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<document id="48147EFDB2C1F52945282B3DEFA53371" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC4BB12FF1881AEFD9CFDE6" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia barclayi Gompel 2020, n. sp." docType="treatment" docUuid="53576D0D-137B-49A7-A58C-C3B7BC8B191B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="154" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="153" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C1E943957436F716D86839EA3B7B3455">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="375A71E0415382F732ED823C26A82781">Gompel, Nicolas</mods:namePart>
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<mods:date id="8DBD0AAFA2EA398B80489F9332AA8387">2020</mods:date>
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<mods:number id="04729675390CB14F52545E4E1CBB2E5B">2020-08-26</mods:number>
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<mods:classification id="473D73E2BA2034B926B9C46BBF777BE5">journal article</mods:classification>
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<mods:identifier id="154E28575DB4D77D379B903210E7A69C" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFC4BB12FF1881AEFD9CFDE6" ID-DOI="http://doi.org/10.5281/zenodo.4404243" ID-GBIF-Taxon="177012530" ID-Zenodo-Dep="4404243" LSID="urn:lsid:zoobank.org:act:53576D0D-137B-49A7-A58C-C3B7BC8B191B" httpUri="http://treatment.plazi.org/id/039187D8FFC4BB12FF1881AEFD9CFDE6" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
<subSubSection id="C3226545FFC4BB13FF1881AEFE29FA01" box="[151,436,1507,1534]" pageId="2" pageNumber="153" type="nomenclature">
<paragraph id="8B8736CEFFC4BB13FF1881AEFE29FA01" blockId="2.[151,436,1507,1571]" box="[151,436,1507,1534]" pageId="2" pageNumber="153">
<heading id="D0CF81A2FFC4BB13FF1881AEFE29FA01" bold="true" box="[151,436,1507,1534]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
<emphasis id="B94CEADCFFC4BB13FF1881AEFE29FA01" bold="true" box="[151,436,1507,1534]" pageId="2" pageNumber="153">
<taxonomicName id="4C384D4DFFC4BB13FF1881AEFEF7FA01" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,362,1507,1534]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="B94CEADCFFC4BB13FF1881AEFEF7FA01" bold="true" box="[151,362,1507,1534]" italics="true" pageId="2" pageNumber="153">Zarcosia barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFC4BB13FEFD81AEFE29FA01" box="[370,436,1508,1534]" pageId="2" pageNumber="153" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC4BB13FF188242FE07F9DC" box="[151,410,1544,1571]" pageId="2" pageNumber="153" type="description">
<paragraph id="8B8736CEFFC4BB13FF188242FE07F9DC" blockId="2.[151,436,1507,1571]" box="[151,410,1544,1571]" pageId="2" pageNumber="153">
(
<figureCitation id="13032A4BFFC4BB13FF108242FE96F9DD" box="[159,267,1544,1570]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="2" pageNumber="153">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC4BB13FE978242FEA2F9DD" box="[280,319,1544,1570]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="2" pageNumber="153">3i,j</figureCitation>
;
<figureCitation id="13032A4BFFC4BB13FEC38242FEF4F9DD" box="[332,361,1544,1570]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="2" pageNumber="153">4a</figureCitation>
;
<figureCitation id="13032A4BFFC4BB13FEFA8242FE0FF9DC" box="[373,402,1544,1571]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="2" pageNumber="153">
<quantity id="4CC09B2BFFC4BB13FEFA8242FE0FF9DC" box="[373,402,1544,1571]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="2" pageNumber="153" unit="g" value="6.0">6g</quantity>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC4BB13FF18821AFB6EF971" pageId="2" pageNumber="153" type="etymology">
<paragraph id="8B8736CEFFC4BB13FF18821AFB6EF971" blockId="2.[151,1437,1616,2039]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF18821AFE83F995" bold="true" box="[151,286,1616,1642]" pageId="2" pageNumber="153">Etymology.</emphasis>
The species is named after Max Barclay in acknowledgment of his unflagging support over two decades to guide me through the insect research collections of the British Museum of Natural History.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC4BB13FF4882D2FABDF94D" box="[199,1312,1687,1715]" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="8B8736CEFFC4BB13FF4882D2FABDF94D" blockId="2.[151,1437,1616,2039]" box="[199,1312,1687,1715]" pageId="2" pageNumber="153">
<materialsCitation id="3B503C93FFC4BB13FF4882D2FA81F94D" ID-GBIF-Occurrence="3020160475" box="[199,1308,1687,1715]" country="Malaysia" location="Gunung Mulu National Park" longitude="114.88333" municipality="Deer Cave" pageId="2" pageNumber="153" specimenCount="1" typeStatus="holotype">
<geoCoordinate id="EE0C5009FFC4BB13FF4882D2FB0FF94C" box="[199,1170,1688,1715]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF4882D2FEFBF94D" bold="true" box="[199,358,1688,1714]" pageId="2" pageNumber="153">
<typeStatus id="5483886CFFC4BB13FF4882D2FE9FF94D" box="[199,258,1688,1714]" pageId="2" pageNumber="153">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC4BB13FEE382D2FE4AF94D" box="[364,471,1688,1714]" name="Malaysia" pageId="2" pageNumber="153">Malaysia</collectingCountry>
: Sarawak,
<location id="8EE76015FFC4BB13FDDF82D2FC0FF94D" LSID="urn:lsid:plazi:treatment:039187D8FFC4BB12FF1881AEFD9CFDE6:8EE76015FFC4BB13FDDF82D2FC0FF94D" box="[592,914,1688,1714]" country="Malaysia" longitude="114.88333" municipality="Deer Cave" name="Gunung Mulu National Park" pageId="2" pageNumber="153">Gunung Mulu National Park</location>
,
<collectingMunicipality id="6BE3ACB4FFC4BB13FC1182D3FB88F94D" box="[926,1045,1688,1715]" pageId="2" pageNumber="153">Deer Cave</collectingMunicipality>
[04°05’’ N
</geoCoordinate>
<geoCoordinate id="EE0C5009FFC4BB13FB1682D2FA89F94D" box="[1177,1300,1687,1715]" degrees="114" direction="east" minutes="53" orientation="longitude" pageId="2" pageNumber="153" precision="925" value="114.88333">114° 53’ E</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC4BB13FF4882F6FDD0F929" box="[199,589,1724,1750]" pageId="2" pageNumber="153" type="distribution">
<paragraph id="8B8736CEFFC4BB13FF4882F6FDD0F929" blockId="2.[151,1437,1616,2039]" box="[199,589,1724,1750]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF4882F6FEFCF929" bold="true" box="[199,353,1724,1750]" pageId="2" pageNumber="153">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC4BB13FEE882F6FE53F929" box="[359,462,1724,1750]" name="Malaysia" pageId="2" pageNumber="153">Malaysia</collectingCountry>
(
<figureCitation id="13032A4BFFC4BB13FE5282F6FDA1F929" box="[477,572,1724,1750]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="2" pageNumber="153">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC4BB12FF4882AAFE51FEBE" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="description">
<paragraph id="8B8736CEFFC4BB13FF4882AAFC31F8E1" blockId="2.[151,1437,1616,2039]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF4882AAFEC4F905" bold="true" box="[199,345,1760,1786]" pageId="2" pageNumber="153">Description.</emphasis>
(
<figureCitation id="13032A4BFFC4BB13FEE982AAFE43F904" box="[358,478,1760,1787]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="2" pageNumber="153">Figure 3i,j</figureCitation>
). Body length.
<specimenCount id="9D3EFD47FFC4BB13FD0882ABFD02F904" box="[647,671,1761,1787]" pageId="2" pageNumber="153" type="male">♂</specimenCount>
: 1.74±
<quantity id="4CC09B2BFFC4BB13FD6782AAFCD0F904" box="[744,845,1760,1787]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="153" unit="mm" value="0.02">0.02 mm</quantity>
(n=2);
<specimenCount id="9D3EFD47FFC4BB13FC1182ABFC28F904" box="[926,949,1761,1787]" pageId="2" pageNumber="153" type="female">♀</specimenCount>
: 1.96±
<quantity id="4CC09B2BFFC4BB13FB8F82AAFBF9F904" box="[1024,1124,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="153" unit="mm" value="0.13">0.13 mm</quantity>
(n=7). Dark brown, densely covered with silver applied pubescence. Appendages orange to brown.
</paragraph>
<paragraph id="8B8736CEFFC4BB13FF488362FE6DF898" blockId="2.[151,1437,1616,2039]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF488362FE99F8BD" box="[199,260,1832,1858]" italics="true" pageId="2" pageNumber="153">Head</emphasis>
. Wider than pronotum in males, as wide as pronotum in females. Palpi orange, last article securiform. Integument mat, finely rugose.
</paragraph>
<paragraph id="8B8736CEFFC4BB13FF48833AFBD0F851" blockId="2.[151,1437,1616,2039]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF48833AFEACF875" box="[199,305,1904,1930]" italics="true" pageId="2" pageNumber="153">Antennae</emphasis>
. A fourth to a third longer than elytra in males, shorter than elytra in females, orange, slightly and progressively thicker toward apex. Antennomere 4 and higher with a slight serration.
</paragraph>
<paragraph id="8B8736CEFFC4BB13FF4883F3FEB4F809" blockId="2.[151,1437,1616,2039]" pageId="2" pageNumber="153">
<emphasis id="B94CEADCFFC4BB13FF4883F3FEABF82D" box="[199,310,1977,2002]" italics="true" pageId="2" pageNumber="153">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFC4BB13FECC83F2FE2EF82C" box="[323,435,1976,2003]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="2" pageNumber="153">
Figure
<quantity id="4CC09B2BFFC4BB13FE1B83F2FE2EF82C" box="[404,435,1976,2003]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="2" pageNumber="153" unit="g" value="6.0">6g</quantity>
</figureCitation>
). Subcylindrical, convex, slightly longer than wide, marked by a strong lateral constriction on each side.
</paragraph>
<paragraph id="8B8736CEFFC5BB12FF4884DDFCB0FF2A" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="154">
<emphasis id="B94CEADCFFC5BB12FF4884DDFE90FF4E" box="[199,269,151,177]" italics="true" pageId="3" pageNumber="154">Elytra</emphasis>
. Length:
<specimenCount id="9D3EFD47FFC5BB12FEF784DDFE0DFF4E" box="[376,400,151,177]" pageId="3" pageNumber="154" type="male">♂</specimenCount>
: 1.20±
<quantity id="4CC09B2BFFC5BB12FE5484DDFDA2FF4E" box="[475,575,151,177]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="3" pageNumber="154" unit="mm" value="0.07">0.07 mm</quantity>
(n=2);
<specimenCount id="9D3EFD47FFC5BB12FD1D84DDFD34FF4E" box="[658,681,151,177]" pageId="3" pageNumber="154" type="female">♀</specimenCount>
: 1.21±
<quantity id="4CC09B2BFFC5BB12FD7C84DDFCC4FF4E" box="[755,857,151,177]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="154" unit="mm" value="0.08">0.08 mm</quantity>
(n=7). Finely and densely punctuated, covered with double pubescence. Moderately convex, depressed on disc.
</paragraph>
<paragraph id="8B8736CEFFC5BB12FF488495FB5EFF06" blockId="3.[151,1437,151,537]" box="[199,1219,223,249]" pageId="3" pageNumber="154">
<emphasis id="B94CEADCFFC5BB12FF488495FF61FF07" box="[199,252,223,248]" italics="true" pageId="3" pageNumber="154">Legs</emphasis>
. No notable sexual characters. Male foretibia with a faint sinuation of their inner side.
</paragraph>
<paragraph id="8B8736CEFFC5BB12FF488549FE51FEBE" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="154">
<emphasis id="B94CEADCFFC5BB12FF488549FEA9FEE2" box="[199,308,259,285]" italics="true" pageId="3" pageNumber="154">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFC5BB12FECB8549FE2BFEE2" box="[324,438,259,285]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="3" pageNumber="154">Figure 4a</figureCitation>
). Simple, narrow, progressively tapered, slightly and regularly bent in lateral view. Appendages visible near apex.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC5BB12FF488501FD9CFDE6" pageId="3" pageNumber="154" type="materials_examined">
<paragraph id="8B8736CEFFC5BB12FF488501FD9CFDE6" blockId="3.[151,1437,151,537]" pageId="3" pageNumber="154">
<emphasis id="B94CEADCFFC5BB12FF488501FEEBFE9B" bold="true" box="[199,374,330,357]" pageId="3" pageNumber="154">Type material.</emphasis>
<materialsCitation id="3B503C93FFC5BB12FEF28501FC50FE76" ID-GBIF-Occurrence="3020160423" collectingDate="1978-05-06" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="360" location="Gunung Mulu National Park" longitude="114.88333" municipality="Deer Cave" pageId="3" pageNumber="154" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="5483886CFFC5BB12FEF28501FE78FE9A" box="[381,485,331,357]" pageId="3" pageNumber="154" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFC5BB12FE788501FD93FE9A" box="[503,526,331,357]" pageId="3" pageNumber="154" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC5BB12FD958501FDEEFE9A" box="[538,627,331,357]" pageId="3" pageNumber="154">BMNH</collectionCode>
).
<emphasis id="B94CEADCFFC5BB12FD098500FD60FE9B" bold="true" box="[646,765,330,356]" pageId="3" pageNumber="154">
<collectingCountry id="F32F765EFFC5BB12FD098500FD6AFE9B" box="[646,759,330,356]" name="Malaysia" pageId="3" pageNumber="154">Malaysia</collectingCountry>
:
</emphasis>
Sarawak,
<location id="8EE76015FFC5BB12FCFB8501FB25FE9A" LSID="urn:lsid:plazi:treatment:039187D8FFC4BB12FF1881AEFD9CFDE6:8EE76015FFC5BB12FCFB8501FB25FE9A" box="[884,1208,331,357]" country="Malaysia" longitude="114.88333" municipality="Deer Cave" name="Gunung Mulu National Park" pageId="3" pageNumber="154">Gunung Mulu National Park</location>
,
<collectingMunicipality id="6BE3ACB4FFC5BB12FB4B8501FAA3FE9A" box="[1220,1342,331,357]" pageId="3" pageNumber="154">Deer Cave</collectingMunicipality>
,
<geoCoordinate id="EE0C5009FFC5BB12FAC58501FF36FE76" pageId="3" pageNumber="154">04°07’’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC5BB12FF3B8525FEB0FE76" box="[180,301,366,393]" degrees="114" direction="east" minutes="53" orientation="longitude" pageId="3" pageNumber="154" precision="925" value="114.88333">114° 53’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFC5BB12FEB28525FE15FE76" box="[317,392,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">
<elevation id="0015D1FDFFC5BB12FEB28525FE15FE76" box="[317,392,367,393]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">360 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC5BB12FE2E8525FDC1FE76" box="[417,604,367,393]" pageId="3" pageNumber="154">P. M. Hammond</collectorName>
&amp;
<collectorName id="26CD5318FFC5BB12FD0C8525FCBFFE76" box="[643,802,367,393]" pageId="3" pageNumber="154">J. E. Marshall</collectorName>
leg.)
<date id="FF86100EFFC5BB12FCEA8525FC50FE76" box="[869,973,367,393]" pageId="3" pageNumber="154" value="1978-05-06">
<collectingDate id="EFC2E9E6FFC5BB12FCEA8525FC50FE76" box="[869,973,367,393]" pageId="3" pageNumber="154" value="1978-05-06">6.V.1978</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B503C93FFC5BB12FC568525FE31FE2E" ID-GBIF-Occurrence="3020160336" collectingDate="1978-05-06" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="360" location="Gunung Mulu National Park" longitude="114.88333" municipality="Deer Cave" pageId="3" pageNumber="154" specimenCount="4" specimenCount-female="3" typeStatus="paratype">
<typeStatus id="5483886CFFC5BB12FC568525FBD7FE76" box="[985,1098,367,393]" pageId="3" pageNumber="154" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B94CEADCFFC5BB12FBDA8524FB5FFE77" bold="true" box="[1109,1218,366,392]" pageId="3" pageNumber="154">
<collectingCountry id="F32F765EFFC5BB12FBDA8524FB5FFE77" box="[1109,1218,366,392]" name="Malaysia" pageId="3" pageNumber="154">Malaysia</collectingCountry>
</emphasis>
: Sarawak,
<location id="8EE76015FFC5BB12FACD8525FEE6FE52" LSID="urn:lsid:plazi:treatment:039187D8FFC4BB12FF1881AEFD9CFDE6:8EE76015FFC5BB12FACD8525FEE6FE52" country="Malaysia" longitude="114.88333" municipality="Deer Cave" name="Gunung Mulu National Park" pageId="3" pageNumber="154">Gunung Mulu National Park</location>
,
<collectingMunicipality id="6BE3ACB4FFC5BB12FE0785D9FD9EFE52" box="[392,515,403,429]" pageId="3" pageNumber="154">Deer Cave</collectingMunicipality>
,
<geoCoordinate id="EE0C5009FFC5BB12FD9F85D9FDE3FE52" box="[528,638,403,429]" pageId="3" pageNumber="154">04°07’’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC5BB12FD0985D9FC9DFE52" box="[646,768,402,429]" degrees="114" direction="east" minutes="53" orientation="longitude" pageId="3" pageNumber="154" precision="925" value="114.88333">114° 53’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFC5BB12FC9F85D9FCC6FE52" box="[784,859,403,429]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">
<elevation id="0015D1FDFFC5BB12FC9F85D9FCC6FE52" box="[784,859,403,429]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">360 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC5BB12FCFB85D9FBB2FE52" box="[884,1071,403,429]" pageId="3" pageNumber="154">P. M. Hammond</collectorName>
&amp;
<collectorName id="26CD5318FFC5BB12FBD985D9FB68FE52" box="[1110,1269,403,429]" pageId="3" pageNumber="154">J. E. Marshall</collectorName>
leg.)
<date id="FF86100EFFC5BB12FAB785D9FA01FE52" box="[1336,1436,403,429]" pageId="3" pageNumber="154" value="1978-05-06">
<collectingDate id="EFC2E9E6FFC5BB12FAB785D9FA01FE52" box="[1336,1436,403,429]" pageId="3" pageNumber="154" value="1978-05-06">6.V.1978</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC5BB12FF2F85FDFF56FE2E" box="[160,203,439,465]" pageId="3" pageNumber="154" type="female">3 ♀</specimenCount>
<typeStatus id="5483886CFFC5BB12FF5D85FDFEDDFE2E" box="[210,320,439,465]" pageId="3" pageNumber="154" type="paratype">paratypes</typeStatus>
,
<collectionCode id="ED29AE0BFFC5BB12FEC485FDFE39FE2E" box="[331,420,439,465]" pageId="3" pageNumber="154">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B503C93FFC5BB12FE3985FDFC56FE0A" ID-GBIF-Occurrence="3020160441" collectingDate="1978-05-13" collectionCode="BMNH" collectorName="P. M. Hammond &amp; J. E. Marshall" country="Malaysia" elevation="360" latitude="4.116667" location="Gunung Mulu National Park" longLatPrecision="1307" longitude="114.916664" municipality="Clearwater Cave" pageId="3" pageNumber="154" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F32F765EFFC5BB12FE3985FDFD84FE2E" box="[438,537,439,465]" name="Malaysia" pageId="3" pageNumber="154">Sarawak</collectingCountry>
,
<location id="8EE76015FFC5BB12FDAA85FDFCFAFE2E" LSID="urn:lsid:plazi:treatment:039187D8FFC4BB12FF1881AEFD9CFDE6:8EE76015FFC5BB12FDAA85FDFCFAFE2E" box="[549,871,439,465]" country="Malaysia" latitude="4.116667" longLatPrecision="1307" longitude="114.916664" municipality="Clearwater Cave" name="Gunung Mulu National Park" pageId="3" pageNumber="154">Gunung Mulu National Park</location>
,
<collectingMunicipality id="6BE3ACB4FFC5BB12FCFD85FDFBADFE2E" box="[882,1072,439,465]" pageId="3" pageNumber="154">Clearwater Cave</collectingMunicipality>
,
<geoCoordinate id="EE0C5009FFC5BB12FBB385FDFB3CFE2E" box="[1084,1185,439,465]" degrees="04" direction="north" minutes="07" orientation="latitude" pageId="3" pageNumber="154" precision="925" value="4.116667">04°07’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC5BB12FB2785FCFA85FE2E" box="[1192,1304,438,465]" degrees="114" direction="east" minutes="55" orientation="longitude" pageId="3" pageNumber="154" precision="925" value="114.916664">114°55’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFC5BB12FAA885FDFAEDFE2E" box="[1319,1392,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">
<elevation id="0015D1FDFFC5BB12FAA885FDFAEDFE2E" box="[1319,1392,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="154" unit="m" value="360.0">360 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC5BB12FA0785FDFEABFE0A" pageId="3" pageNumber="154">P. M. Hammond</collectorName>
&amp;
<collectorName id="26CD5318FFC5BB12FED28591FE60FE0A" box="[349,509,475,501]" pageId="3" pageNumber="154">J. E. Marshall</collectorName>
leg.)
<date id="FF86100EFFC5BB12FDCE8591FD2EFE0A" box="[577,691,475,501]" pageId="3" pageNumber="154" value="1978-05-13">
<collectingDate id="EFC2E9E6FFC5BB12FDCE8591FD2EFE0A" box="[577,691,475,501]" pageId="3" pageNumber="154" value="1978-05-13">13.V.1978</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC5BB12FD488591FD6CFE0A" box="[711,753,475,501]" pageId="3" pageNumber="154" type="female">1 ♀</specimenCount>
<typeStatus id="5483886CFFC5BB12FD758591FCC0FE0A" box="[762,861,475,501]" pageId="3" pageNumber="154" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC5BB12FCE58591FC5FFE0A" box="[874,962,475,501]" pageId="3" pageNumber="154">BMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B503C93FFC5BB12FC588591FE60FDE6" ID-GBIF-Occurrence="3020160381" collectingDate="1978-05-20" collectionCode="BMNH, NGPC" country="Malaysia" pageId="3" pageNumber="154" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<date id="FF86100EFFC5BB12FC588591FBD7FE0A" box="[983,1098,475,501]" pageId="3" pageNumber="154" value="1978-05-20">
<collectingDate id="EFC2E9E6FFC5BB12FC588591FBD7FE0A" box="[983,1098,475,501]" pageId="3" pageNumber="154" value="1978-05-20">20.V.1978</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFC5BB12FBD48591FB15FE0A" box="[1115,1160,475,501]" pageId="3" pageNumber="154" type="female">2 ♀</specimenCount>
<typeStatus id="5483886CFFC5BB12FB1E8591FB62FE0A" box="[1169,1279,475,501]" pageId="3" pageNumber="154" type="paratype">paratypes</typeStatus>
,
<collectionCode id="ED29AE0BFFC5BB12FA838591FAF9FE0A" box="[1292,1380,475,501]" pageId="3" pageNumber="154">BMNH</collectionCode>
;
<specimenCount id="9D3EFD47FFC5BB12FAFF8591FA01FE0A" box="[1392,1436,475,501]" pageId="3" pageNumber="154" type="female">1 ♀</specimenCount>
<typeStatus id="5483886CFFC5BB12FF1885B5FF67FDE6" box="[151,250,511,537]" pageId="3" pageNumber="154" type="paratype">paratype</typeStatus>
,
<specimenCount id="9D3EFD47FFC5BB12FE8985B5FEADFDE6" box="[262,304,511,537]" pageId="3" pageNumber="154" type="male">1 ♂</specimenCount>
<typeStatus id="5483886CFFC5BB12FEB885B5FE07FDE6" box="[311,410,511,537]" pageId="3" pageNumber="154" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC5BB12FE2A85B5FE68FDE6" box="[421,501,511,537]" pageId="3" pageNumber="154">NGPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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