279 lines
57 KiB
XML
279 lines
57 KiB
XML
<document id="74FD4D0802D635A2FBC19CD6FDF901B4" ID-DOI="10.1093/zoolinnean/zlad076" ID-ISSN="0024-4082" ID-Zenodo-Dep="10470514" ID-ZooBank="9A4BC231-A3D0-4335-AFCB-8A99DEF3C953C" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1704719650067" checkinUser="plazi" docAuthor="Sánchez-Fenollosa, Sergio, Verdú, Francisco J. & Cobos, Alberto" docDate="2023" docId="039087FB39185138BF844C12FF28C47D" docLanguage="en" docName="zlad076.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad076" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Oblitosaurus bunnueli Sánchez-Fenollosa & Verdú & Cobos 2023, gen. et sp. nov." docType="treatment" docVersion="3" lastPageNumber="1021" masterDocId="FFA9FF83391A5130BC464B22FFC2C119" masterDocTitle="The largest ornithopod (Dinosauria: Ornithischia) from the Upper Jurassic of Europe sheds light on the evolutionary history of basal ankylopollexians" masterLastPageNumber="1033" masterPageNumber="1013" pageNumber="1015" updateTime="1705322194026" updateUser="guilherme">
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<mods:titleInfo id="D17187E838B51641BBDD31EF0D5BF3E4">
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<mods:title id="CC4A0FB8A5E9469C493A70D75151FEE2">The largest ornithopod (Dinosauria: Ornithischia) from the Upper Jurassic of Europe sheds light on the evolutionary history of basal ankylopollexians</mods:title>
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<mods:roleTerm id="58A7E1549B1B353E03B843E020E95BFF">Author</mods:roleTerm>
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<mods:namePart id="E7B5BA2C84926B1FBA992001157BD4B1">Sánchez-Fenollosa, Sergio</mods:namePart>
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<mods:affiliation id="3D1DF773F78F607A105E8F932CEDD8E3">Fundación Conjunto Paleontológico de Teruel-Dinópolis / Museo Aragonés de Paleontología, Av. Sagunto s / n, 44002, Teruel, Spain</mods:affiliation>
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<mods:nameIdentifier id="43F1799A193231814228CD7FE95718F7" type="email">sfenollosa@fundaciondinopolis.org</mods:nameIdentifier>
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<mods:roleTerm id="2230E088DA68E76E73F56CDEBD284D87">Author</mods:roleTerm>
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<mods:namePart id="1A638046DFD4621520371536669E21DE">Verdú, Francisco J.</mods:namePart>
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<mods:affiliation id="0172156BAF07B163DD99614FF31D3C8D">Fundación Conjunto Paleontológico de Teruel-Dinópolis / Museo Aragonés de Paleontología, Av. Sagunto s / n, 44002, Teruel, Spain</mods:affiliation>
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<mods:namePart id="A4BF4C325FB8D2104C81D37D15A06C5C">Cobos, Alberto</mods:namePart>
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<mods:affiliation id="4D5E66DF62ACA12E005FEFA313B2ACE8">Fundación Conjunto Paleontológico de Teruel-Dinópolis / Museo Aragonés de Paleontología, Av. Sagunto s / n, 44002, Teruel, Spain</mods:affiliation>
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<mods:typeOfResource id="60419A678551E7C58E2A1CF237A3B583">text</mods:typeOfResource>
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<mods:title id="011B9ACB7799925D6D4E4C33A96785C3">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="7F1EF0B54709B17735B688B8B48E8CBE">
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<mods:date id="2769B3FB45FBB03859A8D094B31B37B4">2023</mods:date>
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<mods:number id="CE1A22582767ED98A9933FAF6DDFEDC5">2023-07-18</mods:number>
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<mods:number id="6C189A9F671A4E5F5F766EAF5CDA32AC">199</mods:number>
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<mods:url id="A60DFC5AABFAB86C18967E4972B07878">http://dx.doi.org/10.1093/zoolinnean/zlad076</mods:url>
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<mods:classification id="D0B190F553DE675A3001F0CADD76CBF4">journal article</mods:classification>
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<mods:identifier id="F6E6C3AF37BA499E30393A3FD24A34E5" type="DOI">10.1093/zoolinnean/zlad076</mods:identifier>
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<mods:identifier id="3347290214DC22991A468F38CD602B13" type="ISSN">0024-4082</mods:identifier>
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<treatment id="039087FB39185138BF844C12FF28C47D" LSID="urn:lsid:plazi:treatment:039087FB39185138BF844C12FF28C47D" httpUri="http://treatment.plazi.org/id/039087FB39185138BF844C12FF28C47D" lastPageId="8" lastPageNumber="1021" pageId="2" pageNumber="1015">
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<subSubSection id="C323656639185132BF844C12FAFBC652" box="[962,1337,1840,1867]" pageId="2" pageNumber="1015" type="nomenclature">
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<paragraph id="8B8636ED39185132BF844C12FAFBC652" blockId="2.[962,1337,1840,1867]" box="[962,1337,1840,1867]" pageId="2" pageNumber="1015">
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<heading id="D0CE818139185132BF844C12FAFBC652" bold="true" box="[962,1337,1840,1867]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1015" reason="2">
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<emphasis id="B94DEAFF39185132BF844C12FAFBC652" bold="true" box="[962,1337,1840,1867]" pageId="2" pageNumber="1015">
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<taxonomicName id="4C394D6E39185132BF844C12FB5DC652" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[962,1183,1840,1867]" family="Ankylopollexia" genus="Oblitosaurus" kingdom="Animalia" order="Ornithischia" pageId="2" pageNumber="1015" rank="species" species="bunnueli" status="gen. et sp. nov.">
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<emphasis id="B94DEAFF39185132BF844C12FB5DC652" bold="true" box="[962,1183,1840,1867]" italics="true" pageId="2" pageNumber="1015">Oblitosaurus bunnueli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E578439185132B8E24C13FAFBC652" box="[1188,1337,1841,1867]" pageId="2" pageNumber="1015" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C323656639185132B8004C7FFB74C66E" box="[1094,1206,1885,1911]" pageId="2" pageNumber="1015" type="description">
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<paragraph id="8B8636ED39185132B8004C7FFB74C66E" blockId="2.[1094,1206,1885,1911]" box="[1094,1206,1885,1911]" pageId="2" pageNumber="1015">
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<emphasis id="B94DEAFF39185132B8004C7FFB74C66E" bold="true" box="[1094,1206,1885,1911]" pageId="2" pageNumber="1015">
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(
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<figureCitation id="13022A6839185132B8174C7FFB68C66E" box="[1105,1194,1885,1911]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStart-6="Figure 8" captionStartId-0="3.[113,178,1661,1685]" captionStartId-1="4.[130,195,1450,1474]" captionStartId-2="5.[112,177,1382,1406]" captionStartId-3="6.[129,194,987,1011]" captionStartId-4="7.[115,180,1342,1366]" captionStartId-5="8.[129,194,664,688]" captionStartId-6="9.[113,178,1534,1558]" captionTargetBox-0="[231,1342,144,1633]" captionTargetBox-1="[161,1441,144,1422]" captionTargetBox-2="[146,1426,144,1355]" captionTargetBox-3="[241,1361,144,959]" captionTargetBox-4="[274,1298,144,1314]" captionTargetBox-5="[321,1281,144,636]" captionTargetBox-6="[210,1362,144,1506]" captionTargetId-0="figure-79@3.[231,1342,144,1633]" captionTargetId-1="figure-195@4.[161,1441,144,1422]" captionTargetId-2="figure-303@5.[146,1426,144,1355]" captionTargetId-3="figure-592@6.[241,1361,144,959]" captionTargetId-4="figure-308@7.[274,1298,144,1314]" captionTargetId-5="figure-739@8.[321,1281,144,636]" captionTargetId-6="figure-205@9.[210,1362,144,1506]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Figure 2. Less dentary tooth (CPT-1440) (A–C) and right ungual pollex of the manus (CPT-1444) (D–I) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Dentary tooth in: A, lingual; B, distal; C, labial views.Ungual pollex in: D, dorsal; E, medial; F, proximal; G, ventral; H, lateral; and I, distal views.Abbreviations:ac, accessory ridge; gr, groove; pr, primary ridge; sr, secondary ridge.Dentary tooth scale bar = 1 cm. Ungual pollex scale bar = 5 cm." captionText-1="Figure 3. Less femur (MAP-8290) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm." captionText-2="Figure 4. Less tibia (MAP-8291) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Tibia in: A, lateral; B, anterior; C, medial; D, posterior; E, proximal; and F, distal views.Abbreviations:cc, cnemial crest; fc, fibular condyle; ic, inner condyle; lm, lateral malleolus; mm, medial malleolus.Scale bar = 20 cm." captionText-3="Figure 5. Less fibula (MAP-8292) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Fibula in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: cp, cranial process.Scale bar = 10 cm." captionText-4="Figure 6. Less astragalus (MAP-8293) (A–F) and less calcaneum (MAP-8294) (G–L) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Astragalus in: A, anterior; B, medial; C, dorsal; D, posterior; E, lateral; and F, ventral views. Calcaneum in: G, anterior; H, medial; I, dorsal; J, posterior; K, lateral; and L, ventral views. Abbreviations: ap, ascending process; ff,facet for distal end of fibula; flm, facet for lateral malleolus of tibia; fmm, facet for medial malleolus of tibia; ss, facet for lateral malleolus of tibia.Scale bar = 10 cm." captionText-5="Figure 7. Less metatarsal IV (MAP-8295) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity.Scale bar = 10 cm." captionText-6="Figure 8. Less pedal phalanges of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Phalanx 1 of digit II (MAP-8296) (A–E) in: A, dorsal; B, lateral; C, plantar; D, proximal; and E, distal views. Phalanx 3 of digit IV (MAP-8297) (F–J) in: F, dorsal; G, lateral; H, plantar; I, proximal; and J, distal views. Ungual phalanx of digit II (MAP-8298) (K–N) in: K, dorsal; L, plantar; M, lateral; and N, medial views. Ungual phalanx of digit IV (MAP-8299) (O–R) in: O, dorsal; P, plantar; º, lateral; and R, medial views. Abbreviations: clf, collateral ligament fovea; dl, dorsal lip; egr, extensor groove; fl, flap; lc, lateral condyle; mc, medial condyle; nvs, neurovascular sulci. Scale bar = 10 cm." figureDoi-0="http://doi.org/10.5281/zenodo.10470518" figureDoi-1="http://doi.org/10.5281/zenodo.10470520" figureDoi-2="http://doi.org/10.5281/zenodo.10470524" figureDoi-3="http://doi.org/10.5281/zenodo.10470526" figureDoi-4="http://doi.org/10.5281/zenodo.10470528" figureDoi-5="http://doi.org/10.5281/zenodo.10470530" figureDoi-6="http://doi.org/10.5281/zenodo.10470532" httpUri-0="https://zenodo.org/record/10470518/files/figure.png" httpUri-1="https://zenodo.org/record/10470520/files/figure.png" httpUri-2="https://zenodo.org/record/10470524/files/figure.png" httpUri-3="https://zenodo.org/record/10470526/files/figure.png" httpUri-4="https://zenodo.org/record/10470528/files/figure.png" httpUri-5="https://zenodo.org/record/10470530/files/figure.png" httpUri-6="https://zenodo.org/record/10470532/files/figure.png" pageId="2" pageNumber="1015">Figs 2–8</figureCitation>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323656639185132BF7F4CBCFA02C6CC" pageId="2" pageNumber="1015" type="etymology">
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<paragraph id="8B8636ED39185132BF7F4CBCFA02C6CC" blockId="2.[825,1474,1950,2005]" pageId="2" pageNumber="1015">
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<emphasis id="B94DEAFF39185132BF7F4CBCFC6AC6AF" box="[825,936,1950,1974]" italics="true" pageId="2" pageNumber="1015">Etymology:</emphasis>
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The specific name
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<taxonomicName id="4C394D6E39185132B8314CBCFB09C6AF" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[1143,1227,1950,1974]" genus="Oblitosaurus" pageId="2" pageNumber="1015" rank="species" species="bunnueli">
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<emphasis id="B94DEAFF39185132B8314CBCFB09C6AF" box="[1143,1227,1950,1974]" italics="true" pageId="2" pageNumber="1015">bunnueli</emphasis>
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</taxonomicName>
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honours Luis Buñuel, a prestigious Spanish film director born in the province of Teruel.
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</paragraph>
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</subSubSection>
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<caption id="DF46666539195133BC374D5FFDFFC7F0" ID-DOI="http://doi.org/10.5281/zenodo.10470518" ID-Zenodo-Dep="10470518" httpUri="https://zenodo.org/record/10470518/files/figure.png" pageId="3" pageNumber="1016" startId="3.[113,178,1661,1685]" targetBox="[231,1342,144,1633]" targetPageId="3" targetType="figure">
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<paragraph id="8B8636ED39195133BC374D5FFDFFC7F0" blockId="3.[113,1445,1661,1769]" pageId="3" pageNumber="1016">
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<emphasis id="B94DEAFF39195133BC374D5FFF0BC78C" bold="true" box="[113,201,1661,1685]" pageId="3" pageNumber="1016">Figure 2.</emphasis>
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Less dentary tooth (CPT-1440) (A–C) and right ungual pollex of the manus (CPT-1444) (D–I) of the holotype specimen of
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<taxonomicName id="4C394D6E39195133BC374DBBFEFAC7A8" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[113,312,1689,1713]" genus="Oblitosaurus" pageId="3" pageNumber="1016" rank="species" species="bunnueli" status="gen. et sp. nov">
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<emphasis id="B94DEAFF39195133BC374DBBFEFAC7A8" box="[113,312,1689,1713]" italics="true" pageId="3" pageNumber="1016">Oblitosaurus bunnueli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E578439195133BD7B4DB8FE03C7A8" box="[317,449,1690,1714]" pageId="3" pageNumber="1016" rank="species">gen. et sp. nov</taxonomicNameLabel>
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.. Dentary tooth in: A, lingual; B, distal; C, labial views. Ungual pollex in: D, dorsal; E, medial; F, proximal; G, ventral; H, lateral; and I, distal views. Abbreviations: ac, accessory ridge; gr, groove; pr, primary ridge; sr, secondary ridge. Dentary tooth scale bar = 1 cm. Ungual pollex scale bar = 5 cm.
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</paragraph>
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</caption>
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<subSubSection id="C323656639195134BC374C06FE5AC784" lastPageId="4" lastPageNumber="1017" pageId="3" pageNumber="1016" type="materials_examined">
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<paragraph id="8B8636ED39195133BC374C06FB33C642" blockId="3.[113,763,1828,1977]" lastBlockId="3.[810,1459,1828,1883]" pageId="3" pageNumber="1016">
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<materialsCitation id="3B513CB039195133BC374C06FB33C642" pageId="3" pageNumber="1016" specimenCount="1" typeStatus="holotype">
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<emphasis id="B94DEAFF39195133BC374C06FF13C625" box="[113,209,1828,1852]" italics="true" pageId="3" pageNumber="1016">
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<typeStatus id="5482884F39195133BC374C06FF0EC625" box="[113,204,1828,1852]" pageId="3" pageNumber="1016" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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A dentary tooth (CPT-1440), a right ungual pollex from manus (CPT-1444), and an almost complete less hindlimb: femur (MAP-8290), tibia (MAP-8291), fibula (MAP-8292), astragalus (MAP-8293), calcaneum (MAP-8294), metatarsal IV (MAP-8295), phalanx 1 of digit II (MAP-8296), phalanx 3 of digit IV (MAP-8297), ungual phalanx of digit II (MAP-8298), and ungual phalanx of digit IV (MAP-8299).
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</materialsCitation>
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</paragraph>
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<paragraph id="8B8636ED39195134BF6C4CA0FE5AC784" blockId="3.[810,1459,1922,1978]" lastBlockId="4.[129,777,1606,1693]" lastPageId="4" lastPageNumber="1017" pageId="3" pageNumber="1016">
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<emphasis id="B94DEAFF39195133BF6C4CA0FBE6C683" box="[810,1060,1922,1946]" italics="true" pageId="3" pageNumber="1016">Locality and horizon:</emphasis>
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Barrihonda-El Humero (RD-10) site in the municipality of Riodeva, province of Teruel,
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<collectingRegion id="49FDF80F391E5134BCC74D64FF0CC747" box="[129,206,1606,1630]" country="Spain" name="Aragon" pageId="4" pageNumber="1017">Aragón</collectingRegion>
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,
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<collectingCountry id="F32E767D391E5134BCA54D64FEDDC747" box="[227,287,1606,1630]" name="Spain" pageId="4" pageNumber="1017">Spain</collectingCountry>
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. South-Iberian Basin, Villar del Arzobispo Formation, Riodeva Facies Association, Upper Jurassic (upper Kimmeridgian–Tithonian).
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</paragraph>
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</subSubSection>
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<caption id="DF466665391E5134BCC44E88FF38C70F" ID-DOI="http://doi.org/10.5281/zenodo.10470520" ID-Zenodo-Dep="10470520" httpUri="https://zenodo.org/record/10470520/files/figure.png" pageId="4" pageNumber="1017" startId="4.[130,195,1450,1474]" targetBox="[161,1441,144,1422]" targetPageId="4" targetType="figure">
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<paragraph id="8B8636ED391E5134BCC44E88FF38C70F" blockId="4.[129,1409,1450,1559]" pageId="4" pageNumber="1017">
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<emphasis id="B94DEAFF391E5134BCC44E88FF18C4DB" bold="true" box="[130,218,1450,1474]" pageId="4" pageNumber="1017">Figure 3.</emphasis>
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Less femur (MAP-8290) of the holotype specimen of
|
||
<taxonomicName id="4C394D6E391E5134BE914E88FC5CC4DB" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[727,926,1450,1474]" genus="Oblitosaurus" pageId="4" pageNumber="1017" rank="species" species="bunnueli" status="gen. et sp. nov">
|
||
<emphasis id="B94DEAFF391E5134BE914E88FC5CC4DB" box="[727,926,1450,1474]" italics="true" pageId="4" pageNumber="1017">Oblitosaurus bunnueli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E5784391E5134BFE74E89FBE7C4DB" box="[929,1061,1451,1475]" pageId="4" pageNumber="1017" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3236566391E5134BCC74DEAFC72C764" pageId="4" pageNumber="1017" type="diagnosis">
|
||
<paragraph id="8B8636ED391E5134BCC74DEAFC72C764" blockId="4.[129,778,1735,1978]" lastBlockId="4.[825,1474,1606,1661]" pageId="4" pageNumber="1017">
|
||
<emphasis id="B94DEAFF391E5134BCC74DEAFF25C7C6" box="[129,231,1736,1759]" italics="true" pageId="4" pageNumber="1017">Diagnosis:</emphasis>
|
||
Ankylopollexian ornithopod diagnosed by the following autapomorphies (* when dubious due to taphonomic distortions): femur with (i*) a wide and robust lesser trochanter that almost covers the lateral surface of the greater trochanter; and (ii) a sub-circular distal medial condyle; with (iii) a pronounced and narrow lateromedial ridge on its ventral surface; tibia with (iv) an internal condyle projected dorsally above the dorsal margin of the cnemial crest; and metatarsal IV with (v) sub-triangular (or ellipsoid) morphology of the proximal articulation facet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236566391E5138B80E4D83FF28C47D" lastPageId="8" lastPageNumber="1021" pageId="4" pageNumber="1017" type="description">
|
||
<paragraph id="8B8636ED391E5134B80E4D83FB76C7A3" blockId="4.[1096,1204,1697,1722]" box="[1096,1204,1697,1722]" pageId="4" pageNumber="1017">
|
||
<emphasis id="B94DEAFF391E5134B80E4D83FB76C7A3" box="[1096,1204,1697,1722]" italics="true" pageId="4" pageNumber="1017">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391E5135BF7F4DEAFD20C7E0" blockId="4.[825,1475,1735,1979]" lastBlockId="5.[113,763,1511,1785]" lastPageId="5" lastPageNumber="1018" pageId="4" pageNumber="1017">
|
||
<emphasis id="B94DEAFF391E5134BF7F4DEAFC5FC7C6" box="[825,925,1736,1759]" italics="true" pageId="4" pageNumber="1017">Dentition:</emphasis>
|
||
CPT-1440 (
|
||
<figureCitation id="13022A68391E5134B86D4DE5FB54C7C6" box="[1067,1174,1735,1759]" captionStart="Figure 2" captionStartId="3.[113,178,1661,1685]" captionTargetBox="[231,1342,144,1633]" captionTargetId="figure-79@3.[231,1342,144,1633]" captionTargetPageId="3" captionText="Figure 2. Less dentary tooth (CPT-1440) (A–C) and right ungual pollex of the manus (CPT-1444) (D–I) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Dentary tooth in: A, lingual; B, distal; C, labial views.Ungual pollex in: D, dorsal; E, medial; F, proximal; G, ventral; H, lateral; and I, distal views.Abbreviations:ac, accessory ridge; gr, groove; pr, primary ridge; sr, secondary ridge.Dentary tooth scale bar = 1 cm. Ungual pollex scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10470518" httpUri="https://zenodo.org/record/10470518/files/figure.png" pageId="4" pageNumber="1017">Fig. 2A–C</figureCitation>
|
||
) is a less dentary tooth. This well-preserved tooth is strongly labially curved. The crown is shield-shaped and the enamel is restricted to the lingual surface. The primary ridge is posteriorly offset and bifurcates toward the root. Two incomplete and less prominent secondary ridges are positioned mesially to the primary ridge. The one nearest to the primary ridge also bifurcates at the end of the crown. In addition, there are two faint and shorter accessory ridges located distally and mesially, respectively. There are no ridges on the nonenamelled labial face of the crown. The carinae present tongueshaped denticles with smooth margins from mid-crown height to apex. There is a worn facet, almost restricted to the labial surface. There is no cingulum separating the crown and root on the lingual surface. The root is incomplete and circular in shape in the section near the tip. There is a shallow depression on each side of the teeth (more distally evident) that extends from the root to the crown to accommodate contiguous growing teeth.
|
||
</paragraph>
|
||
<caption id="DF466665391F5135BC364E44FE09C4AF" ID-DOI="http://doi.org/10.5281/zenodo.10470524" ID-Zenodo-Dep="10470524" httpUri="https://zenodo.org/record/10470524/files/figure.png" pageId="5" pageNumber="1018" startId="5.[112,177,1382,1406]" targetBox="[146,1426,144,1355]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8636ED391F5135BC364E44FE09C4AF" blockId="5.[112,1443,1382,1463]" pageId="5" pageNumber="1018">
|
||
<emphasis id="B94DEAFF391F5135BC364E44FF0AC466" bold="true" box="[112,200,1382,1407]" pageId="5" pageNumber="1018">Figure 4.</emphasis>
|
||
Less tibia (MAP-8291) of the holotype specimen of
|
||
<taxonomicName id="4C394D6E391F5135BEF14E45FCBCC466" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[695,894,1383,1407]" genus="Oblitosaurus" pageId="5" pageNumber="1018" rank="species" species="bunnueli" status="gen. et sp. nov">
|
||
<emphasis id="B94DEAFF391F5135BEF14E45FCBCC466" box="[695,894,1383,1407]" italics="true" pageId="5" pageNumber="1018">Oblitosaurus bunnueli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E5784391F5135BFC24E45FBCAC467" box="[900,1032,1383,1407]" pageId="5" pageNumber="1018" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. Tibia in: A, lateral; B, anterior; C, medial; D, posterior; E, proximal; and F, distal views. Abbreviations: cc, cnemial crest; fc, fibular condyle; ic, inner condyle; lm, lateral malleolus; mm, medial malleolus. Scale bar = 20 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636ED391F5135BC374C38FA76C707" blockId="5.[113,763,1818,1968]" lastBlockId="5.[810,1460,1510,1566]" pageId="5" pageNumber="1018">
|
||
<emphasis id="B94DEAFF391F5135BC374C38FEEFC62B" box="[113,301,1818,1842]" italics="true" pageId="5" pageNumber="1018">Manual phalanges:</emphasis>
|
||
The right ungual phalanx of the digit I CPT-1444 (
|
||
<figureCitation id="13022A68391F5135BCFA4C18FEE6C64B" box="[188,292,1850,1874]" captionStart="Figure 2" captionStartId="3.[113,178,1661,1685]" captionTargetBox="[231,1342,144,1633]" captionTargetId="figure-79@3.[231,1342,144,1633]" captionTargetPageId="3" captionText="Figure 2. Less dentary tooth (CPT-1440) (A–C) and right ungual pollex of the manus (CPT-1444) (D–I) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Dentary tooth in: A, lingual; B, distal; C, labial views.Ungual pollex in: D, dorsal; E, medial; F, proximal; G, ventral; H, lateral; and I, distal views.Abbreviations:ac, accessory ridge; gr, groove; pr, primary ridge; sr, secondary ridge.Dentary tooth scale bar = 1 cm. Ungual pollex scale bar = 5 cm." figureDoi="http://doi.org/10.5281/zenodo.10470518" httpUri="https://zenodo.org/record/10470518/files/figure.png" pageId="5" pageNumber="1018">Fig. 2D–I</figureCitation>
|
||
) is robust, short, sub-conical, and pointed (despite the tip is not being preserved). Furthermore, it has a sub-circular and concave proximal articular facet, which suggests a certain degree of mobility at the articulation with the proximal phalanx. There are two nail grooves: a curved and deep medial groove, and a much fainter (almost inappreciable) lateral groove.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391F5136BF6C4D62FD40C468" blockId="5.[810,1460,1599,1968]" lastBlockId="6.[129,778,1087,1393]" lastPageId="6" lastPageNumber="1019" pageId="5" pageNumber="1018">
|
||
<emphasis id="B94DEAFF391F5135BF6C4D62FCADC74E" box="[810,879,1600,1623]" italics="true" pageId="5" pageNumber="1018">Femur:</emphasis>
|
||
MAP-8290 (
|
||
<figureCitation id="13022A68391F5135BFB94D1DFBFEC74E" box="[1023,1084,1599,1623]" captionStart="Figure 3" captionStartId="4.[130,195,1450,1474]" captionTargetBox="[161,1441,144,1422]" captionTargetId="figure-195@4.[161,1441,144,1422]" captionTargetPageId="4" captionText="Figure 3. Less femur (MAP-8290) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470520" httpUri="https://zenodo.org/record/10470520/files/figure.png" pageId="5" pageNumber="1018">Fig. 3</figureCitation>
|
||
) is an almost entire less femur but lacking part of the diaphysis below the fourth trochanter. It is robust and slightly curved posteriorly in medial/lateral view. Both epiphyses are lateromedially expanded relative to the diaphysis. The femoral head is bulbous and broad with a shallow concavity on the posterior face, separated from the greater trochanter by a wide depression. The greater trochanter is dorsally convex and projects dorsolaterally above the lesser trochanter. The spike-like lesser trochanter is wide and robust, oblique in dorsal view, and covers almost the entire lateral surface of the greater trochanter (
|
||
<figureCitation id="13022A68391F5135BF724C5AFCBCC688" box="[820,894,1912,1937]" captionStart="Figure 3" captionStartId="4.[130,195,1450,1474]" captionTargetBox="[161,1441,144,1422]" captionTargetId="figure-195@4.[161,1441,144,1422]" captionTargetPageId="4" captionText="Figure 3. Less femur (MAP-8290) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470520" httpUri="https://zenodo.org/record/10470520/files/figure.png" pageId="5" pageNumber="1018">Fig. 3D</figureCitation>
|
||
). Moreover, it is separated from the greater trochanter by a wide cless. The fourth trochanter is fragmented and incomplete at the distal end; however, the preserved part is large, sub-triangular, and located in the proximal-midpart of the diaphysis. There is a sharp but low ridge running diagonally from the lesser trochanter to the distal end, but it does not reach the medial condyle. At the distal end, the medial and lateral condyles are separated by a deep U-shaped intercondylar extensor groove. The medial condyle is sub-circular in ventral view (
|
||
<figureCitation id="13022A68391C5136BD874FD9FDCAC40A" box="[449,520,1275,1299]" captionStart="Figure 3" captionStartId="4.[130,195,1450,1474]" captionTargetBox="[161,1441,144,1422]" captionTargetId="figure-195@4.[161,1441,144,1422]" captionTargetPageId="4" captionText="Figure 3. Less femur (MAP-8290) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470520" httpUri="https://zenodo.org/record/10470520/files/figure.png" pageId="6" pageNumber="1019">Fig. 3F</figureCitation>
|
||
), with no protrusion extending to the intercondylar flexor groove, and has a marked but narrow transversal ridge, which projects ventrally (
|
||
<figureCitation id="13022A68391C5136BEC24E1BFD09C448" box="[644,715,1337,1361]" captionStart="Figure 3" captionStartId="4.[130,195,1450,1474]" captionTargetBox="[161,1441,144,1422]" captionTargetId="figure-195@4.[161,1441,144,1422]" captionTargetPageId="4" captionText="Figure 3. Less femur (MAP-8290) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Femur in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: egr, extensor groove; fgr, flexor groove; ffl, femoral head; gt, greater trochanter; lc, lateral condyle; lt, lesser trochanter; mc, medial condyle; mcr, medial condylar ridge; r, ridge; 4t, fourth trochanter. Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470520" httpUri="https://zenodo.org/record/10470520/files/figure.png" pageId="6" pageNumber="1019">Fig. 3B</figureCitation>
|
||
). The posterior half of the lateral condyle is not preserved.
|
||
</paragraph>
|
||
<caption id="DF466665391C5136BCC748F9FBA7C517" ID-DOI="http://doi.org/10.5281/zenodo.10470526" ID-Zenodo-Dep="10470526" httpUri="https://zenodo.org/record/10470526/files/figure.png" pageId="6" pageNumber="1019" startId="6.[129,194,987,1011]" targetBox="[241,1361,144,959]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8B8636ED391C5136BCC748F9FBA7C517" blockId="6.[129,1402,987,1039]" pageId="6" pageNumber="1019">
|
||
<emphasis id="B94DEAFF391C5136BCC748F9FF1AC2EA" bold="true" box="[129,216,987,1011]" pageId="6" pageNumber="1019">Figure 5.</emphasis>
|
||
Less fibula (MAP-8292) of the holotype specimen of
|
||
<taxonomicName id="4C394D6E391C5136BE9448F9FC5BC2EA" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[722,921,987,1011]" genus="Oblitosaurus" pageId="6" pageNumber="1019" rank="species" species="bunnueli" status="gen. et sp. nov">
|
||
<emphasis id="B94DEAFF391C5136BE9448F9FC5BC2EA" box="[722,921,987,1011]" italics="true" pageId="6" pageNumber="1019">Oblitosaurus bunnueli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E5784391C5136BFD848F9FBE0C2EB" box="[926,1058,987,1011]" pageId="6" pageNumber="1019" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. Fibula in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: cp, cranial process. Scale bar = 10 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636ED391C5136BCC74EB0FB92C42B" blockId="6.[128,778,1426,1982]" lastBlockId="6.[825,1474,1086,1330]" pageId="6" pageNumber="1019">
|
||
<emphasis id="B94DEAFF391C5136BCC74EB0FF7EC4B3" box="[129,188,1426,1450]" italics="true" pageId="6" pageNumber="1019">Tibia:</emphasis>
|
||
MAP-8291 (
|
||
<figureCitation id="13022A68391C5136BD034EB0FEB9C4B3" box="[325,379,1426,1450]" captionStart="Figure 4" captionStartId="5.[112,177,1382,1406]" captionTargetBox="[146,1426,144,1355]" captionTargetId="figure-303@5.[146,1426,144,1355]" captionTargetPageId="5" captionText="Figure 4. Less tibia (MAP-8291) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Tibia in: A, lateral; B, anterior; C, medial; D, posterior; E, proximal; and F, distal views.Abbreviations:cc, cnemial crest; fc, fibular condyle; ic, inner condyle; lm, lateral malleolus; mm, medial malleolus.Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470524" httpUri="https://zenodo.org/record/10470524/files/figure.png" pageId="6" pageNumber="1019">Fig. 4</figureCitation>
|
||
) is an almost complete less tibia, shorter than the femur. The typical rotation present in the proximal end of the tibia in ornithopods cannot be observed here probably due to taphonomic reasons. The anterior, posterior, and dorsal surfaces of the proximal epiphysis are slightly eroded, but the cortical bone can be observed. However, the medial and lateral surfaces are eroded; therefore, its morphology should be considered with caution. The proximal epiphysis is slightly expanded both anteroposteriorly and lateromedially (
|
||
<figureCitation id="13022A68391C5136BDB84DAEFD81C7BC" box="[510,579,1676,1701]" captionStart="Figure 4" captionStartId="5.[112,177,1382,1406]" captionTargetBox="[146,1426,144,1355]" captionTargetId="figure-303@5.[146,1426,144,1355]" captionTargetPageId="5" captionText="Figure 4. Less tibia (MAP-8291) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Tibia in: A, lateral; B, anterior; C, medial; D, posterior; E, proximal; and F, distal views.Abbreviations:cc, cnemial crest; fc, fibular condyle; ic, inner condyle; lm, lateral malleolus; mm, medial malleolus.Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470524" httpUri="https://zenodo.org/record/10470524/files/figure.png" pageId="6" pageNumber="1019">Fig. 4E</figureCitation>
|
||
). The cnemial crest is short, curved, robust, and laterally separated from the fibular (or lateral) condyle by a gentle concave margin. The eroded fibular condyle is located in the median position. The medial edge of the proximal end is gently sinuous. Moreover, the internal condyle projects dorsally above the cnemial crest (
|
||
<figureCitation id="13022A68391C5136BE254C0BFD09C658" box="[611,715,1833,1857]" captionStart="Figure 4" captionStartId="5.[112,177,1382,1406]" captionTargetBox="[146,1426,144,1355]" captionTargetId="figure-303@5.[146,1426,144,1355]" captionTargetPageId="5" captionText="Figure 4. Less tibia (MAP-8291) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Tibia in: A, lateral; B, anterior; C, medial; D, posterior; E, proximal; and F, distal views.Abbreviations:cc, cnemial crest; fc, fibular condyle; ic, inner condyle; lm, lateral malleolus; mm, medial malleolus.Scale bar = 20 cm." figureDoi="http://doi.org/10.5281/zenodo.10470524" httpUri="https://zenodo.org/record/10470524/files/figure.png" pageId="6" pageNumber="1019">Fig. 4B, D</figureCitation>
|
||
). The diaphysis is large, presumably cylindrical, and expands drastically toward the distal end. The distal end is much wider lateromedially than the proximal end anteroposteriorly. In particular, the lateral malleolus is wider and projects more distally than the medial one.
|
||
<emphasis id="B94DEAFF391C5136BF7F4F1CFCBDC54F" box="[825,895,1086,1110]" italics="true" pageId="6" pageNumber="1019">Fibula:</emphasis>
|
||
MAP-8292 (
|
||
<figureCitation id="13022A68391C5136B8484F1CFB85C54F" box="[1038,1095,1086,1110]" captionStart="Figure 5" captionStartId="6.[129,194,987,1011]" captionTargetBox="[241,1361,144,959]" captionTargetId="figure-592@6.[241,1361,144,959]" captionTargetPageId="6" captionText="Figure 5. Less fibula (MAP-8292) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Fibula in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: cp, cranial process.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470526" httpUri="https://zenodo.org/record/10470526/files/figure.png" pageId="6" pageNumber="1019">Fig. 5</figureCitation>
|
||
) is a slender less fibula that is shorter than the tibia. Both the proximal and distal regions are expanded, especially the proximal region, which has an angular cranial process. The fibula is twisted along its length. The lateral surface of the proximal end is slightly convex, whereas the medial surface is concave to receive the fibular condyle of the tibia. The dorsal margin of the fibula slopes slightly from posteroventral to anterodorsal in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391C5136BF7F4E71FB6DC79D" blockId="6.[825,1474,1363,1669]" pageId="6" pageNumber="1019">
|
||
<emphasis id="B94DEAFF391C5136BF7F4E71FC65C472" box="[825,935,1363,1387]" italics="true" pageId="6" pageNumber="1019">Astragalus:</emphasis>
|
||
MAP-8293 (
|
||
<figureCitation id="13022A68391C5136B8704E71FB59C472" box="[1078,1179,1363,1387]" captionStart="Figure 6" captionStartId="7.[115,180,1342,1366]" captionTargetBox="[274,1298,144,1314]" captionTargetId="figure-308@7.[274,1298,144,1314]" captionTargetPageId="7" captionText="Figure 6. Less astragalus (MAP-8293) (A–F) and less calcaneum (MAP-8294) (G–L) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Astragalus in: A, anterior; B, medial; C, dorsal; D, posterior; E, lateral; and F, ventral views. Calcaneum in: G, anterior; H, medial; I, dorsal; J, posterior; K, lateral; and L, ventral views. Abbreviations: ap, ascending process; ff,facet for distal end of fibula; flm, facet for lateral malleolus of tibia; fmm, facet for medial malleolus of tibia; ss, facet for lateral malleolus of tibia.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470528" httpUri="https://zenodo.org/record/10470528/files/figure.png" pageId="6" pageNumber="1019">Fig. 6A–F</figureCitation>
|
||
) is a complete less astragalus with a general sub-triangular shape. The dorsal surface has two slightly concave facets for the medial and lateral malleoli of the tibia. These facets are separated by a ridge, and the medial facet is wider both anteroposteriorly and lateromedially than the lateral facet. A prominent triangular ascending process is observed on the posterior margin (
|
||
<figureCitation id="13022A68391C5136B8584D2DFBABC73E" box="[1054,1129,1551,1575]" captionStart="Figure 6" captionStartId="7.[115,180,1342,1366]" captionTargetBox="[274,1298,144,1314]" captionTargetId="figure-308@7.[274,1298,144,1314]" captionTargetPageId="7" captionText="Figure 6. Less astragalus (MAP-8293) (A–F) and less calcaneum (MAP-8294) (G–L) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Astragalus in: A, anterior; B, medial; C, dorsal; D, posterior; E, lateral; and F, ventral views. Calcaneum in: G, anterior; H, medial; I, dorsal; J, posterior; K, lateral; and L, ventral views. Abbreviations: ap, ascending process; ff,facet for distal end of fibula; flm, facet for lateral malleolus of tibia; fmm, facet for medial malleolus of tibia; ss, facet for lateral malleolus of tibia.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470528" httpUri="https://zenodo.org/record/10470528/files/figure.png" pageId="6" pageNumber="1019">Fig. 6C</figureCitation>
|
||
). Both the anterior and posterior margins do not face each other enclosing articular facets, so the astragalus is very open in dorsal view. In contrast to the dorsal surface, the ventral surface is convex.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391C5136BF7F4D8EFAAEC6A6" blockId="6.[825,1475,1708,1983]" pageId="6" pageNumber="1019">
|
||
<emphasis id="B94DEAFF391C5136BF7F4D8EFC6DC7DD" box="[825,943,1708,1732]" italics="true" pageId="6" pageNumber="1019">
|
||
<taxonomicName id="4C394D6E391C5136BF7F4D8EFC6BC7DD" authorityName="Townes" authorityYear="1971" box="[825,937,1708,1732]" class="Insecta" family="Ichneumonidae" genus="Calcaneum" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="1019" phylum="Arthropoda" rank="genus">Calcaneum</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
MAP-8294 (
|
||
<figureCitation id="13022A68391C5136B87D4D8EFB60C7DD" box="[1083,1186,1708,1732]" captionStart="Figure 6" captionStartId="7.[115,180,1342,1366]" captionTargetBox="[274,1298,144,1314]" captionTargetId="figure-308@7.[274,1298,144,1314]" captionTargetPageId="7" captionText="Figure 6. Less astragalus (MAP-8293) (A–F) and less calcaneum (MAP-8294) (G–L) of the holotype specimen of Oblitosaurus bunnueligen. et sp. nov.. Astragalus in: A, anterior; B, medial; C, dorsal; D, posterior; E, lateral; and F, ventral views. Calcaneum in: G, anterior; H, medial; I, dorsal; J, posterior; K, lateral; and L, ventral views. Abbreviations: ap, ascending process; ff,facet for distal end of fibula; flm, facet for lateral malleolus of tibia; fmm, facet for medial malleolus of tibia; ss, facet for lateral malleolus of tibia.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470528" httpUri="https://zenodo.org/record/10470528/files/figure.png" pageId="6" pageNumber="1019">Fig. 6G–L</figureCitation>
|
||
) is a complete less calcaneus with an overall hemispherical shape. The dorsal surface is divided into fibular and tibial facets, separated from each other by a pronounced oblique lateromedial ridge. The fibular facet is larger than the tibial facet. The tibial facet has a strongly concave surface and the fibular facet has an almost straight one. Both facets present straight margins in lateral view. There is a depression on the lateral side, with a gentle ridge separating it into two halves. The anteroventral surface is markedly convex and thick.
|
||
</paragraph>
|
||
<caption id="DF466665391D5137BC354E1CFB5DC4B3" ID-DOI="http://doi.org/10.5281/zenodo.10470528" ID-Zenodo-Dep="10470528" httpUri="https://zenodo.org/record/10470528/files/figure.png" pageId="7" pageNumber="1020" startId="7.[115,180,1342,1366]" targetBox="[274,1298,144,1314]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B8636ED391D5137BC354E1CFA55C46B" blockId="7.[113,1439,1342,1450]" pageId="7" pageNumber="1020">
|
||
<emphasis id="B94DEAFF391D5137BC354E1CFF09C44F" bold="true" box="[115,203,1342,1366]" pageId="7" pageNumber="1020">Figure 6.</emphasis>
|
||
Less astragalus (MAP-8293) (A–F) and less calcaneum (MAP-8294) (G–L) of the holotype specimen of
|
||
<taxonomicName id="4C394D6E391D5137B8ED4E1CFAB0C44F" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[1195,1394,1342,1366]" genus="Oblitosaurus" pageId="7" pageNumber="1020" rank="species" species="bunnueli" status="gen. et sp. nov">
|
||
<emphasis id="B94DEAFF391D5137B8ED4E1CFAB0C44F" box="[1195,1394,1342,1366]" italics="true" pageId="7" pageNumber="1020">Oblitosaurus bunnueli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E5784391D5137B9334E1DFF0BC46B" pageId="7" pageNumber="1020" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. Astragalus in: A, anterior; B, medial; C, dorsal; D, posterior; E, lateral; and F, ventral views.
|
||
<taxonomicName id="4C394D6E391D5137B8774E78FB5FC46B" authorityName="Townes" authorityYear="1971" box="[1073,1181,1370,1394]" class="Insecta" family="Ichneumonidae" genus="Calcaneum" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="1025" phylum="Arthropoda" rank="genus">Calcaneum</taxonomicName>
|
||
in: G, anterior; H, medial;
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391D5137BC374E55FB5DC4B3" blockId="7.[113,1439,1342,1450]" pageId="7" pageNumber="1020">I, dorsal; J, posterior; K, lateral; and L, ventral views. Abbreviations: ap, ascending process; ff, facet for distal end of fibula; flm, facet for lateral malleolus of tibia; fmm, facet for medial malleolus of tibia; ss, facet for lateral malleolus of tibia. Scale bar = 10 cm.</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636ED391D5137BC374EF8FC6DC708" blockId="7.[113,763,1498,1961]" lastBlockId="7.[810,1459,1498,1553]" pageId="7" pageNumber="1020">
|
||
<emphasis id="B94DEAFF391D5137BC374EF8FEC2C4EB" box="[113,256,1498,1522]" italics="true" pageId="7" pageNumber="1020">Metatarsal IV:</emphasis>
|
||
MAP-8295 (
|
||
<figureCitation id="13022A68391D5137BDCB4EF8FE07C4EB" box="[397,453,1498,1522]" captionStart="Figure 7" captionStartId="8.[129,194,664,688]" captionTargetBox="[321,1281,144,636]" captionTargetId="figure-739@8.[321,1281,144,636]" captionTargetPageId="8" captionText="Figure 7. Less metatarsal IV (MAP-8295) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470530" httpUri="https://zenodo.org/record/10470530/files/figure.png" pageId="7" pageNumber="1020">Fig. 7</figureCitation>
|
||
) comprises a long and slender less metatarsal IV. In anterior and posterior views, the medial margin is convex, forming the medial tuberosity, while the lateral margin is concave. Both the proximal and distal ends are expanded. The former has a concave, anteroposteriorly compressed, and sub-triangular articular facet (
|
||
<figureCitation id="13022A68391D5137BE0B4D55FD59C797" box="[589,667,1655,1679]" captionStart="Figure 7" captionStartId="8.[129,194,664,688]" captionTargetBox="[321,1281,144,636]" captionTargetId="figure-739@8.[321,1281,144,636]" captionTargetPageId="8" captionText="Figure 7. Less metatarsal IV (MAP-8295) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470530" httpUri="https://zenodo.org/record/10470530/files/figure.png" pageId="7" pageNumber="1020">Fig. 7E</figureCitation>
|
||
) with a lateromedial width much greater than the anteroposterior, so the lateral edge diverges from the medial one in anterior and posterior views (
|
||
<figureCitation id="13022A68391D5137BD594DF7FE4AC7F4" box="[287,392,1749,1773]" captionStart="Figure 7" captionStartId="8.[129,194,664,688]" captionTargetBox="[321,1281,144,636]" captionTargetId="figure-739@8.[321,1281,144,636]" captionTargetPageId="8" captionText="Figure 7. Less metatarsal IV (MAP-8295) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470530" httpUri="https://zenodo.org/record/10470530/files/figure.png" pageId="7" pageNumber="1020">Fig. 7A, C</figureCitation>
|
||
). Moreover, there is a concave facet in the medial margin for articulating with the metatarsal III. In lateral and medial views, the anterior margin of the diaphysis is straight, but the posterior margin is concave. The distal end is sub-circular in lateral and medial views, but it is sub-rectangular in distal view, being longer than wide. Both the medial and lateral collateral ligament foveae at the distal end are deeply depressed, especially on the lateral side (
|
||
<figureCitation id="13022A68391D5137B82A4EF8FB75C4EB" box="[1132,1207,1498,1522]" captionStart="Figure 7" captionStartId="8.[129,194,664,688]" captionTargetBox="[321,1281,144,636]" captionTargetId="figure-739@8.[321,1281,144,636]" captionTargetPageId="8" captionText="Figure 7. Less metatarsal IV (MAP-8295) of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity.Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470530" httpUri="https://zenodo.org/record/10470530/files/figure.png" pageId="7" pageNumber="1020">Fig. 7F</figureCitation>
|
||
). The condyles are not well-defined.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED391D5138BF6C4D14FF28C256" blockId="7.[810,1460,1590,1959]" lastBlockId="8.[129,778,792,1380]" lastPageId="8" lastPageNumber="1021" pageId="7" pageNumber="1020">
|
||
<emphasis id="B94DEAFF391D5137BF6C4D14FC0DC757" box="[810,975,1590,1614]" italics="true" pageId="7" pageNumber="1020">Pedal phalanges:</emphasis>
|
||
Four phalanges belonging to the less pes have been recovered. These include the phalanx 1 of digit II (MAP-8296), ungual phalanx of digit II (MAP-8298), phalanx 3 of digit IV (MAP-8297), and ungual phalanx of digit IV (MAP-8299). The phalanx 1 of digit II (MAP-8296,
|
||
<figureCitation id="13022A68391D5137B8F74D91FAD4C7D5" box="[1201,1302,1715,1740]" captionStart="Figure 8" captionStartId="9.[113,178,1534,1558]" captionTargetBox="[210,1362,144,1506]" captionTargetId="figure-205@9.[210,1362,144,1506]" captionTargetPageId="9" captionText="Figure 8. Less pedal phalanges of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Phalanx 1 of digit II (MAP-8296) (A–E) in: A, dorsal; B, lateral; C, plantar; D, proximal; and E, distal views. Phalanx 3 of digit IV (MAP-8297) (F–J) in: F, dorsal; G, lateral; H, plantar; I, proximal; and J, distal views. Ungual phalanx of digit II (MAP-8298) (K–N) in: K, dorsal; L, plantar; M, lateral; and N, medial views. Ungual phalanx of digit IV (MAP-8299) (O–R) in: O, dorsal; P, plantar; º, lateral; and R, medial views. Abbreviations: clf, collateral ligament fovea; dl, dorsal lip; egr, extensor groove; fl, flap; lc, lateral condyle; mc, medial condyle; nvs, neurovascular sulci. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470532" httpUri="https://zenodo.org/record/10470532/files/figure.png" pageId="7" pageNumber="1020">Fig. 8A–E</figureCitation>
|
||
) is longer than tall, with an elongated diaphysis. This phalanx is keyholdershaped in both lateral and medial views. Its proximal articular facet is concave, with an almost straight plantar margin and convex dorsal margin. On the other hand, the distal end has two prominent condyles separated by a deep extensor and flexor intercondylar grooves in the median position. The medial condyle is more prominent than the lateral condyle. There are two well-developed collateral ligament foveae on both sides of the distal end.
|
||
</paragraph>
|
||
<caption id="DF46666539125138BCC749BAFE1CC3F1" ID-DOI="http://doi.org/10.5281/zenodo.10470530" ID-Zenodo-Dep="10470530" httpUri="https://zenodo.org/record/10470530/files/figure.png" pageId="8" pageNumber="1021" startId="8.[129,194,664,688]" targetBox="[321,1281,144,636]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B8636ED39125138BCC749BAFE1CC3F1" blockId="8.[129,1446,664,744]" pageId="8" pageNumber="1021">
|
||
<emphasis id="B94DEAFF39125138BCC749BAFF1AC3A9" bold="true" box="[129,216,664,688]" pageId="8" pageNumber="1021">Figure 7.</emphasis>
|
||
Less metatarsal IV (MAP-8295) of the holotype specimen of
|
||
<taxonomicName id="4C394D6E39125138BF5D49BAFC20C3A9" authority="Sánchez-Fenollosa & Verdú & Cobos, 2023" authorityName="Sánchez-Fenollosa & Verdú & Cobos" authorityYear="2023" box="[795,994,664,688]" genus="Oblitosaurus" pageId="8" pageNumber="1021" rank="species" species="bunnueli" status="gen. et sp. nov">
|
||
<emphasis id="B94DEAFF39125138BF5D49BAFC20C3A9" box="[795,994,664,688]" italics="true" pageId="8" pageNumber="1021">Oblitosaurus bunnueli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E578439125138BFA149BBFBA9C3A9" box="[999,1131,665,689]" pageId="8" pageNumber="1021" rank="species">gen. et sp. nov</taxonomicNameLabel>
|
||
.. Metatarsal IV in: A, anterior; B, medial; C, posterior; D, lateral; E, proximal; and F, distal views. Abbreviations: clf, collateral ligament fovea; f-III, facet for metatarsal III; mt, medial tuberosity. Scale bar = 10 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8636ED39125138BCDA4875FE2DC512" blockId="8.[129,778,792,1380]" pageId="8" pageNumber="1021">
|
||
The ungual phalanx of digit II (MAP-8298,
|
||
<figureCitation id="13022A6839125138BE334875FD26C276" box="[629,740,855,879]" captionStart="Figure 8" captionStartId="9.[113,178,1534,1558]" captionTargetBox="[210,1362,144,1506]" captionTargetId="figure-205@9.[210,1362,144,1506]" captionTargetPageId="9" captionText="Figure 8. Less pedal phalanges of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Phalanx 1 of digit II (MAP-8296) (A–E) in: A, dorsal; B, lateral; C, plantar; D, proximal; and E, distal views. Phalanx 3 of digit IV (MAP-8297) (F–J) in: F, dorsal; G, lateral; H, plantar; I, proximal; and J, distal views. Ungual phalanx of digit II (MAP-8298) (K–N) in: K, dorsal; L, plantar; M, lateral; and N, medial views. Ungual phalanx of digit IV (MAP-8299) (O–R) in: O, dorsal; P, plantar; º, lateral; and R, medial views. Abbreviations: clf, collateral ligament fovea; dl, dorsal lip; egr, extensor groove; fl, flap; lc, lateral condyle; mc, medial condyle; nvs, neurovascular sulci. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470532" httpUri="https://zenodo.org/record/10470532/files/figure.png" pageId="8" pageNumber="1021">Fig. 8K–N</figureCitation>
|
||
) is elongated, slightly curved, and presumably pointed (the tip is not preserved). This phalanx has a sub-circular (resembling an irregular and rounded pentagon) and concave proximal articular facet. The lateral and medial surfaces have neurovascular sulci along their entire length, with a flap.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED39125138BCDA4F31FE2EC5B1" blockId="8.[129,778,792,1380]" pageId="8" pageNumber="1021">
|
||
In contrast to the phalanx 1 of digit II, the phalanx 3 of digit IV (MAP-8297,
|
||
<figureCitation id="13022A6839125138BD294F10FE10C553" box="[367,466,1074,1098]" captionStart="Figure 8" captionStartId="9.[113,178,1534,1558]" captionTargetBox="[210,1362,144,1506]" captionTargetId="figure-205@9.[210,1362,144,1506]" captionTargetPageId="9" captionText="Figure 8. Less pedal phalanges of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Phalanx 1 of digit II (MAP-8296) (A–E) in: A, dorsal; B, lateral; C, plantar; D, proximal; and E, distal views. Phalanx 3 of digit IV (MAP-8297) (F–J) in: F, dorsal; G, lateral; H, plantar; I, proximal; and J, distal views. Ungual phalanx of digit II (MAP-8298) (K–N) in: K, dorsal; L, plantar; M, lateral; and N, medial views. Ungual phalanx of digit IV (MAP-8299) (O–R) in: O, dorsal; P, plantar; º, lateral; and R, medial views. Abbreviations: clf, collateral ligament fovea; dl, dorsal lip; egr, extensor groove; fl, flap; lc, lateral condyle; mc, medial condyle; nvs, neurovascular sulci. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470532" httpUri="https://zenodo.org/record/10470532/files/figure.png" pageId="8" pageNumber="1021">Fig. 8F–J</figureCitation>
|
||
) has no diaphysis, and it is proximodistally compressed. The proximal articular facet is concave and a dorsal lip arises from the dorsal margin. The condyles are well-developed at the distal end.
|
||
</paragraph>
|
||
<paragraph id="8B8636ED39125138BCDA4F8DFF28C47D" blockId="8.[129,778,792,1380]" pageId="8" pageNumber="1021">
|
||
Finally, the ungual phalanx of digit IV (MAP-8299,
|
||
<figureCitation id="13022A6839125138BEA54F8DFF02C5FE" captionStart="Figure 8" captionStartId="9.[113,178,1534,1558]" captionTargetBox="[210,1362,144,1506]" captionTargetId="figure-205@9.[210,1362,144,1506]" captionTargetPageId="9" captionText="Figure 8. Less pedal phalanges of the holotype specimen of Oblitosaurus bunnueli gen. et sp. nov.. Phalanx 1 of digit II (MAP-8296) (A–E) in: A, dorsal; B, lateral; C, plantar; D, proximal; and E, distal views. Phalanx 3 of digit IV (MAP-8297) (F–J) in: F, dorsal; G, lateral; H, plantar; I, proximal; and J, distal views. Ungual phalanx of digit II (MAP-8298) (K–N) in: K, dorsal; L, plantar; M, lateral; and N, medial views. Ungual phalanx of digit IV (MAP-8299) (O–R) in: O, dorsal; P, plantar; º, lateral; and R, medial views. Abbreviations: clf, collateral ligament fovea; dl, dorsal lip; egr, extensor groove; fl, flap; lc, lateral condyle; mc, medial condyle; nvs, neurovascular sulci. Scale bar = 10 cm." figureDoi="http://doi.org/10.5281/zenodo.10470532" httpUri="https://zenodo.org/record/10470532/files/figure.png" pageId="8" pageNumber="1021">Fig. 8O–R</figureCitation>
|
||
) has a concave and sub-circular proximal articular facet. Unlike the ungual phalanx of digit II (MAP-8298), it is shorter, slender, more curved and has shallower medial and lateral neurovascular sulci. Moreover, it has flaps and is pointed at the distal end.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |