,315,1734,1760]" pageId="18" pageNumber="485">Diagnosis.</emphasis>
Animal small, BL
<quantity id="4CC59B3DF011FFAFFDCFF940FD8FF970" box="[517,601,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="485" unit="mm" value="5.5">5.5 mm</quantity>
, low elevated, valves rounded, not beaked; tail valve mucro posterior, postmucronal slope slightly concave; tegmentum sculptured with small granules arranged in longitudinal rows in pleural areas of intermediate and tail valves, more or less in radial rows in head valve and quincuncially in other areas, including jugal area; each granule with one megalaesthete and 12 micraesthetes; dorsal scales sculptured with 11 to 12 fine, partly double riblets; marginal and sutural needles with ribs; central teeth of radula wide, major lateral tooth of radula unicuspidate, with very week knob on inner margin of head.
</paragraph>
</subSubSection>
<subSubSection id="C3276553F011FFA9FF0DF818FD6EF945" lastPageId="20" lastPageNumber="487" pageId="18" pageNumber="485" type="description">
<paragraph id="8B8236D8F011FFAFFF0DF818FD97F84C" blockId="18.[151,1437,1482,2013]" pageId="18" pageNumber="485">
<emphasis id="B949EACAF011FFAFFF0DF818FE87F829" bold="true" box="[199,337,1950,1976]" pageId="18" pageNumber="485">Description.</emphasis>
<typeStatus id="5486887AF011FFAFFE9CF819FE6FF828" box="[342,441,1951,1977]" pageId="18" pageNumber="485" type="holotype">Holotype</typeStatus>
(BL
<quantity id="4CC59B3DF011FFAFFE27F818FDE9F828" box="[493,575,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="485" unit="mm" value="5.5">5.5 mm</quantity>
) elongate oval in outline. Shell rounded, low elevated (elevation ratio 0.33), valves not beaked. Tegmentum white in colour.
</paragraph>
<paragraph id="8B8236D8F010FFAEFF0DFF11FE0BFEF4" blockId="19.[151,1437,151,357]" pageId="19" pageNumber="486">Head valve semicircular, tegmentum sculpture with small, roundish granules arranged in more or less radial rows. Intermediate valves rectangular, lateral areas not raised, anterior and posterior margins nearly straight. Pleural areas sculptured with round or oval granules arranged in longitudinal rows, jugal and lateral areas with granules arranged in quincunx. Tail valve narrower than head valve, mucro posterior, postmucronal slope slightly concave. Antemucronal area sculptured like central area of intermediate valves, postmucronal area sculptured like lateral areas. Each granule with one megalaesthete and 1012 micraesthetes.</paragraph>
<caption id="DF426650F010FFAEFF5DF8FAFBB4F84D" ID-DOI="http://doi.org/10.5281/zenodo.4425224" ID-Zenodo-Dep="4425224" httpUri="https://zenodo.org/record/4425224/files/figure.png" pageId="19" pageNumber="486" startId="19.[151,250,1916,1940]" targetBox="[204,1381,403,1906]" targetPageId="19">
<paragraph id="8B8236D8F010FFAEFF5DF8FAFBB4F84D" blockId="19.[151,1436,1916,2013]" pageId="19" pageNumber="486">
<emphasis id="B949EACAF010FFAEFF5DF8FAFEF7F805" bold="true" box="[151,289,1916,1940]" pageId="19" pageNumber="486">FIGURE 21.</emphasis>
<taxonomicName id="4C3D4D5BF010FFAEFEE2F8FAFDF7F805" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[296,545,1916,1940]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="19" pageNumber="486" phylum="Mollusca" rank="species" species="costatoacus" status="sp. nov.">
<emphasis id="B949EACAF010FFAEFEE2F8FAFDF7F805" box="[296,545,1916,1940]" italics="true" pageId="19" pageNumber="486">Leptochiton costatoacus</emphasis>
</taxonomicName>
<emphasis id="B949EACAF010FFAEFDE2F8FBFDB7F805" bold="true" box="[552,609,1917,1941]" pageId="19" pageNumber="486">
<taxonomicNameLabel id="A27A57B1F010FFAEFDE2F8FBFDB7F805" box="[552,609,1917,1941]" pageId="19" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2007-35506), BL 5.5 mm: A. Valve I, dorsal view; B. Valve III, dorsal view; C. Valve IV, dorsal view; D. Valve VIII, dorsal view; E. Valve IV, detail of tegmentum in central and lateral areas; F. Valve III, rostral view; G. Valve VIII, lateral view; H. valve V, ventral view.
</paragraph>
</caption>
<caption id="DF426650F017FFA9FF5DFA8DFAE0FAD9" ID-DOI="http://doi.org/10.5281/zenodo.4425226" ID-Zenodo-Dep="4425226" httpUri="https://zenodo.org/record/4425226/files/figure.png" pageId="20" pageNumber="487" startId="20.[151,250,1291,1315]" targetBox="[168,1416,186,1262]" targetPageId="20">
<paragraph id="8B8236D8F017FFA9FF5DFA8DFAE0FAD9" blockId="20.[151,1436,1291,1352]" pageId="20" pageNumber="487">
<emphasis id="B949EACAF017FFA9FF5DFA8DFEF6FAB5" bold="true" box="[151,288,1291,1316]" pageId="20" pageNumber="487">FIGURE 22.</emphasis>
<taxonomicName id="4C3D4D5BF017FFA9FEECFA8AFDCBFAB2" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[294,541,1292,1316]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="costatoacus" status="sp. nov.">
<emphasis id="B949EACAF017FFA9FEECFA8AFDCBFAB2" box="[294,541,1292,1316]" italics="true" pageId="20" pageNumber="487">Leptochiton costatoacus</emphasis>
</taxonomicName>
<emphasis id="B949EACAF017FFA9FDE9FA8AFD8AFAB2" bold="true" box="[547,604,1292,1316]" pageId="20" pageNumber="487">
<taxonomicNameLabel id="A27A57B1F017FFA9FDE9FA8AFD8AFAB2" box="[547,604,1292,1316]" pageId="20" pageNumber="487" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, Philippines, holotype (MNHN-IM-2007-35506), BL 5.5 mm: A. Dorsal and ventral scales and marginal needles; B. Radula; C. Sutural scales and needles; D. Central and first lateral teeth of radula.
</paragraph>
</caption>
<paragraph id="8B8236D8F017FFA9FF0DFAF0FE88FA25" blockId="20.[151,1437,1398,2000]" pageId="20" pageNumber="487">Articulamentum moderately developed, apophyses small, widely separated, subtriangular in valves IIIII, more or less trapezoidal in tail valve.</paragraph>
<paragraph id="8B8236D8F017FFA9FF0DFA38FAFBF9D5" blockId="20.[151,1437,1398,2000]" pageId="20" pageNumber="487">
Girdle very narrow, dorsally densely beset with elongate, slightly curved, pointed scales, sculptured with eleven to twelve fine riblets (60 x 43 μm), long sutural scales up to 110 μm, with 9 riblets, sutural needles long and ribbed, (250 x 20 μm), ribbed marginal needles (240 x
<quantity id="4CC59B3DF017FFA9FD62F980FD3AF9B0" box="[680,748,1542,1569]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="487" unit="mm" value="25.0">25µm</quantity>
) same size but wider. Two rows of ventral scales near margin (71 x 30 μm) with two short ribs in upper half, other ventral scales lanceolate, flat, smooth (90 x 23 μm).
</paragraph>
<paragraph id="8B8236D8F017FFA9FF0DF9C8FC01F9F9" blockId="20.[151,1437,1398,2000]" box="[199,983,1614,1640]" pageId="20" pageNumber="487">
<materialsCitation id="3B553C85F017FFA9FF0DF9C8FC01F9F9" ID-GBIF-Occurrence="3044017324" box="[199,983,1614,1640]" collectionCode="VI" pageId="20" pageNumber="487" specimenCount="1" typeStatus="holotype">
<typeStatus id="5486887AF017FFA9FF0DF9C8FEFCF9F9" box="[199,298,1614,1640]" pageId="20" pageNumber="487" type="holotype">Holotype</typeStatus>
with ten gills on each side extending from valve
<collectionCode id="ED2CAE1DF017FFA9FCE4F9C8FC9CF9F9" box="[814,842,1614,1640]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="20" pageNumber="487">VI</collectionCode>
to near anus.
</materialsCitation>
</paragraph>
<paragraph id="8B8236D8F017FFA9FF0DF9F4FB52F921" blockId="20.[151,1437,1398,2000]" pageId="20" pageNumber="487">
Radula of the
<typeStatus id="5486887AF017FFA9FE92F9F4FE62F91D" box="[344,436,1650,1676]" pageId="20" pageNumber="487" type="holotype">holotype</typeStatus>
<quantity id="4CC59B3DF017FFA9FE70F9F4FDDCF91C" box="[442,522,1650,1677]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="20" pageNumber="487" unit="mm" value="2.8">2.8 mm</quantity>
long with 26 transverse rows of mature teeth. Central teeth wide with trapezoidal blade. Major lateral teeth with sharply pointed, unidentate head with very weak knob on inner margin.
</paragraph>
<paragraph id="8B8236D8F017FFA9FF0DF93CFD6EF945" blockId="20.[151,1437,1398,2000]" box="[199,696,1722,1748]" pageId="20" pageNumber="487">Gut containing intact foraminifera and detritus.</paragraph>
</subSubSection>
<subSubSection id="C3276553F017FFA9FF0DF958FC47F841" pageId="20" pageNumber="487" type="discussion">
<paragraph id="8B8236D8F017FFA9FF0DF958FC47F841" blockId="20.[151,1437,1398,2000]" pageId="20" pageNumber="487">
<emphasis id="B949EACAF017FFA9FF0DF958FEE0F969" bold="true" box="[199,310,1758,1784]" pageId="20" pageNumber="487">Remarks.</emphasis>
In
<taxonomicName id="4C3D4D5BF017FFA9FE91F958FD8BF969" authority="Sirenko, 2020" authorityName="Sirenko" authorityYear="2020" box="[347,605,1758,1784]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="costatoacus" status="sp. nov.">
<emphasis id="B949EACAF017FFA9FE91F958FD8BF969" box="[347,605,1758,1784]" italics="true" pageId="20" pageNumber="487">Leptochiton costatoacus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57B1F017FFA9FDAFF958FD49F969" box="[613,671,1758,1784]" pageId="20" pageNumber="487" rank="species">n. sp.</taxonomicNameLabel>
, the granules are located in longitudinal rows in the central areas of the intermediate valves and in the antemucronal area of the tail valve. Regarding the rest of the features of girdle, radula and other areas of the shell (jugal area, postmucronal area of the tail valve and the most part of head valve) the new species is quite close to species of the
<taxonomicName id="4C3D4D5BF017FFA9FE17F8CDFD83F8F5" authorityName="Dall" authorityYear="1878" box="[477,597,1866,1892]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="belknapi">
<emphasis id="B949EACAF017FFA9FE17F8CDFD83F8F5" box="[477,597,1866,1892]" italics="true" pageId="20" pageNumber="487">L. belknapi</emphasis>
</taxonomicName>
group. Of all the species of this group the new species most closely resembles
<taxonomicName id="4C3D4D5BF017FFA9FF5DF8E8FDDAF819" authority="Sirenko, 2016" authorityName="Sirenko" authorityYear="2016" box="[151,524,1902,1928]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="fijiensis">
<emphasis id="B949EACAF017FFA9FF5DF8E8FEB8F819" box="[151,366,1902,1928]" italics="true" pageId="20" pageNumber="487">Leptochiton fijiensis</emphasis>
Sirenko, 2016
</taxonomicName>
, but differs by having a posterior mucro (vs. anterior mucro in
<taxonomicName id="4C3D4D5BF017FFA9FB63F8E9FAC0F819" authorityName="Sirenko" authorityYear="2016" box="[1193,1302,1902,1928]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="fijiensis">
<emphasis id="B949EACAF017FFA9FB63F8E9FAC0F819" box="[1193,1302,1902,1928]" italics="true" pageId="20" pageNumber="487">L. fijiensis</emphasis>
</taxonomicName>
), 1112 ribs in dorsal scales (vs. 67 ribs in
<taxonomicName id="4C3D4D5BF017FFA9FE15F815FD9DF83D" authorityName="Sirenko" authorityYear="2016" box="[479,587,1938,1964]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="fijiensis">
<emphasis id="B949EACAF017FFA9FE15F815FD9DF83D" box="[479,587,1938,1964]" italics="true" pageId="20" pageNumber="487">L. fijiensis</emphasis>
</taxonomicName>
), ribbed sutural needles (vs. smooth needles in
<taxonomicName id="4C3D4D5BF017FFA9FBF6F815FB7EF83D" authorityName="Sirenko" authorityYear="2016" box="[1084,1192,1938,1964]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="fijiensis">
<emphasis id="B949EACAF017FFA9FBF6F815FB9DF83D" box="[1084,1099,1939,1964]" italics="true" pageId="20" pageNumber="487">L</emphasis>
.
<emphasis id="B949EACAF017FFA9FB9DF814FB7EF83D" box="[1111,1192,1938,1964]" italics="true" pageId="20" pageNumber="487">fijiensis</emphasis>
</taxonomicName>
) and longitudinal rows of granules in pleural areas (granules arrange in quincunx in
<taxonomicName id="4C3D4D5BF017FFA9FCDFF831FC57F841" authorityName="Sirenko" authorityYear="2016" box="[789,897,1974,2000]" class="Polyplacophora" family="Leptochitonidae" genus="Leptochiton" kingdom="Animalia" order="Lepidopleurida" pageId="20" pageNumber="487" phylum="Mollusca" rank="species" species="fijiensis">
<emphasis id="B949EACAF017FFA9FCDFF831FC57F841" box="[789,897,1974,2000]" italics="true" pageId="20" pageNumber="487">L. fijiensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>