151 lines
17 KiB
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151 lines
17 KiB
XML
<document id="EDF3BF2480AE627CD7CC76A2DE634424" ID-DOI="10.1080/00222933.2018.1526345" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747340" ID-ZooBank="51DA9869-D786-4A8A-8C1F-62DC1BB67923" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620671110245" checkinUser="carolina" docAuthor="Ocaña, Oscar & Çinar, Melih Ertan" docDate="2018" docId="039187AEFFF30833FDE76A13FC29FD8D" docLanguage="en" docName="JNATHIST.52.35-36.2243-2282.pdf" docOrigin="Journal of Natural History 52 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2018.1526345" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Marmara Ocaña & Çinar 2018, gen. nov." docType="treatment" docVersion="3" lastPageNumber="2246" masterDocId="FFA8FFD6FFF00837FFFA6E27FFC4FFA9" masterDocTitle="Descriptions of two new genera, six new species and three new records of Anthozoa (Cnidaria) from the Sea of Marmara" masterLastPageNumber="2282" masterPageNumber="2243" pageNumber="2245" updateTime="1698914570521" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="04303CA0F39F6B0FCDC36BF48DE57C90">Descriptions of two new genera, six new species and three new records of Anthozoa (Cnidaria) from the Sea of Marmara</mods:title>
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<mods:namePart id="0730D0329FE76BDA7293AAE1ABAB6C6D">Ocaña, Oscar</mods:namePart>
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<mods:namePart id="E04F9305BA6DF94BB2DD16C902837C35">Çinar, Melih Ertan</mods:namePart>
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<mods:title id="2966160E1425E7AEAB355AFA615D085B">Journal of Natural History</mods:title>
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<mods:part id="B2655D5524F24DCD914F1F22A8BB19B1">
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<mods:date id="28E9B5C9A8930B4F3964CCE12683B9C6">2018</mods:date>
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<mods:number id="4D4DCC73B6874F1A44C03AC26C4BA78C">2018-10-23</mods:number>
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<mods:number id="64F369D963A63DD8AE9AC691BD874A46">52</mods:number>
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<mods:number id="EAE732D1E6A8397E7ED4AACB773867E4">35 - 36</mods:number>
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<mods:url id="636B3ADA555E57A83AE9B3CDEAA69EEE">http://dx.doi.org/10.1080/00222933.2018.1526345</mods:url>
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<subSubSection id="C3226533FFF30834FDE76A13FCC3FBE7" box="[541,775,1076,1102]" pageId="3" pageNumber="2245" type="nomenclature">
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<paragraph id="8B8736B8FFF30834FDE76A13FCC3FBE7" blockId="3.[490,825,1008,1102]" box="[541,775,1076,1102]" pageId="3" pageNumber="2245">
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<heading id="D0CF81D4FFF30834FDE76A13FCC3FBE7" box="[541,775,1076,1102]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="2245" reason="2">
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<taxonomicName id="4C384D3BFFF30834FDE76A13FD4BFBE7" ID-CoL="8V668" authority="Ocaña & Çinar, 2018" authorityName="Ocaña & Çinar" authorityYear="2018" box="[541,655,1076,1102]" class="Insecta" family="Gracillariidae" genus="Marmara" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="2245" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94CEAAAFFF30834FDE76A13FD4BFBE7" bold="true" box="[541,655,1076,1102]" italics="true" pageId="3" pageNumber="2245">Marmara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F57D1FFF30834FD626A12FCC3FBE7" box="[664,775,1077,1102]" pageId="3" pageNumber="2245" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226533FFF30834FF5A6AADFCAEFAA3" pageId="3" pageNumber="2245" type="diagnosis">
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<paragraph id="8B8736B8FFF30834FF5A6AADFED0FB0A" blockId="3.[160,1156,1162,1290]" box="[160,276,1162,1187]" pageId="3" pageNumber="2245">
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<emphasis id="B94CEAAAFFF30834FF5A6AADFED0FB0A" bold="true" box="[160,276,1162,1187]" italics="true" pageId="3" pageNumber="2245">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B8736B8FFF30834FF5A6A8AFCAEFAA3" blockId="3.[160,1156,1162,1290]" pageId="3" pageNumber="2245">
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<taxonomicName id="4C384D3BFFF30834FF5A6A8AFED9FB6C" authorityName="Gosse" authorityYear="1858" box="[160,285,1197,1221]" class="Anthozoa" family="Sagartiidae" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="2245" phylum="Cnidaria" rank="family">Sagartiidae</taxonomicName>
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without suckers or tenaculi, with strong endodermic musculature along column. Visible and conspicuous muscular belts along column and capitulum. Sphincter mesogloeal mainly present in upper part of column.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226533FFF30834FF5A6B19FE24FAD0" pageId="3" pageNumber="2245" type="type_taxon">
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<paragraph id="8B8736B8FFF30834FF5A6B19FEF0FAFE" blockId="3.[160,480,1342,1401]" box="[160,308,1342,1367]" pageId="3" pageNumber="2245">
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<emphasis id="B94CEAAAFFF30834FF5A6B19FEF0FAFE" bold="true" box="[160,308,1342,1367]" italics="true" pageId="3" pageNumber="2245">
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<typeStatus id="5483881AFFF30834FF5A6B19FF1CFAFE" box="[160,216,1342,1367]" pageId="3" pageNumber="2245">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="8B8736B8FFF30834FF5A6B47FE24FAD0" blockId="3.[160,480,1342,1401]" box="[160,480,1376,1401]" pageId="3" pageNumber="2245">
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<taxonomicName id="4C384D3BFFF30834FF5A6B47FEBBFAD0" authority="Ocaña & Çinar, 2018" authorityName="Ocaña & Çinar" authorityYear="2018" box="[160,383,1376,1401]" class="Insecta" family="Gracillariidae" genus="Marmara" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="2245" phylum="Arthropoda" rank="species" species="musculata" status="sp. nov.">
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<emphasis id="B94CEAAAFFF30834FF5A6B47FEBBFAD0" box="[160,383,1376,1401]" italics="true" pageId="3" pageNumber="2245">Marmara musculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F57D1FFF30834FE726B46FE24FAD0" box="[392,480,1377,1401]" pageId="3" pageNumber="2245" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3226533FFF30834FF5A6B8AFC9AFA41" box="[160,862,1453,1512]" pageId="3" pageNumber="2245" type="etymology">
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<paragraph id="8B8736B8FFF30834FF5A6B8AFED9FA6F" blockId="3.[160,862,1453,1512]" box="[160,285,1453,1478]" pageId="3" pageNumber="2245">
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<emphasis id="B94CEAAAFFF30834FF5A6B8AFED9FA6F" bold="true" box="[160,285,1453,1478]" italics="true" pageId="3" pageNumber="2245">Etymology</emphasis>
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</paragraph>
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<paragraph id="8B8736B8FFF30834FF5A6BF7FC9AFA41" blockId="3.[160,862,1453,1512]" box="[160,862,1488,1512]" pageId="3" pageNumber="2245">
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The generic name refers to the
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<typeStatus id="5483881AFFF30834FDF66BF7FDFAFA41" box="[524,574,1488,1512]" pageId="3" pageNumber="2245">type</typeStatus>
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locality, Sea of
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<taxonomicName id="4C384D3BFFF30834FD0E6BF7FC9EFA41" authorityName="Ocaña & Çinar" authorityYear="2018" box="[756,858,1488,1512]" class="Insecta" family="Gracillariidae" genus="Marmara" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="2278" phylum="Arthropoda" rank="genus">Marmara</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3226533FFF30833FF5A683BFC29FD8D" lastPageId="4" lastPageNumber="2246" pageId="3" pageNumber="2245" type="discussion">
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<paragraph id="8B8736B8FFF30834FF5A683BFEC2F99C" blockId="3.[160,1156,1564,1727]" box="[160,262,1564,1589]" pageId="3" pageNumber="2245">
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<emphasis id="B94CEAAAFFF30834FF5A683BFEC2F99C" bold="true" box="[160,262,1564,1589]" italics="true" pageId="3" pageNumber="2245">Remarks</emphasis>
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</paragraph>
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<paragraph id="8B8736B8FFF30834FF5A6818FE7FF916" blockId="3.[160,1156,1564,1727]" pageId="3" pageNumber="2245">
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The absence of suckers on the body wall represents a close affinity of this new genus with the genera
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<taxonomicName id="4C384D3BFFF30834FEA46846FD59F9D3" authority="Carlgren 1943" authorityName="Carlgren" authorityYear="1943" box="[350,669,1633,1658]" class="Anthozoa" family="Sagartiidae" genus="Sagartianthus" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="2245" phylum="Cnidaria" rank="genus">
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<emphasis id="B94CEAAAFFF30834FEA46846FE32F9D3" box="[350,502,1633,1658]" italics="true" pageId="3" pageNumber="2245">Sagartianthus</emphasis>
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<bibRefCitation id="EFA94B49FFF30834FE056845FD59F9D3" author="Carlgren O" box="[511,669,1634,1658]" pageId="3" pageNumber="2245" pagination="1 - 43" refId="ref14931" refString="Carlgren O. 1943. East-Asiatic Corallimorpharia and Actiniaria. Kungl Svenska Vetensk Akad Handl. 20: 1 - 43." type="journal article" year="1943">Carlgren 1943</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="4C384D3BFFF30834FD516846FBC6F9D3" authority="Stephenson 1918" authorityName="Stephenson" authorityYear="1918" box="[683,1026,1633,1658]" class="Anthozoa" family="Sagartiidae" genus="Artemidactis" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="2245" phylum="Cnidaria" rank="genus">
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<emphasis id="B94CEAAAFFF30834FD516846FCF6F9D3" box="[683,818,1633,1658]" italics="true" pageId="3" pageNumber="2245">Artemidactis</emphasis>
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Stephenson 1918
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</taxonomicName>
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,
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<taxonomicName id="4C384D3BFFF30834FBF46846FEE9F934" authority="Fischer 1889" authorityName="Fischer" authorityYear="1889" class="Anthozoa" family="Sagartiidae" genus="Actinothoe" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="2245" phylum="Cnidaria" rank="genus">
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<emphasis id="B94CEAAAFFF30834FBF46846FB40F9D3" box="[1038,1156,1633,1658]" italics="true" pageId="3" pageNumber="2245">Actinothoe</emphasis>
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Fischer 1889
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</taxonomicName>
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and
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<taxonomicName id="4C384D3BFFF30834FE9C68A3FDBAF934" authority="Carlgren 1938" authorityName="Carlgren" authorityYear="1938" box="[358,638,1668,1693]" class="Anthozoa" family="Sagartiidae" genus="Anthothoe" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="2245" phylum="Cnidaria" rank="genus">
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<emphasis id="B94CEAAAFFF30834FE9C68A3FE1CF934" box="[358,472,1668,1693]" italics="true" pageId="3" pageNumber="2245">Anthothoe</emphasis>
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<bibRefCitation id="EFA94B49FFF30834FE1A68A2FDBAF934" author="Carlgren O" box="[480,638,1669,1693]" pageId="3" pageNumber="2245" pagination="1 - 148" refId="ref14837" refString="Carlgren O. 1938. South African Actiniaria and Zoantharia. Kungl svenska Vettensk Akad Handl. 17 (3): 1 - 148." type="journal article" year="1938">Carlgren 1938</bibRefCitation>
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</taxonomicName>
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. However, unlike the other genera, the genus
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<taxonomicName id="4C384D3BFFF30834FF5A6881FEC1F916" authority="Ocaña & Çinar, 2018" authorityName="Ocaña & Çinar" authorityYear="2018" box="[160,261,1702,1727]" class="Insecta" family="Gracillariidae" genus="Marmara" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="2245" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B94CEAAAFFF30834FF5A6881FEC1F916" box="[160,261,1702,1727]" italics="true" pageId="3" pageNumber="2245">Marmara</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F57D1FFF30834FEF46880FEBCF916" box="[270,376,1703,1727]" pageId="3" pageNumber="2245" rank="genus">gen. nov.</taxonomicNameLabel>
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has c
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</paragraph>
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<paragraph id="8B8736B8FFF40833FF3A6EA9FC29FD8D" blockId="4.[160,1156,142,548]" pageId="4" pageNumber="2246">
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onspicuous muscular belts of endodermic musculature along the column, which is a peculiar anatomical feature only known in this genus. In the contracted/relaxed stages of the
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<typeStatus id="5483881AFFF40833FF136EF3FEDFFF45" box="[233,283,212,236]" pageId="4" pageNumber="2246">type</typeStatus>
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species
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<taxonomicName id="4C384D3BFFF40833FE806EF4FD93FF45" authority="Ocaña & Çinar, 2018" authorityName="Ocaña & Çinar" authorityYear="2018" box="[378,599,211,236]" class="Insecta" family="Gracillariidae" genus="Marmara" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="2246" phylum="Arthropoda" rank="species" species="musculata" status="sp. nov.">
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<emphasis id="B94CEAAAFFF40833FE806EF4FD93FF45" box="[378,599,211,236]" italics="true" pageId="4" pageNumber="2246">Marmara musculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F57D1FFF40833FDA46EF3FD73FF45" box="[606,695,212,236]" pageId="4" pageNumber="2246" rank="species">sp. nov.</taxonomicNameLabel>
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, two well differentiated areas are formed along the body: the upper part has tentacles and pharynx, constituting a bud-like structure; the stalk part includes the rest of the column, forming a long and cord-like structure, which has never been recorded in any
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<taxonomicName id="4C384D3BFFF40833FD216F1BFC80FEFD" authorityName="Hertwig" authorityYear="1882" box="[731,836,316,340]" class="Anthozoa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="2246" phylum="Cnidaria" rank="order">Actiniaria</taxonomicName>
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before (see,
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<bibRefCitation id="EFA94B49FFF40833FC266F1BFBB8FEFD" author="Carlgren O" box="[988,1148,316,340]" pageId="4" pageNumber="2246" pagination="1 - 121" refId="ref14953" refString="Carlgren O. 1949. A Survey of the Ptychodactiaria, Corallimorpharia and Actiniaria. Kung Sven Vetens Handlingar. 1: 1 - 121." type="journal article" year="1949">Carlgren 1949</bibRefCitation>
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) (
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<figureCitation id="13032A3DFFF40833FF5D6F79FED7FEDF" box="[167,275,350,374]" captionStart="Figure 2" captionStartId="4.[160,225,1675,1698]" captionTargetBox="[162,1152,850,1644]" captionTargetId="figure-183@4.[162,1153,850,1644]" captionTargetPageId="4" captionText="Figure 2. Marmara musculata sp. nov. (a–d) Retracted and protruded fixed specimens showing characteristic colour pattern and external morphology. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4747344" httpUri="https://zenodo.org/record/4747344/files/figure.png" pageId="4" pageNumber="2246">Figures 2</figureCitation>
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(d) and 3(c)). This amazing differentiation is possibly due to the burrowing habit displayed by this new species. This morphology looks like an extraordinary adaptation to sandy bottoms to elongate the column and extend its mouth to ambient water while the base is still in the burrow. We failed to identify the scapulus, but cannot assure the absence of such structure due to the retraction stage of the material; scapulus is present in the other closely related genera (
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<emphasis id="B94CEAAAFFF40833FD486C2CFC1BFD8D" box="[690,991,523,548]" italics="true" pageId="4" pageNumber="2246">
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<taxonomicName id="4C384D3BFFF40833FD486C2CFC8FFD8D" authorityName="Carlgren" authorityYear="1943" box="[690,843,523,548]" class="Anthozoa" family="Sagartiidae" genus="Sagartianthus" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="2246" phylum="Cnidaria" rank="genus">Sagartianthus</taxonomicName>
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,
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<taxonomicName id="4C384D3BFFF40833FCA26C2CFC1BFD8D" authorityName="Stephenson" authorityYear="1918" box="[856,991,523,548]" class="Anthozoa" family="Sagartiidae" genus="Artemidactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="2246" phylum="Cnidaria" rank="genus">Artemidactis</taxonomicName>
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</emphasis>
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).
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</paragraph>
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</subSubSection>
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||
</treatment>
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||
</document> |