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<document id="F3895654346F49607FBBAE3A812C4E22" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFC2BB14FF1881F1FC9CFC7E" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia brunnea Gompel 2020, n. sp." docType="treatment" docUuid="E5FE45C9-1723-45CE-83A3-CB9982F0C062" docUuidSource="ZooBank" docVersion="8" lastPageNumber="156" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="155" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E8466F9B2445910351CD166B78CA3402">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="FFA5F17C4D4832D81646610029DA5563">Gompel, Nicolas</mods:namePart>
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<mods:date id="001B8702374FF273F46AF8B49BBE0427">2020</mods:date>
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<mods:number id="E3C1FAD94148D15B6794110C797917AE">2020-08-26</mods:number>
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<mods:classification id="2A5BBDD90DDDEB87F0340C2B3C6D7C74">journal article</mods:classification>
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<mods:identifier id="FE130ED6BB4F646573ADFFF49BE5DE20" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFC2BB14FF1881F1FC9CFC7E" ID-DOI="http://doi.org/10.5281/zenodo.4404247" ID-GBIF-Taxon="177012536" ID-Zenodo-Dep="4404247" LSID="urn:lsid:zoobank.org:act:E5FE45C9-1723-45CE-83A3-CB9982F0C062" httpUri="http://treatment.plazi.org/id/039187D8FFC2BB14FF1881F1FC9CFC7E" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
<subSubSection id="C3226545FFC2BB15FF1881F1FE24FA2A" box="[151,441,1467,1494]" pageId="4" pageNumber="155" type="nomenclature">
<paragraph id="8B8736CEFFC2BB15FF1881F1FE24FA2A" blockId="4.[151,456,1467,1530]" box="[151,441,1467,1494]" pageId="4" pageNumber="155">
<heading id="D0CF81A2FFC2BB15FF1881F1FE24FA2A" bold="true" box="[151,441,1467,1494]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<emphasis id="B94CEADCFFC2BB15FF1881F1FE24FA2A" bold="true" box="[151,441,1467,1494]" pageId="4" pageNumber="155">
<taxonomicName id="4C384D4DFFC2BB15FF1881F1FEF2FA29" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,367,1467,1494]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="B94CEADCFFC2BB15FF1881F1FEF2FA29" bold="true" box="[151,367,1467,1494]" italics="true" pageId="4" pageNumber="155">Zarcosia brunnea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57A7FFC2BB15FEF981F1FE24FA2A" box="[374,441,1467,1493]" pageId="4" pageNumber="155" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3226545FFC2BB15FF188195FE55FA05" box="[151,456,1503,1530]" pageId="4" pageNumber="155" type="description">
<paragraph id="8B8736CEFFC2BB15FF188195FE55FA05" blockId="4.[151,456,1467,1530]" box="[151,456,1503,1530]" pageId="4" pageNumber="155">
(
<figureCitation id="13032A4BFFC2BB15FF108195FE96FA06" box="[159,267,1503,1529]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="4" pageNumber="155">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FE978195FEB3FA06" box="[280,302,1503,1529]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="4" pageNumber="155">
<quantity id="4CC09B2BFFC2BB15FE978195FEB3FA06" box="[280,302,1503,1529]" metricMagnitude="3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="155" unit="t" value="3.0">3t</quantity>
</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FECB8195FEFBFA06" box="[324,358,1503,1529]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="4" pageNumber="155">4w</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FEF38195FE05FA05" box="[380,408,1503,1530]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="4" pageNumber="155">5s</figureCitation>
;
<figureCitation id="13032A4BFFC2BB15FE2B8195FE5CFA05" box="[420,449,1503,1530]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="4" pageNumber="155">6o</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF18826DFB09F9BE" box="[151,1172,1575,1602]" pageId="4" pageNumber="155" type="etymology">
<paragraph id="8B8736CEFFC2BB15FF18826DFB09F9BE" blockId="4.[151,1437,1575,2033]" box="[151,1172,1575,1602]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF18826DFE83F9BE" bold="true" box="[151,286,1575,1601]" pageId="4" pageNumber="155">Etymology.</emphasis>
The species is named in reference to the overall brown color of the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF488201FF24F976" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8B8736CEFFC2BB15FF488201FF24F976" blockId="4.[151,1437,1575,2033]" pageId="4" pageNumber="155">
<materialsCitation id="3B503C93FFC2BB15FF488201FF28F976" ID-GBIF-Occurrence="3020160399" country="Sri Lanka" county="Victoria Rendenigala Ramtembe Sanctuary" latitude="7.2166667" location="Central" longLatPrecision="1303" longitude="80.95" municipality="Kandy district" pageId="4" pageNumber="155" specimenCount="1" typeStatus="holotype">
<emphasis id="B94CEADCFFC2BB15FF488201FEF8F99A" bold="true" box="[199,357,1611,1637]" pageId="4" pageNumber="155">
<typeStatus id="5483886CFFC2BB15FF488201FE9FF99A" box="[199,258,1611,1637]" pageId="4" pageNumber="155">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFC2BB15FEE48201FE40F99A" box="[363,477,1611,1637]" name="Sri Lanka" pageId="4" pageNumber="155">Sri Lanka</collectingCountry>
:
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,
<collectingMunicipality id="6BE3ACB4FFC2BB15FDC88201FD74F99A" box="[583,745,1611,1637]" pageId="4" pageNumber="155">Kandy district</collectingMunicipality>
,
<collectingCounty id="62E64E42FFC2BB15FD7C8201FB4BF99A" box="[755,1238,1611,1637]" pageId="4" pageNumber="155">Victoria Rendenigala Ramtembe Sanctuary</collectingCounty>
[
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<geoCoordinate id="EE0C5009FFC2BB15FADD8201FF30F976" degrees="80" direction="east" minutes="57" orientation="longitude" pageId="4" pageNumber="155" precision="925" value="80.95">80°57’ E</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB15FF4882D9FDC9F952" box="[199,596,1683,1709]" pageId="4" pageNumber="155" type="distribution">
<paragraph id="8B8736CEFFC2BB15FF4882D9FDC9F952" blockId="4.[151,1437,1575,2033]" box="[199,596,1683,1709]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4882D9FEFCF952" bold="true" box="[199,353,1683,1709]" pageId="4" pageNumber="155">Distribution.</emphasis>
<collectingCountry id="F32F765EFFC2BB15FEE882D9FE48F952" box="[359,469,1683,1709]" name="Sri Lanka" pageId="4" pageNumber="155">Sri Lanka</collectingCountry>
(
<figureCitation id="13032A4BFFC2BB15FE6B82D9FDDEF952" box="[484,579,1683,1709]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="4" pageNumber="155">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC2BB14FF4882FDFC6FFE0A" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="description">
<paragraph id="8B8736CEFFC2BB15FF4882FDFD26F90A" blockId="4.[151,1437,1575,2033]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4882FDFEC4F92E" bold="true" box="[199,345,1719,1745]" pageId="4" pageNumber="155">Description.</emphasis>
(
<figureCitation id="13032A4BFFC2BB15FEE682FDFE48F92E" box="[361,469,1719,1745]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="4" pageNumber="155">
Figure
<quantity id="4CC09B2BFFC2BB15FE3382FDFE48F92E" box="[444,469,1719,1745]" metricMagnitude="3" metricUnit="kg" metricValue="3.0" pageId="4" pageNumber="155" unit="t" value="3.0">3t</quantity>
</figureCitation>
). Female unknown. Body length.
<specimenCount id="9D3EFD47FFC2BB15FCD782F2FCEDF92D" box="[856,880,1720,1746]" pageId="4" pageNumber="155" type="male">♂</specimenCount>
: 2.80±
<quantity id="4CC09B2BFFC2BB15FC3182FDFBBEF92D" box="[958,1059,1719,1746]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="155" unit="mm" value="0.13">0.13 mm</quantity>
(n=3). Robust, sturdy, uniformly reddish brown, covered with double pubescence.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF4882B5FDBBF8C2" blockId="4.[151,1437,1575,2033]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4882B5FE99F8E6" box="[199,260,1791,1817]" italics="true" pageId="4" pageNumber="155">Head</emphasis>
. Transverse, wider than pronotum, covered with blond pubescence. Punctation fine and very dense. Palpi dark orange, last article securiform.
</paragraph>
<paragraph id="8B8736CEFFC2BB15FF48830DFDB7F87A" blockId="4.[151,1437,1575,2033]" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF48830DFEACF89E" box="[199,305,1863,1889]" italics="true" pageId="4" pageNumber="155">Antennae</emphasis>
. Strong, shorter than elytra, antennomere 3–10 short, at most twice longer than wide, conical. Covered with hirsute blond pubescence.
</paragraph>
<paragraph id="8B8736CEFFC2BB14FF4883DAFE2DFF2A" blockId="4.[151,1437,1575,2033]" lastBlockId="5.[151,1437,151,897]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
<emphasis id="B94CEADCFFC2BB15FF4883DAFEABF856" box="[199,310,1936,1961]" italics="true" pageId="4" pageNumber="155">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFC2BB15FEC883C5FE27F855" box="[327,442,1935,1962]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="4" pageNumber="155">Figure 6o</figureCitation>
). Globose, as long as wide, disc convex. Sides diverging outward from the base to the apical third, then converging markedly inward toward apical margin anteriorly. Densely punctuated, space between points smaller than the diameters of points. Covered with double pubescence, moderately long blond hairs borne from the punctation and pulverulent interspersed hairs. Base bi-sinuated with a dense fringe of golden pubescence anterior to the scutellum.
</paragraph>
<paragraph id="8B8736CEFFC3BB14FF488495FCACFEBD" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF488495FE90FF06" box="[199,269,223,249]" italics="true" pageId="5" pageNumber="156">Elytra</emphasis>
. Length:
<specimenCount id="9D3EFD47FFC3BB14FEF88495FE12FF06" box="[375,399,223,249]" pageId="5" pageNumber="156" type="male">♂</specimenCount>
: 1.92±
<quantity id="4CC09B2BFFC3BB14FE558495FDA3FF06" box="[474,574,223,249]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="156" unit="mm" value="0.03">0.03 mm</quantity>
(n=3). Densely punctuated, space between points larger than the points themselves, covered with double pubescence. Moderately convex, depressed on disc. Apex with a few long hairs (similar to
<taxonomicName id="4C384D4DFFC3BB14FF3B856DFEFBFEBE" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[180,358,295,321]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="srilankaensis" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FF3B856DFEFBFEBE" box="[180,358,295,321]" italics="true" pageId="5" pageNumber="156">Z. srilankaensis</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FEE3856DFE36FEBE" bold="true" box="[364,427,295,321]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FEE3856DFE36FEBE" box="[364,427,295,321]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<figureCitation id="13032A4BFFC3BB14FE36856DFDBAFEBE" box="[441,551,294,321]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="5" pageNumber="156">Figure 5h</figureCitation>
, but much fewer hairs).
</paragraph>
<paragraph id="8B8736CEFFC3BB14FF488501FE87FE52" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF488501FF61FE9B" box="[199,252,331,356]" italics="true" pageId="5" pageNumber="156">Legs</emphasis>
. Short, sturdy. Sexual characters: foretibia with a sharp tooth on their inner side (
<figureCitation id="13032A4BFFC3BB14FB0C8501FB72FE9A" box="[1155,1263,330,357]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="5" pageNumber="156">Figure 5s</figureCitation>
). Mesothoracic femora with an acute apical tooth, straight and pointing outward (similar to that of
<taxonomicName id="4C384D4DFFC3BB14FBD28525FA05FE76" authority="(Pic, 1912)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Pic" baseAuthorityYear="1912" box="[1117,1432,367,393]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="dentatifemur">
<emphasis id="B94CEADCFFC3BB14FBD28525FA91FE76" box="[1117,1292,367,393]" italics="true" pageId="5" pageNumber="156">Z. dentatifemur</emphasis>
(Pic, 1912)
</taxonomicName>
,
<figureCitation id="13032A4BFFC3BB14FF1885D9FE90FE52" box="[151,269,402,429]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="5" pageNumber="156">Figure 5w</figureCitation>
).
</paragraph>
<paragraph id="8B8736CEFFC3BB14FF4885FDFC6FFE0A" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF4885FDFEA9FE2E" box="[199,308,439,465]" italics="true" pageId="5" pageNumber="156">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFC3BB14FECB85FDFE23FE2E" box="[324,446,439,465]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="5" pageNumber="156">Figure 4w</figureCitation>
). Sturdy, sides regularly parallel in dorsal view, strongly convergent in the apical fifth, ended with a small rounded lobe. Strongly and regularly bent in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFC3BB14FF4885B5FC9CFC7E" pageId="5" pageNumber="156" type="materials_examined">
<paragraph id="8B8736CEFFC3BB14FF4885B5FCDCFD56" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF4885B5FEEAFDE7" bold="true" box="[199,375,510,537]" pageId="5" pageNumber="156">Type material.</emphasis>
<materialsCitation id="3B503C93FFC3BB14FE0E85B4FC47FDC2" ID-GBIF-Occurrence="3020160426" collectingDate="1999-08-23" collectionCode="UCDC" collectorName="M. &amp; J. Wasbauer" country="Sri Lanka" county="Victoria Rendenigala Ramtembe Sanctuary" elevation="445" latitude="7.2166667" location="Central" longLatPrecision="1303" longitude="80.95" municipality="Kandy district" pageId="5" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B94CEADCFFC3BB14FE0E85B4FE61FDE6" bold="true" box="[385,508,510,537]" pageId="5" pageNumber="156">
<collectingCountry id="F32F765EFFC3BB14FE0E85B4FE61FDE6" box="[385,508,510,537]" name="Sri Lanka" pageId="5" pageNumber="156">Sri Lanka</collectingCountry>
</emphasis>
:
<typeStatus id="5483886CFFC3BB14FD8285B5FDE8FDE6" box="[525,629,511,537]" pageId="5" pageNumber="156" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFC3BB14FD0785B5FD02FDE6" box="[648,671,511,537]" pageId="5" pageNumber="156" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFC3BB14FD2185B5FC9CFDE6" box="[686,769,511,537]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="5" pageNumber="156" type="Museum">UCDC</collectionCode>
).
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,
<collectingMunicipality id="6BE3ACB4FFC3BB14FCF785B5FB80FDE6" box="[888,1053,511,537]" pageId="5" pageNumber="156">Kandy district</collectingMunicipality>
,
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,
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(
<quantity id="4CC09B2BFFC3BB14FE738668FDDAFDC2" box="[508,583,546,573]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="5" pageNumber="156" unit="m" value="445.0">
<elevation id="0015D1FDFFC3BB14FE738668FDDAFDC2" box="[508,583,546,573]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="5" pageNumber="156" unit="m" value="445.0">445 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC3BB14FDEE8669FCA9FDC1" box="[609,820,547,574]" pageId="5" pageNumber="156">M. &amp; J. Wasbauer</collectorName>
)
<date id="FF86100EFFC3BB14FCCB8669FC47FDC2" box="[836,986,547,573]" pageId="5" pageNumber="156" value="1999-08-23">
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</date>
</materialsCitation>
.
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<typeStatus id="5483886CFFC3BB14FC688669FBC5FDC2" box="[999,1112,547,573]" pageId="5" pageNumber="156" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B94CEADCFFC3BB14FBEB8668FB43FDC2" bold="true" box="[1124,1246,546,573]" pageId="5" pageNumber="156">
<collectingCountry id="F32F765EFFC3BB14FBEB8668FB43FDC2" box="[1124,1246,546,573]" name="Sri Lanka" pageId="5" pageNumber="156">Sri Lanka</collectingCountry>
</emphasis>
:
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,
<collectingMunicipality id="6BE3ACB4FFC3BB14FADD8669FF75FD9E" pageId="5" pageNumber="156">Kandy district</collectingMunicipality>
,
<collectingCounty id="62E64E42FFC3BB14FF7C860DFD46FD9E" box="[243,731,583,609]" pageId="5" pageNumber="156">Victoria Rendenigala Ramtembe Sanctuary</collectingCounty>
,
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<geoCoordinate id="EE0C5009FFC3BB14FCD9860CFC24FD9E" box="[854,953,582,609]" degrees="80" direction="east" minutes="57" orientation="longitude" pageId="5" pageNumber="156" precision="925" value="80.95">80°57’ E</geoCoordinate>
(
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<elevation id="0015D1FDFFC3BB14FC46860CFB8EFD9E" box="[969,1043,582,609]" metricMagnitude="2" metricUnit="m" metricValue="4.45" pageId="5" pageNumber="156" unit="m" value="445.0">445 m</elevation>
</quantity>
) (
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)
<date id="FF86100EFFC3BB14FA85860DFA01FD9E" box="[1290,1436,583,609]" pageId="5" pageNumber="156" value="1999-08-23">
<collectingDate id="EFC2E9E6FFC3BB14FA85860DFA01FD9E" box="[1290,1436,583,609]" pageId="5" pageNumber="156" value="1999-08-23">23.VIII.1999</collectingDate>
</date>
(
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<typeStatus id="5483886CFFC3BB14FF5F8621FEAEFD7A" box="[208,307,619,645]" pageId="5" pageNumber="156" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC3BB14FEB28621FE10FD7A" box="[317,397,619,645]" pageId="5" pageNumber="156">NGPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B503C93FFC3BB14FE108621FDAEFD56" ID-GBIF-Occurrence="3020160397" collectingDate="1999-08-23" collectorName="M. &amp; J. Wasbauer" country="Sri Lanka" county="Victoria Rendenigala Ramtembe Sanctuary" elevation="135" latitude="7.2" location="Central" longLatPrecision="1303" longitude="80.95" municipality="Kandy district" pageId="5" pageNumber="156" specimenCount="1" typeStatus="paratype">
<location id="8EE76015FFC3BB14FE108621FE69FD7A" LSID="urn:lsid:plazi:treatment:039187D8FFC2BB14FF1881F1FC9CFC7E:8EE76015FFC3BB14FE108621FE69FD7A" box="[415,500,619,645]" country="Sri Lanka" county="Victoria Rendenigala Ramtembe Sanctuary" latitude="7.2" longLatPrecision="1303" longitude="80.95" municipality="Kandy district" name="Central" pageId="5" pageNumber="156">Central</location>
,
<collectingMunicipality id="6BE3ACB4FFC3BB14FE708621FD3DFD7A" box="[511,672,619,645]" pageId="5" pageNumber="156">Kandy district</collectingMunicipality>
,
<collectingCounty id="62E64E42FFC3BB14FD258621FB11FD7A" box="[682,1164,619,645]" pageId="5" pageNumber="156">Victoria Rendenigala Ramtembe Sanctuary</collectingCounty>
,
<geoCoordinate id="EE0C5009FFC3BB14FB198621FB66FD7A" box="[1174,1275,619,645]" degrees="07" direction="north" minutes="12" orientation="latitude" pageId="5" pageNumber="156" precision="925" value="7.2">07°12’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFC3BB14FA8E8620FAFEFD7A" box="[1281,1379,618,645]" degrees="80" direction="east" minutes="57" orientation="longitude" pageId="5" pageNumber="156" precision="925" value="80.95">80°57’ E</geoCoordinate>
(
<quantity id="4CC09B2BFFC3BB14FAFE8620FF33FD56" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="156" unit="m" value="135.0">
<elevation id="0015D1FDFFC3BB14FAFE8620FF33FD56" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="156" unit="m" value="135.0">135 m</elevation>
</quantity>
) (
<collectorName id="26CD5318FFC3BB14FF4986C5FE0EFD55" box="[198,403,655,682]" pageId="5" pageNumber="156">M. &amp; J. Wasbauer</collectorName>
)
<date id="FF86100EFFC3BB14FE2E86C5FDAEFD56" box="[417,563,655,681]" pageId="5" pageNumber="156" value="1999-08-23">
<collectingDate id="EFC2E9E6FFC3BB14FE2E86C5FDAEFD56" box="[417,563,655,681]" pageId="5" pageNumber="156" value="1999-08-23">23.VIII.1999</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B503C93FFC3BB14FDB586C5FCA3FD56" ID-GBIF-Occurrence="3020160406" box="[570,830,655,681]" collectionCode="UCDC" country="Sri Lanka" pageId="5" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
(
<specimenCount id="9D3EFD47FFC3BB14FDCA86C5FDF0FD56" box="[581,621,655,681]" pageId="5" pageNumber="156" type="male">1 ♂</specimenCount>
<typeStatus id="5483886CFFC3BB14FDFB86C5FD4AFD56" box="[628,727,655,681]" pageId="5" pageNumber="156" type="paratype">paratype</typeStatus>
,
<collectionCode id="ED29AE0BFFC3BB14FD6C86C5FCABFD56" box="[739,822,655,681]" country="USA" httpUri="http://grbio.org/cool/kd99-229r" name="R. M. Bohart Museum of Entomology" pageId="5" pageNumber="156" type="Museum">UCDC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736CEFFC3BB14FF4886F9FC9CFC7E" blockId="5.[151,1437,151,897]" pageId="5" pageNumber="156">
<emphasis id="B94CEADCFFC3BB14FF4886F9FF62FD32" bold="true" box="[199,255,691,717]" pageId="5" pageNumber="156">Note</emphasis>
. This species was collected together with
<taxonomicName id="4C384D4DFFC3BB14FD4086F9FC60FD32" authority="(Pic, 1912)" authorityName="Gompel" authorityYear="2020" baseAuthorityName="Pic" baseAuthorityYear="1912" box="[719,1021,691,717]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="dentatifemur">
<emphasis id="B94CEADCFFC3BB14FD4086F9FCE7FD32" box="[719,890,691,717]" italics="true" pageId="5" pageNumber="156">Z. dentatifemur</emphasis>
(Pic, 1912)
</taxonomicName>
,
<taxonomicName id="4C384D4DFFC3BB14FB8786F9FB01FD32" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[1032,1180,691,717]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="spinifemur" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FB8786F9FB01FD32" box="[1032,1180,691,717]" italics="true" pageId="5" pageNumber="156">Z. spinifemur</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FB2D86F9FB42FD32" bold="true" box="[1186,1247,691,717]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FB2D86F9FB42FD32" box="[1186,1247,691,717]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C384D4DFFC3BB14FB6386F9FA01FD32" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[1260,1436,691,717]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="srilankaensis" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FB6386F9FA01FD32" box="[1260,1436,691,717]" italics="true" pageId="5" pageNumber="156">Z. srilankaensis</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FF18869DFF48FD0E" bold="true" box="[151,213,727,753]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FF18869DFF48FD0E" box="[151,213,727,753]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C384D4DFFC3BB14FE86869DFE03FD0E" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[265,414,727,753]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FE86869DFE03FD0E" box="[265,414,727,753]" italics="true" pageId="5" pageNumber="156">Z. intermedia</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FE2B869DFE7FFD0E" bold="true" box="[420,482,727,753]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FE2B869DFE7FFD0E" box="[420,482,727,753]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
The sample contained a large series of females, which I could not assign to the respective species. In particular,
<taxonomicName id="4C384D4DFFC3BB14FE4E86B1FDA1FCEA" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[449,572,763,789]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="brunnea" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FE4E86B1FDA1FCEA" box="[449,572,763,789]" italics="true" pageId="5" pageNumber="156">Z. brunnea</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FDCB86B1FD19FCEA" bold="true" box="[580,644,763,789]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FDCB86B1FD19FCEA" box="[580,644,763,789]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C384D4DFFC3BB14FD1C86B1FCDBFCEA" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[659,838,763,789]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="srilankaensis" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FD1C86B1FCDBFCEA" box="[659,838,763,789]" italics="true" pageId="5" pageNumber="156">Z. srilankaensis</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FCC286B1FC10FCEA" bold="true" box="[845,909,763,789]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FCC286B1FC10FCEA" box="[845,909,763,789]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C384D4DFFC3BB14FC4A86B1FBC0FCEA" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[965,1117,763,789]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B94CEADCFFC3BB14FC4A86B1FBC0FCEA" box="[965,1117,763,789]" italics="true" pageId="5" pageNumber="156">Z. intermedia</emphasis>
</taxonomicName>
<emphasis id="B94CEADCFFC3BB14FBEA86B1FB39FCEA" bold="true" box="[1125,1188,763,789]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A27F57A7FFC3BB14FBEA86B1FB39FCEA" box="[1125,1188,763,789]" pageId="5" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
form a very homogeneous group. Their males are easily distinguished by stable sexual characters (aedeagus, teeth on the tibiae, hooks on the femora), but in spite of extensive comparisons of the females collected with them, I could not decide whether these females represented one or more taxa, nor which.
</paragraph>
</subSubSection>
</treatment>
</document>
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