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<document id="BBB65949785870F339EDF5807E2693F8" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC2BB15FF1884DDFAEFFC59" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia bedosae Gompel 2020, n. sp." docType="treatment" docUuid="330A5BB9-B773-4698-B644-5DA3EAC95433" docUuidSource="ZooBank" docVersion="8" lastPageNumber="155" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="155" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="92962664D2ABC0C45100A149C4FB3867">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="B433E18512048BA2709CB7A74BCF9DA3">Gompel, Nicolas</mods:namePart>
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<mods:date id="ACFF56816DD2EA742435938E8D1E5BA4">2020</mods:date>
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<mods:number id="9938F67BAE449687B304036A94BC8C9E">2020-08-26</mods:number>
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<mods:classification id="82521A6000143175DD2BAC41FBEDA0E3">journal article</mods:classification>
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<mods:identifier id="D528B026C3F6374D1F527365051B09DC" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFC2BB15FF1884DDFAEFFC59" ID-DOI="http://doi.org/10.5281/zenodo.4404245" ID-GBIF-Taxon="177012535" ID-Zenodo-Dep="4404245" LSID="urn:lsid:zoobank.org:act:330A5BB9-B773-4698-B644-5DA3EAC95433" httpUri="http://treatment.plazi.org/id/039187D8FFC2BB15FF1884DDFAEFFC59" lastPageNumber="155" pageId="4" pageNumber="155">
<subSubSection id="C3226545FFC2BB15FF1884DDFE2FFF4E" box="[151,434,151,178]" pageId="4" pageNumber="155" type="nomenclature">
<paragraph id="8B8736CEFFC2BB15FF1884DDFE2FFF4E" blockId="4.[151,434,151,214]" box="[151,434,151,178]" pageId="4" pageNumber="155">
<heading id="D0CF81A2FFC2BB15FF1884DDFE2FFF4E" bold="true" box="[151,434,151,178]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<emphasis id="B94CEADCFFC2BB15FF1884DDFE2FFF4E" bold="true" box="[151,434,151,178]" pageId="4" pageNumber="155">
<taxonomicName id="4C384D4DFFC2BB15FF1884DDFEF5FF4D" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,360,151,178]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="B94CEADCFFC2BB15FF1884DDFEF5FF4D" bold="true" box="[151,360,151,178]" italics="true" pageId="4" pageNumber="155">Zarcosia bedosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFC2BB15FEFF84DDFE2FFF4E" box="[368,434,151,177]" pageId="4" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF1884F1FE08FF29" box="[151,405,187,214]" pageId="4" pageNumber="155" type="description">
<paragraph id="8B8736CEFFC2BB15FF1884F1FE08FF29" blockId="4.[151,434,151,214]" box="[151,405,187,214]" pageId="4" pageNumber="155">
(
<figureCitation id="13032A4BFFC2BB15FF1084F1FE96FF2A" box="[159,267,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="4" pageNumber="155">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FE9784F1FEA2FF2A" box="[280,319,187,213]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="4" pageNumber="155">
<quantity id="4CC09B2BFFC2BB15FE9784F1FEA2FF2A" box="[280,319,187,213]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="155" unit="m" value="3.0">3m</quantity>
</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FEC484F1FEF7FF2A" box="[331,362,187,213]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="4" pageNumber="155">4b</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FEF984F1FE11FF29" box="[374,396,187,214]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="4" pageNumber="155">6i</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF188549FAF9FEE2" box="[151,1380,259,286]" pageId="4" pageNumber="155" type="etymology">
<paragraph id="8B8736CEFFC2BB15FF188549FAF9FEE2" blockId="4.[151,1437,259,934]" box="[151,1380,259,286]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF188549FE83FEE2" bold="true" box="[151,286,259,285]" pageId="4" pageNumber="155">Etymology.</emphasis>
The species is named after Anne Bedos in recognition of her efforts to collect material for others.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF48856DFB71FEBE" box="[199,1260,294,322]" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8B8736CEFFC2BB15FF48856DFB71FEBE" blockId="4.[151,1437,259,934]" box="[199,1260,294,322]" pageId="4" pageNumber="155">
<materialsCitation id="3B503C93FFC2BB15FF48856DFB74FEBE" ID-GBIF-Occurrence="3020160469" box="[199,1257,294,322]" country="Laos" latitude="17.966667" location="Tham Thon" longLatPrecision="1277" longitude="104.51667" municipality="Ban Thongkha" pageId="4" pageNumber="155" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
<emphasis id="B94CEADCFFC2BB15FF48856DFEFBFEBE" bold="true" box="[199,358,295,321]" pageId="4" pageNumber="155">
<typeStatus id="5483886CFFC2BB15FF48856DFE9FFEBE" box="[199,258,295,321]" pageId="4" pageNumber="155">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC2BB15FEE38562FE3AFEBD" box="[364,423,296,322]" name="Laos" pageId="4" pageNumber="155">Laos</collectingCountry>
:
<collectingRegion id="49FCF82CFFC2BB15FE3D856DFD33FEBE" box="[434,686,295,321]" country="Laos" name="Khammouan" pageId="4" pageNumber="155">Khammouan province</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC2BB15FD368562FCFCFEBE" box="[697,865,295,322]" pageId="4" pageNumber="155">Ban Thôngkha</collectingMunicipality>
,
<location id="8EE76015FFC2BB15FCE3856DFC72FEBE" LSID="urn:lsid:plazi:treatment:039187D8FFC2BB15FF1884DDFAEFFC59:8EE76015FFC2BB15FCE3856DFC72FEBE" box="[876,1007,295,321]" country="Laos" latitude="17.966667" longLatPrecision="1277" longitude="104.51667" municipality="Ban Thongkha" name="Tham Thon" pageId="4" pageNumber="155" stateProvince="Khammouan">Tham Thon</location>
[
<geoCoordinate id="EE0C5009FFC2BB15FC72856DFBF9FEBD" box="[1021,1124,295,322]" degrees="17" direction="north" minutes="58" orientation="latitude" pageId="4" pageNumber="155" precision="925" value="17.966667">17°58’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC2BB15FBE3856DFB7CFEBE" box="[1132,1249,294,321]" degrees="104" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="155" precision="925" value="104.51667">104°31’ E</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF488501FD81FE9A" box="[199,540,331,358]" pageId="4" pageNumber="155" type="distribution">
<paragraph id="8B8736CEFFC2BB15FF488501FD81FE9A" blockId="4.[151,1437,259,934]" box="[199,540,331,358]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF488501FEFCFE9A" bold="true" box="[199,353,331,357]" pageId="4" pageNumber="155">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC2BB15FEE88506FE00FE99" box="[359,413,332,358]" name="Laos" pageId="4" pageNumber="155">Laos</collectingCountry>
(
<figureCitation id="13032A4BFFC2BB15FE238501FD96FE9A" box="[428,523,331,357]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="4" pageNumber="155">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF488525FD18FCC5" pageId="4" pageNumber="155" type="description">
<paragraph id="8B8736CEFFC2BB15FF488525FC83FE52" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF488525FEC4FE76" bold="true" box="[199,345,367,393]" pageId="4" pageNumber="155">Description.</emphasis>
(
<figureCitation id="13032A4BFFC2BB15FEE88525FE7DFE76" box="[359,480,367,393]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="4" pageNumber="155">
Figure
<quantity id="4CC09B2BFFC2BB15FE368525FE7DFE76" box="[441,480,367,393]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="155" unit="m" value="3.0">3m</quantity>
</figureCitation>
). Female unknown. Body length.
<specimenCount id="9D3EFD47FFC2BB15FCD4853AFCEDFE75" box="[859,880,368,394]" pageId="4" pageNumber="155" type="male">♂</specimenCount>
<typeStatus id="5483886CFFC2BB15FCF88525FC40FE76" box="[887,989,367,393]" pageId="4" pageNumber="155" type="holotype">holotype</typeStatus>
:
<quantity id="4CC09B2BFFC2BB15FC678525FBDCFE75" box="[1000,1089,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="155" unit="mm" value="2.5">2.5 mm</quantity>
. Black, mat, covered with very short, pulverulent, silver pubescence. Appendages brown.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF4885FDFB2FFE0A" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4885FDFE99FE2E" box="[199,260,439,465]" italics="true" pageId="4" pageNumber="155">Head</emphasis>
. Moderately transverse, wider than pronotum. Palpi yellowish, last article securiform. Integument mat, coarsely rugose, grainy. Occiput lined with a fringe of dense and short silver hairs posteriorly.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF4885B5FC31FDE6" blockId="4.[151,1437,259,934]" box="[199,940,511,538]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4885B5FEACFDE6" box="[199,305,511,537]" italics="true" pageId="4" pageNumber="155">Antennae</emphasis>
. A fifth longer than elytra, entirely dark brown, filiform.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF48866EFB5DFD9E" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF48866EFEABFDC2" box="[199,310,548,573]" italics="true" pageId="4" pageNumber="155">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFC2BB15FECB8669FE30FDC1" box="[324,429,547,574]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="4" pageNumber="155">Figure 6i</figureCitation>
). Convex, slightly longer than wide, sides parallel from the base to the posterior two thirds, then progressively convergent anteriorly. Sides marked with lateral impressions and callosities.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF488621FB3AFD32" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF488621FE90FD7A" box="[199,269,619,645]" italics="true" pageId="4" pageNumber="155">Elytra</emphasis>
. Length:
<quantity id="4CC09B2BFFC2BB15FEF48621FE79FD79" box="[379,484,619,646]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="155" unit="mm" value="1.75">1.75 mm</quantity>
. Punctation regular, as dense as on the pronotum, with larger points. Space between points mat, alutaceous. Disc subconvex, with two broad depression in the anterior third. Pubescence dense and short, comprised of very short hairs borne from the points and interspersed pulverulent hairs.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF488692FCE9FD0E" blockId="4.[151,1437,259,934]" box="[199,884,727,754]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF488692FF61FD0E" box="[199,252,728,753]" italics="true" pageId="4" pageNumber="155">Legs</emphasis>
. Particularly elongated. No other notable modifications.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF4886B1FD18FCC5" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4886B1FEA9FCEA" box="[199,308,763,789]" italics="true" pageId="4" pageNumber="155">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFC2BB15FECC86B1FE28FCE9" box="[323,437,763,790]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="4" pageNumber="155">Figure 4b</figureCitation>
). Sturdy, sides weakly convergent in dorsal view and suddenly tapered apically. Strongly convex in lateral view, bent upward at apex.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF488709FAEFFC59" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8B8736CEFFC2BB15FF488709FC43FC7E" blockId="4.[151,1437,259,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF488709FEE5FCA2" bold="true" box="[199,376,835,861]" pageId="4" pageNumber="155">Type material.</emphasis>
<materialsCitation id="3B503C93FFC2BB15FE0D8709FC43FC7E" ID-GBIF-Occurrence="3020160453" collectingDate="1998-02-12" collectionCode="MNHN, A" collectorName="L. Deharveng &amp; Bedos" country="Laos" elevation="230" latitude="17.966667" location="Tham Thon" longLatPrecision="1277" longitude="104.51667" municipality="Ban Thongkha" pageId="4" pageNumber="155" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="5483886CFFC2BB15FE0D8709FE77FCA2" box="[386,490,835,861]" pageId="4" pageNumber="155" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFC2BB15FE718709FD88FCA2" box="[510,533,835,861]" pageId="4" pageNumber="155" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC2BB15FDAA870EFD1DFCA1" box="[549,640,836,862]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="155">MNHN</collectionCode>
).
<emphasis id="B94CEADCFFC2BB15FD1A8709FD52FCA2" bold="true" box="[661,719,835,861]" pageId="4" pageNumber="155">
<collectingCountry id="F32F765EFFC2BB15FD1A8709FD52FCA2" box="[661,719,835,861]" name="Laos" pageId="4" pageNumber="155">Laos</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFC2BB15FD6F8709FC7DFCA2" box="[736,992,835,861]" country="Laos" name="Khammouan" pageId="4" pageNumber="155">Khammouan province</collectingRegion>
,
<collectingMunicipality id="6BE3ACB4FFC2BB15FC60870EFB07FCA2" box="[1007,1178,835,862]" pageId="4" pageNumber="155">Ban Thôngkha</collectingMunicipality>
,
<location id="8EE76015FFC2BB15FB278709FAB3FCA2" LSID="urn:lsid:plazi:treatment:039187D8FFC2BB15FF1884DDFAEFFC59:8EE76015FFC2BB15FB278709FAB3FCA2" box="[1192,1326,835,861]" country="Laos" latitude="17.966667" longLatPrecision="1277" longitude="104.51667" municipality="Ban Thongkha" name="Tham Thon" pageId="4" pageNumber="155" stateProvince="Khammouan">Tham Thon</location>
(
<quantity id="4CC09B2BFFC2BB15FACF8709FA12FCA1" box="[1344,1423,835,862]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="155" unit="m" value="230.0">
<elevation id="0015D1FDFFC2BB15FACF8709FA12FCA1" box="[1344,1423,835,862]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="155" unit="m" value="230.0">230 m</elevation>
</quantity>
), cave, by hand,
<geoCoordinate id="EE0C5009FFC2BB15FECF872DFE38FC7D" box="[320,421,871,898]" degrees="17" direction="north" minutes="58" orientation="latitude" pageId="4" pageNumber="155" precision="925" value="17.966667">17°58’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC2BB15FE22872DFD80FC7D" box="[429,541,871,898]" degrees="104" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="155" precision="925" value="104.51667">104°31’ E</geoCoordinate>
(
<collectorName id="26CD5318FFC2BB15FDA2872DFD55FC7E" box="[557,712,871,897]" pageId="4" pageNumber="155">L. Deharveng</collectorName>
&amp;
<collectionCode id="ED29AE0BFFC2BB15FD65872DFC9CFC7E" box="[746,769,871,897]" pageId="4" pageNumber="155">A</collectionCode>
.
<collectorName id="26CD5318FFC2BB15FC83872DFCC9FC7D" box="[780,852,871,898]" pageId="4" pageNumber="155">Bedos</collectorName>
)
<date id="FF86100EFFC2BB15FCED872DFC47FC7E" box="[866,986,871,897]" pageId="4" pageNumber="155" value="1998-02-12">
<collectingDate id="EFC2E9E6FFC2BB15FCED872DFC47FC7E" box="[866,986,871,897]" pageId="4" pageNumber="155" value="1998-02-12">12.II.1998</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF4887C1FAEFFC59" blockId="4.[151,1437,259,934]" box="[199,1394,907,934]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4887C1FF62FC5A" bold="true" box="[199,255,907,933]" pageId="4" pageNumber="155">Note</emphasis>
. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.
</paragraph>
</subSubSection>
</treatment>
</document>
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