treatments-xml/data/03/90/87/039087E231764124FF47724176E2FAA8.xml

<document id="0F6C47CBCE091BC77F8B29EA459521EF" ID-DOI="10.11646/zootaxa.3686.2.8" ID-GBIF-Dataset="58cfde7c-f31e-4e9e-aa6f-379c72896270" ID-ISSN="1175-5326" ID-Zenodo-Dep="221504" ID-ZooBank="23C42922-B568-4B62-AC5B-F9E8FED4AEB1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460647451992" checkinUser="plazi" docAuthor="Dehling, J. Maximilian &amp; Dehling, D. Matthias" docDate="2013" docId="039087E231764124FF47724176E2FAA8" docLanguage="en" docName="zt03686p288.pdf" docOrigin="Zootaxa 3686 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Philautus kakipanjang, sp. n." docType="treatment" docVersion="11" lastPageNumber="286" masterDocId="FFA9FF9A3177412DFFD0746B7526FFFC" masterDocTitle="A new montane species of Philautus (Amphibia: Anura: Rhacophoridae) from western Sarawak, Malaysia, Borneo" masterLastPageNumber="288" masterPageNumber="277" pageNumber="278" updateTime="1698337185733" updateUser="plazi">
<mods:mods id="9B269FC778794313025E1F955F6C3518" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3AB61176869E0978E470D82D0F8FC1EB">
<mods:title id="83DC764332422662CC946172932E827E">A new montane species of Philautus (Amphibia: Anura: Rhacophoridae) from western Sarawak, Malaysia, Borneo</mods:title>
</mods:titleInfo>
<mods:name id="A73EEE915EF512E21399B109254CA92C" type="personal">
<mods:role id="2A34F2493D30C1B8C7B48B9CE4AEAF1D">
<mods:roleTerm id="591A77FCD13FFED9296E2FC3FA9E16D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF839E09BBE2AF1C4B21531F151A039D">Dehling, J. Maximilian</mods:namePart>
</mods:name>
<mods:name id="84E2993CB5D9DAE8E1289C5C4E2A1C96" type="personal">
<mods:role id="8586CCCF6AAF290C1E4EA91149765110">
<mods:roleTerm id="6108D7D7C2823126AD2DDAA9A85BB94F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="407F146635AED72B747B99CEF40BFDAC">Dehling, D. Matthias</mods:namePart>
</mods:name>
<mods:typeOfResource id="48107D05787AB403D97B7BFBC48C6414">text</mods:typeOfResource>
<mods:relatedItem id="40BC2D47A83C6956A110BB3E3A8BE21A" type="host">
<mods:titleInfo id="088BBA96269FA0E6652BBC40D9695824">
<mods:title id="205B72837AEA6221D18AF31D6DEE6876">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7FFB2428E47B5AC1800853AEF582331A">
<mods:date id="92737A7CC4E4B1AF59F69E4368F69DE9">2013</mods:date>
<mods:detail id="DD771598AB43FD76D93645F2E50DB639" type="pubDate">
<mods:number id="6088815395D101F7610C61F9EA7727EC">2013-07-11</mods:number>
</mods:detail>
<mods:detail id="2930DFB9F1CCEB352BF117DE55E3BBE6" type="volume">
<mods:number id="F35A9E2CFD1ED568F8CE304640494E2F">3686</mods:number>
</mods:detail>
<mods:detail id="EDC4D456707994ABB76114A32EAFB23C" type="issue">
<mods:number id="E3FCFC24C3C3D8F1F0CC484CD7EF8A24">2</mods:number>
</mods:detail>
<mods:extent id="20698A6EF211FD1E8E7D8B874E4F2BC8" unit="page">
<mods:start id="525190195D7B0A6DEEF03BA189F24D28">277</mods:start>
<mods:end id="A4E8F3BC5697BB72ED062F24286A576B">288</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FDC8042FD66B94789BE241B1B4F6EE5C">journal article</mods:classification>
<mods:identifier id="0A6D1C7B2D0B7904FE94FF2AF43C705E" type="DOI">10.11646/zootaxa.3686.2.8</mods:identifier>
<mods:identifier id="5D81D219C24A80A3EFFE621D7D7BF116" type="GBIF-Dataset">58cfde7c-f31e-4e9e-aa6f-379c72896270</mods:identifier>
<mods:identifier id="5E2EAE7789272F76C4B965A362927757" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="34B3C52C71EAA107EDE53F27C75514FE" type="Zenodo-Dep">221504</mods:identifier>
<mods:identifier id="30E5905CFA07A659093670BF1000C5DA" type="ZooBank">23C42922-B568-4B62-AC5B-F9E8FED4AEB1</mods:identifier>
</mods:mods>
<treatment id="039087E231764124FF47724176E2FAA8" ID-DOI="http://doi.org/10.5281/zenodo.5685437" ID-GBIF-Taxon="119572164" ID-Zenodo-Dep="5685437" LSID="urn:lsid:plazi:treatment:039087E231764124FF47724176E2FAA8" httpUri="http://treatment.plazi.org/id/039087E231764124FF47724176E2FAA8" lastPageId="9" lastPageNumber="286" pageId="1" pageNumber="278">
<subSubSection id="C323657F3176412CFF47724174DDF99A" pageId="1" pageNumber="278" type="nomenclature">
<paragraph id="8B8636F43176412CFF47724174D2F9B8" blockId="1.[151,507,1578,1638]" box="[151,500,1578,1604]" pageId="1" pageNumber="278">
<heading id="D0CE81983176412CFF47724174D2F9B8" bold="true" box="[151,500,1578,1604]" fontSize="11" level="1" pageId="1" pageNumber="278" reason="1">
<emphasis id="B94DEAE63176412CFF47724174D2F9B8" bold="true" box="[151,500,1578,1604]" pageId="1" pageNumber="278">
<taxonomicName id="4C394D773176412CFF477241748CF9B8" ID-CoL="4G5H6" authority="Dehling &amp; Dehling, 2013" box="[151,426,1578,1604]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="278" phylum="Chordata" rank="species" species="kakipanjang" status="sp. nov.">
<emphasis id="B94DEAE63176412CFF477241748CF9B8" bold="true" box="[151,426,1578,1604]" italics="true" pageId="1" pageNumber="278">Philautus kakipanjang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E579D3176412CFE62724174D2F9B8" box="[434,500,1578,1604]" pageId="1" pageNumber="278" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8636F43176412CFF47722574DDF99A" blockId="1.[151,507,1578,1638]" box="[151,507,1613,1638]" pageId="1" pageNumber="278">Western long-legged Bush Frog</paragraph>
</subSubSection>
<subSubSection id="C323657F3176412CFF4772FE7743F879" pageId="1" pageNumber="278" type="materials_examined">
<paragraph id="8B8636F43176412CFF4772FE7769F909" blockId="1.[151,1436,1685,2034]" pageId="1" pageNumber="278">
<materialsCitation id="3B513CA93176412CFF4772FE7769F909" ID-GBIF-Occurrence="3462337301" collectingDate="2009-09-25" collectionCode="NMBE" collectorName="J. M. Dehling &amp; D. M. Dehling" country="Malaysia" elevation="780" location="Kubah National Park" pageId="1" pageNumber="278" specimenCode="NMBE 1060433" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
<typeStatus id="548288563176412CFF4772FE7422F951" box="[151,260,1685,1709]" pageId="1" pageNumber="278" type="holotype">
<emphasis id="B94DEAE63176412CFF4772FE7422F951" bold="true" box="[151,260,1685,1709]" pageId="1" pageNumber="278">Holotype</emphasis>
</typeStatus>
: 
<specimenCode id="DB9F9E8F3176412CFEC472FD74E9F951" box="[276,463,1685,1709]" pageId="1" pageNumber="278">
<collectionCode id="ED28AE313176412CFEC472FD7443F951" box="[276,357,1686,1709]" pageId="1" pageNumber="278">NMBE</collectionCode>
1060433
</specimenCode>
(ex JMD 323), 
<specimenCount id="9D3FFD7D3176412CFD5772FE7799F951" box="[647,703,1685,1709]" pageId="1" pageNumber="278" type="adult">adult</specimenCount>
<specimenCount id="9D3FFD7D3176412CFD1772FE7626F951" box="[711,768,1685,1709]" pageId="1" pageNumber="278" type="male">male</specimenCount>
, from below the summit of Gunung Serapi (
<geoCoordinate id="EE0D50333176412CFB2A72FE70BEF951" box="[1274,1432,1685,1709]" pageId="1" pageNumber="278">01°35.261’ N</geoCoordinate>
, 
<geoCoordinate id="EE0D50333176412CFF4772D27419F92D" box="[151,319,1721,1745]" pageId="1" pageNumber="278">110°11.578’ E</geoCoordinate>
; 
<quantity id="4CC19B113176412CFE9E72D274BFF92D" box="[334,409,1721,1745]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="278" unit="m" value="780.0">
<elevation id="0014D1C73176412CFE9E72D274BFF92D" box="[334,409,1721,1745]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="278" unit="m" value="780.0">780 m</elevation>
</quantity>
a.s.l.), 
<location id="8EE6602F3176412CFE2272D277CEF92D" LSID="urn:lsid:plazi:treatment:039087E231764124FF47724176E2FAA8:8EE6602F3176412CFE2272D277CEF92D" box="[498,744,1721,1745]" country="Malaysia" name="Kubah National Park" pageId="1" pageNumber="278" stateProvince="Sarawak">Kubah National Park</location>
, 
<collectingRegion id="49FDF8163176412CFD2772D2767DF92D" box="[759,859,1721,1745]" pageId="1" pageNumber="278">Sarawak</collectingRegion>
, 
<collectingCountry id="F32E76643176412CFCB972D276F4F92D" box="[873,978,1721,1745]" name="Malaysia" pageId="1" pageNumber="278">Malaysia</collectingCountry>
, collected by 
<collectorName id="26CC53223176412CFBA872D1703FF92D" box="[1144,1305,1721,1745]" pageId="1" pageNumber="278">J. M. Dehling</collectorName>
and 
<collectorName id="26CC53223176412CFA8672D175D2F909" pageId="1" pageNumber="278">D. M. Dehling</collectorName>
, 
<date id="FF8710343176412CFF2F72B674FFF909" box="[255,473,1757,1781]" pageId="1" pageNumber="278" value="2009-09-25">
<collectingDate id="EFC3E9DC3176412CFF2F72B674FFF909" box="[255,473,1757,1781]" pageId="1" pageNumber="278" value="2009-09-25">25 September 2009</collectingDate>
</date>
at 0 0 15 h.
</materialsCitation>
</paragraph>
<paragraph id="8B8636F43176412CFF1773697743F879" blockId="1.[151,1436,1685,2034]" pageId="1" pageNumber="278">
<materialsCitation id="3B513CA93176412CFF177369701CF8E5" ID-GBIF-Occurrence="3462337302" box="[199,1338,1793,1818]" collectingDate="2009-09-25" collectionCode="NMBE" collectorName="J. M. Dehling &amp; D. M. Dehling" country="Malaysia" elevation="780" location="Kubah National Park" pageId="1" pageNumber="278" specimenCode="NMBE 1060432" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="paratype">
<typeStatus id="548288563176412CFF1773697419F8E6" box="[199,319,1794,1818]" pageId="1" pageNumber="278" type="paratype">
<emphasis id="B94DEAE63176412CFF1773697419F8E6" bold="true" box="[199,319,1794,1818]" pageId="1" pageNumber="278">Paratypes</emphasis>
</typeStatus>
: 
<collectionCode id="ED28AE313176412CFE8373697482F8E5" box="[339,420,1794,1817]" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="1" pageNumber="278">NMBE</collectionCode>
1060432 (ex JMD 322), adult male, same collection details as for 
<typeStatus id="548288563176412CFB05736A701CF8E5" box="[1237,1338,1793,1817]" pageId="1" pageNumber="278" type="holotype">holotype</typeStatus>
</materialsCitation>
; 
<materialsCitation id="3B513CA93176412CFA9B73697743F879" ID-GBIF-Occurrence="3462337303" collectingDate="2009-10-07" collectionCode="NMBE" collectorName="J. M. Dehling &amp; D. M. Dehling" elevation="885" location="Gunung Gading" pageId="1" pageNumber="278" specimenCode="NMBE 1060427, 1060431" specimenCount="1" stateProvince="Gunung Gading">
<specimenCode id="DB9F9E8F3176412CFA9B736975DFF8C1" pageId="1" pageNumber="278">NMBE 1060427</specimenCode>
– 
<specimenCode id="DB9F9E8F3176412CFED8734E744FF8C1" box="[264,361,1829,1853]" pageId="1" pageNumber="278">1060431</specimenCode>
(ex JMD 461, 466, 472, 483, 488, respectively), five 
<specimenCount id="9D3FFD7D3176412CFC08734E7178F8C1" box="[984,1118,1829,1853]" pageId="1" pageNumber="278" type="adult">adult males</specimenCount>
, from the summit region of 
<collectingRegion id="49FDF8163176412CFF4773227469F89D" box="[151,335,1865,1889]" pageId="1" pageNumber="278">Gunung Gading</collectingRegion>
(
<geoCoordinate id="EE0D50333176412CFEB2732274D6F89D" box="[354,496,1865,1889]" pageId="1" pageNumber="278">1°42.813' N</geoCoordinate>
, 
<geoCoordinate id="EE0D50333176412CFE2F73227783F89D" box="[511,677,1865,1889]" pageId="1" pageNumber="278">119°50.186' E</geoCoordinate>
; 
<quantity id="4CC19B113176412CFD64732277D9F89D" box="[692,767,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="1" pageNumber="278" unit="m" value="885.0">
<elevation id="0014D1C73176412CFD64732277D9F89D" box="[692,767,1865,1889]" metricMagnitude="2" metricUnit="m" metricValue="8.85" pageId="1" pageNumber="278" unit="m" value="885.0">885 m</elevation>
</quantity>
a.s.l.), collected by 
<collectorName id="26CC53223176412CFC20732171B4F89D" box="[1008,1170,1865,1889]" pageId="1" pageNumber="278">J. M. Dehling</collectorName>
and 
<collectorName id="26CC53223176412CFB0073217059F89D" box="[1232,1407,1865,1889]" pageId="1" pageNumber="278">D. M. Dehling</collectorName>
, 
<date id="FF8710343176412CFA5E73217415F879" pageId="1" pageNumber="278" value="2009-10-07">
<collectingDate id="EFC3E9DC3176412CFA5E73217415F879" pageId="1" pageNumber="278" value="2009-10-07">7 October 2009</collectingDate>
</date>
, between 
<date id="FF8710343176412CFE727306776FF879" box="[418,585,1901,1925]" pageId="1" pageNumber="278" value="1900" valueMax="2200">1900 and 2200</date>
h.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C323657F3176412FFF1773FA7468FE76" lastPageId="2" lastPageNumber="279" pageId="1" pageNumber="278" type="diagnosis">
<paragraph id="8B8636F43176412FFF1773FA7468FE76" blockId="1.[151,1436,1685,2034]" lastBlockId="2.[151,1436,151,936]" lastPageId="2" lastPageNumber="279" pageId="1" pageNumber="278">
<emphasis id="B94DEAE63176412CFF1773FA7465F856" bold="true" box="[199,323,1937,1962]" pageId="1" pageNumber="278">Diagnosis:</emphasis>
Currently, there are no morphological synapomorphic characters considered diagnostic for the genus 
<taxonomicName id="4C394D773176412CFF0F73DE746DF831" box="[223,331,1973,1997]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="278" phylum="Chordata" rank="genus">
<emphasis id="B94DEAE63176412CFF0F73DE746DF831" box="[223,331,1973,1997]" italics="true" pageId="1" pageNumber="278">Philautus</emphasis>
</taxonomicName>
. We assign the new species to this genus for showing morphological and bioacoustic resemblance and close genetic relationship to several of the Bornean species assigned to the genus. The new species differs from its congeners by the combination of the following characters: size moderately large (SVL of adult males 
<quantity id="4CC19B113175412FFAFC74FC75EEFF28" metricMagnitude="-2" metricUnit="m" metricValue="2.245" metricValueMax="2.37" metricValueMin="2.12" pageId="2" pageNumber="279" unit="mm" value="22.45" valueMax="23.7" valueMin="21.2">21.2–23.7 mm</quantity>
); hindlimbs long, tibiotarsal joint reaching beyond tip of snout when legs are adpressed to body, TFL/SVL 0.56–0.64; snout acuminate in dorsal view, longer than eye diameter; median lingual process absent; heel without dermal appendage; nuptial pads small, prominent; toes extensively webbed, formula 
<date id="FF8710343175412FFBA5756C71A8FEE3" box="[1141,1166,263,287]" pageId="2" pageNumber="279">
<emphasis id="B94DEAE63175412FFBA5756C71A6FEE3" bold="true" box="[1141,1152,263,287]" pageId="2" pageNumber="279">I</emphasis>
2
</date>
- -2.25 
<emphasis id="B94DEAE63175412FFB1E756C71C2FEE3" bold="true" box="[1230,1252,263,287]" pageId="2" pageNumber="279">II</emphasis>
1.5-2.75 
<emphasis id="B94DEAE63175412FFA91756C7044FEE3" bold="true" box="[1345,1378,263,287]" pageId="2" pageNumber="279">III</emphasis>
1.75- 
<date id="FF8710343175412FFF4775417583FEBE" box="[151,165,298,322]" pageId="2" pageNumber="279">3</date>
<emphasis id="B94DEAE63175412FFF75754075E2FEBF" bold="true" box="[165,196,299,323]" pageId="2" pageNumber="279">IV</emphasis>
2.75-1.5 
<emphasis id="B94DEAE63175412FFEF175407415FEBF" bold="true" box="[289,307,299,323]" pageId="2" pageNumber="279">V</emphasis>
; iris golden in life; advertisement call consisting of a single note with 9–12 pulses, lasting 342–478 ms, with a dominant frequency of 2.0–2.7 kHz, moderate frequency modulation and marked amplitude modulation at 18.1–18.5 °C.
</paragraph>
</subSubSection>
<subSubSection id="C323657F3175412BFF1775FD7193FD13" lastPageId="6" lastPageNumber="283" pageId="2" pageNumber="279" type="description">
<paragraph id="8B8636F43175412FFF1775FD7635FC54" blockId="2.[151,1436,151,936]" pageId="2" pageNumber="279">
<emphasis id="B94DEAE63175412FFF1775FD74F9FE53" bold="true" box="[199,479,406,431]" pageId="2" pageNumber="279">
Description of 
<typeStatus id="548288563175412FFEA875FC74F9FE53" box="[376,479,407,431]" pageId="2" pageNumber="279" type="holotype">holotype</typeStatus>
</emphasis>
: Measurements are provided in 
<tableCitation id="C6BB034F3175412FFC9C75FC7686FE53" box="[844,928,407,431]" captionStart="TABLE 1" captionStartId="2.[151,239,977,1001]" captionTargetBox="[159,1401,1028,1959]" captionTargetPageId="2" captionText="TABLE 1. Measurements of the type series of Philautus kakipanjang sp. n. For abbreviations see Material &amp; Methods." httpUri="http://table.plazi.org/id/DF46667C3175412FFF4777BA70B7FC14" pageId="2" pageNumber="279" tableUuid="DF46667C3175412FFF4777BA70B7FC14">Table 1</tableCitation>
. Body moderately sturdy, widest at temporal region, tapering to groin (
<figureCitation id="13022A713175412FFE6E75D17739FE2E" box="[446,543,442,466]" captionStart="FIGURE 1" captionStartId="3.[151,250,1566,1589]" captionTargetBox="[151,1436,195,1541]" captionTargetId="figure@3.[151,1436,195,1543]" captionTargetPageId="3" captionText="FIGURE 1. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of Philautus kakipanjang (NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of Philautus kakipanjang (NMBE 1060432) in life, showing the colouration during the night." httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="2" pageNumber="279">Figure 1</figureCitation>
); head large (HL/SVL 0.36, HW/SVL 0.41), wider than long (HW/HL 1.15); snout long (SL/HL 0.50), acuminate in dorsal view, rounded in profile (
<figureCitation id="13022A713175412FFC0275B47115FE0B" box="[978,1075,479,503]" captionStart="FIGURE 1" captionStartId="3.[151,250,1566,1589]" captionTargetBox="[151,1436,195,1541]" captionTargetId="figure@3.[151,1436,195,1543]" captionTargetPageId="3" captionText="FIGURE 1. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of Philautus kakipanjang (NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of Philautus kakipanjang (NMBE 1060432) in life, showing the colouration during the night." httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="2" pageNumber="279">Figure 1</figureCitation>
), projecting beyond lower jaw, wider than long (SL/EE 0.86); canthus rostralis distinct, straight-lined in profile, strongly concave in dorsal view; loreal region oblique, concave; nostrils rounded, directed laterally; situated much closer to tip of snout than to eye (EN/NS 1.45), separated from each other by distance subequal to distance between eye and nostril (NN/EN 0.96); eyes directed anterolaterally, protruding, large (ED/HL 0.45); pupil horizontal; eye diameter slightly shorter than snout (ED/SL 0.91); interorbital distance wider than upper eyelid (IO/EW 1.16) and greater than internarial distance (IO/NN 1.28); tympanum distinctly visible, its posterodorsal fourth concealed under thick layer of skin, separated from eye by one-fifth of its diameter; tympanum diameter two-fifths of eye diameter; upper jaw with dentition; choanae small, oval, located far anterolaterally at margins of roof of the mouth, concealed by upper jaw in ventral view; vomer processes and teeth absent; tongue long (
<quantity id="4CC19B113175412FFC58774976C2FCC6" box="[904,996,802,826]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="2" pageNumber="279" unit="mm" value="7.9">7.9 mm</quantity>
) and broad (
<quantity id="4CC19B113175412FFBAB774971F0FCC6" box="[1147,1238,802,826]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="279" unit="mm" value="4.6">4.6 mm</quantity>
at widest point), bilobed for one-fifth of its length, free distally for two-thirds of its length; median lingual process absent; vocal sac single, median, subgular; vocal sac aperture on each side of the mouth, slit-like, long, directed laterally, situated slightly closer to corner of mouth than to base of tongue.
</paragraph>
<caption id="DF46667C3175412FFF4777BA70B7FC14" ID-Table-UUID="DF46667C3175412FFF4777BA70B7FC14" box="[151,1425,977,1001]" httpUri="http://table.plazi.org/id/DF46667C3175412FFF4777BA70B7FC14" pageId="2" pageNumber="279" targetBox="[159,1401,1028,1959]" targetIsTable="true" targetPageId="2">
<paragraph id="8B8636F43175412FFF4777BA70B7FC14" blockId="2.[151,1425,977,1001]" box="[151,1425,977,1001]" pageId="2" pageNumber="279">
<emphasis id="B94DEAE63175412FFF4777BA7424FC15" bold="true" box="[151,258,977,1001]" pageId="2" pageNumber="279">TABLE 1</emphasis>
. Measurements of the type series of 
<taxonomicName id="4C394D773175412FFD5C77BA765AFC14" box="[652,892,977,1000]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="279" phylum="Chordata" rank="species" species="kakipanjang" status="sp. nov.">
<emphasis id="B94DEAE63175412FFD5C77BA765AFC14" box="[652,892,977,1000]" italics="true" pageId="2" pageNumber="279">Philautus kakipanjang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E579D3175412FFC5377BA7698FC15" box="[899,958,977,1001]" pageId="2" pageNumber="279" rank="species">
<emphasis id="B94DEAE63175412FFC5377BA7698FC15" bold="true" box="[899,958,977,1001]" pageId="2" pageNumber="279">sp. n.</emphasis>
</taxonomicNameLabel>
For abbreviations see Material &amp; Methods.
</paragraph>
</caption>
<paragraph id="8B8636F43175412FFF4F706E700EF85B" pageId="2" pageNumber="279">
<table id="F939C4543175BED2FF4F706F705FF85B" box="[159,1401,1028,1959]" gridcols="8" gridrows="23" pageId="2" pageNumber="279">
<tr id="350934B63175BED2FF4F706F705FFBE6" box="[159,1401,1028,1050]" gridrow="0" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F706F75CEFBE6" box="[159,232,1028,1050]" gridcol="0" gridrow="0" pageId="2" pageNumber="279">NMBE</th>
<th id="76D85DCA3175BED2FEEF706F748FFBE6" box="[319,425,1028,1050]" gridcol="1" gridrow="0" pageId="2" pageNumber="279">1060433</th>
<th id="76D85DCA3175BED2FE33706F776BFBE6" box="[483,589,1028,1050]" gridcol="2" gridrow="0" pageId="2" pageNumber="279">1060432</th>
<th id="76D85DCA3175BED2FD58706F77DEFBE6" box="[648,760,1028,1050]" gridcol="3" gridrow="0" pageId="2" pageNumber="279">1060427</th>
<th id="76D85DCA3175BED2FCF8706F76BFFBE6" box="[808,921,1028,1050]" gridcol="4" gridrow="0" pageId="2" pageNumber="279">1060428</th>
<th id="76D85DCA3175BED2FC18706F711FFBE6" box="[968,1081,1028,1050]" gridcol="5" gridrow="0" pageId="2" pageNumber="279">1060429</th>
<th id="76D85DCA3175BED2FBB8706F71FFFBE6" box="[1128,1241,1028,1050]" gridcol="6" gridrow="0" pageId="2" pageNumber="279">1060430</th>
<th id="76D85DCA3175BED2FAD8706F705FFBE6" box="[1288,1401,1028,1050]" gridcol="7" gridrow="0" pageId="2" pageNumber="279">1060431</th>
</tr>
<tr id="350934B63175BED2FF4F7046705FFBBF" box="[159,1401,1069,1091]" gridrow="1" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F704675CEFBBF" box="[159,232,1069,1091]" gridcol="0" gridrow="1" pageId="2" pageNumber="279">status</th>
<td id="76D85DCA3175BED2FEEF7046748FFBBF" box="[319,425,1069,1091]" gridcol="1" gridrow="1" pageId="2" pageNumber="279">holotype</td>
<td id="76D85DCA3175BED2FE337046776BFBBF" box="[483,589,1069,1091]" gridcol="2" gridrow="1" pageId="2" pageNumber="279">paratype</td>
<td id="76D85DCA3175BED2FD58704677DEFBBF" box="[648,760,1069,1091]" gridcol="3" gridrow="1" pageId="2" pageNumber="279">paratype</td>
<td id="76D85DCA3175BED2FCF8704676BFFBBF" box="[808,921,1069,1091]" gridcol="4" gridrow="1" pageId="2" pageNumber="279">paratype</td>
<td id="76D85DCA3175BED2FC187046711FFBBF" box="[968,1081,1069,1091]" gridcol="5" gridrow="1" pageId="2" pageNumber="279">paratype</td>
<td id="76D85DCA3175BED2FBB8704671FFFBBF" box="[1128,1241,1069,1091]" gridcol="6" gridrow="1" pageId="2" pageNumber="279">paratype</td>
<td id="76D85DCA3175BED2FAD87046705FFBBF" box="[1288,1401,1069,1091]" gridcol="7" gridrow="1" pageId="2" pageNumber="279">paratype</td>
</tr>
<tr id="350934B63175BED2FF4F703D705FFB90" box="[159,1401,1110,1132]" gridrow="2" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F703D75CEFB90" box="[159,232,1110,1132]" gridcol="0" gridrow="2" pageId="2" pageNumber="279">origin</th>
<td id="76D85DCA3175BED2FEEF703D748FFB90" box="[319,425,1110,1132]" gridcol="1" gridrow="2" pageId="2" pageNumber="279">Kubah NP</td>
<td id="76D85DCA3175BED2FE33703D776BFB90" box="[483,589,1110,1132]" gridcol="2" gridrow="2" pageId="2" pageNumber="279">Kubah NP</td>
<td id="76D85DCA3175BED2FD58703D77DEFB90" box="[648,760,1110,1132]" gridcol="3" gridrow="2" pageId="2" pageNumber="279">Gading NP</td>
<td id="76D85DCA3175BED2FCF8703D76BFFB90" box="[808,921,1110,1132]" gridcol="4" gridrow="2" pageId="2" pageNumber="279">Gading NP</td>
<td id="76D85DCA3175BED2FC18703D711FFB90" box="[968,1081,1110,1132]" gridcol="5" gridrow="2" pageId="2" pageNumber="279">Gading NP</td>
<td id="76D85DCA3175BED2FBB8703D71FFFB90" box="[1128,1241,1110,1132]" gridcol="6" gridrow="2" pageId="2" pageNumber="279">Gading NP</td>
<td id="76D85DCA3175BED2FAD8703D705FFB90" box="[1288,1401,1110,1132]" gridcol="7" gridrow="2" pageId="2" pageNumber="279">Gading NP</td>
</tr>
<tr id="350934B63175BED2FF4F70EB705FFB69" box="[159,1401,1152,1173]" gridrow="3" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F70EB75CEFB69" box="[159,232,1152,1173]" gridcol="0" gridrow="3" pageId="2" pageNumber="279">SVL</th>
<td id="76D85DCA3175BED2FEEF70EB748FFB69" box="[319,425,1152,1173]" gridcol="1" gridrow="3" pageId="2" pageNumber="279">21.4</td>
<td id="76D85DCA3175BED2FE3370EB776BFB69" box="[483,589,1152,1173]" gridcol="2" gridrow="3" pageId="2" pageNumber="279">22.0</td>
<td id="76D85DCA3175BED2FD5870EB77DEFB69" box="[648,760,1152,1173]" gridcol="3" gridrow="3" pageId="2" pageNumber="279">21.4</td>
<td id="76D85DCA3175BED2FCF870EB76BFFB69" box="[808,921,1152,1173]" gridcol="4" gridrow="3" pageId="2" pageNumber="279">22.3</td>
<td id="76D85DCA3175BED2FC1870EB711FFB69" box="[968,1081,1152,1173]" gridcol="5" gridrow="3" pageId="2" pageNumber="279">23.7</td>
<td id="76D85DCA3175BED2FBB870EB71FFFB69" box="[1128,1241,1152,1173]" gridcol="6" gridrow="3" pageId="2" pageNumber="279">21.4</td>
<td id="76D85DCA3175BED2FAD870EB705FFB69" box="[1288,1401,1152,1173]" gridcol="7" gridrow="3" pageId="2" pageNumber="279">21.2</td>
</tr>
<tr id="350934B63175BED2FF4F70C2705FFB43" box="[159,1401,1193,1215]" gridrow="4" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F70C275CEFB43" box="[159,232,1193,1215]" gridcol="0" gridrow="4" pageId="2" pageNumber="279">TFL</th>
<td id="76D85DCA3175BED2FEEF70C2748FFB43" box="[319,425,1193,1215]" gridcol="1" gridrow="4" pageId="2" pageNumber="279">12.1</td>
<td id="76D85DCA3175BED2FE3370C2776BFB43" box="[483,589,1193,1215]" gridcol="2" gridrow="4" pageId="2" pageNumber="279">12.7</td>
<td id="76D85DCA3175BED2FD5870C277DEFB43" box="[648,760,1193,1215]" gridcol="3" gridrow="4" pageId="2" pageNumber="279">12.7</td>
<td id="76D85DCA3175BED2FCF870C276BFFB43" box="[808,921,1193,1215]" gridcol="4" gridrow="4" pageId="2" pageNumber="279">13.7</td>
<td id="76D85DCA3175BED2FC1870C2711FFB43" box="[968,1081,1193,1215]" gridcol="5" gridrow="4" pageId="2" pageNumber="279">13.6</td>
<td id="76D85DCA3175BED2FBB870C271FFFB43" box="[1128,1241,1193,1215]" gridcol="6" gridrow="4" pageId="2" pageNumber="279">12.6</td>
<td id="76D85DCA3175BED2FAD870C2705FFB43" box="[1288,1401,1193,1215]" gridcol="7" gridrow="4" pageId="2" pageNumber="279">13.6</td>
</tr>
<tr id="350934B63175BED2FF4F70B9705FFB14" box="[159,1401,1234,1256]" gridrow="5" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F70B975CEFB14" box="[159,232,1234,1256]" gridcol="0" gridrow="5" pageId="2" pageNumber="279">FOT</th>
<td id="76D85DCA3175BED2FEEF70B9748FFB14" box="[319,425,1234,1256]" gridcol="1" gridrow="5" pageId="2" pageNumber="279">9.5</td>
<td id="76D85DCA3175BED2FE3370B9776BFB14" box="[483,589,1234,1256]" gridcol="2" gridrow="5" pageId="2" pageNumber="279">9.6</td>
<td id="76D85DCA3175BED2FD5870B977DEFB14" box="[648,760,1234,1256]" gridcol="3" gridrow="5" pageId="2" pageNumber="279">9.4</td>
<td id="76D85DCA3175BED2FCF870B976BFFB14" box="[808,921,1234,1256]" gridcol="4" gridrow="5" pageId="2" pageNumber="279">10.8</td>
<td id="76D85DCA3175BED2FC1870B9711FFB14" box="[968,1081,1234,1256]" gridcol="5" gridrow="5" pageId="2" pageNumber="279">10.5</td>
<td id="76D85DCA3175BED2FBB870B971FFFB14" box="[1128,1241,1234,1256]" gridcol="6" gridrow="5" pageId="2" pageNumber="279">9.9</td>
<td id="76D85DCA3175BED2FAD870B9705FFB14" box="[1288,1401,1234,1256]" gridcol="7" gridrow="5" pageId="2" pageNumber="279">10.3</td>
</tr>
<tr id="350934B63175BED2FF4F7097705FFAEE" box="[159,1401,1276,1298]" gridrow="6" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F709775CEFAEE" box="[159,232,1276,1298]" gridcol="0" gridrow="6" pageId="2" pageNumber="279">TarL</th>
<td id="76D85DCA3175BED2FEEF7097748FFAEE" box="[319,425,1276,1298]" gridcol="1" gridrow="6" pageId="2" pageNumber="279">15.1</td>
<td id="76D85DCA3175BED2FE337097776BFAEE" box="[483,589,1276,1298]" gridcol="2" gridrow="6" pageId="2" pageNumber="279">15.8</td>
<td id="76D85DCA3175BED2FD58709777DEFAEE" box="[648,760,1276,1298]" gridcol="3" gridrow="6" pageId="2" pageNumber="279">15.8</td>
<td id="76D85DCA3175BED2FCF8709776BFFAEE" box="[808,921,1276,1298]" gridcol="4" gridrow="6" pageId="2" pageNumber="279">16.8</td>
<td id="76D85DCA3175BED2FC187097711FFAEE" box="[968,1081,1276,1298]" gridcol="5" gridrow="6" pageId="2" pageNumber="279">17.2</td>
<td id="76D85DCA3175BED2FBB8709771FFFAEE" box="[1128,1241,1276,1298]" gridcol="6" gridrow="6" pageId="2" pageNumber="279">16.2</td>
<td id="76D85DCA3175BED2FAD87097705FFAEE" box="[1288,1401,1276,1298]" gridcol="7" gridrow="6" pageId="2" pageNumber="279">16.7</td>
</tr>
<tr id="350934B63175BED2FF4F714E705FFAC7" box="[159,1401,1317,1339]" gridrow="7" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F714E75CEFAC7" box="[159,232,1317,1339]" gridcol="0" gridrow="7" pageId="2" pageNumber="279">LEG</th>
<td id="76D85DCA3175BED2FEEF714E748FFAC7" box="[319,425,1317,1339]" gridcol="1" gridrow="7" pageId="2" pageNumber="279">36.8</td>
<td id="76D85DCA3175BED2FE33714E776BFAC7" box="[483,589,1317,1339]" gridcol="2" gridrow="7" pageId="2" pageNumber="279">37.4</td>
<td id="76D85DCA3175BED2FD58714E77DEFAC7" box="[648,760,1317,1339]" gridcol="3" gridrow="7" pageId="2" pageNumber="279">37.3</td>
<td id="76D85DCA3175BED2FCF8714E76BFFAC7" box="[808,921,1317,1339]" gridcol="4" gridrow="7" pageId="2" pageNumber="279">41.0</td>
<td id="76D85DCA3175BED2FC18714E711FFAC7" box="[968,1081,1317,1339]" gridcol="5" gridrow="7" pageId="2" pageNumber="279">41.5</td>
<td id="76D85DCA3175BED2FBB8714E71FFFAC7" box="[1128,1241,1317,1339]" gridcol="6" gridrow="7" pageId="2" pageNumber="279">38.4</td>
<td id="76D85DCA3175BED2FAD8714E705FFAC7" box="[1288,1401,1317,1339]" gridcol="7" gridrow="7" pageId="2" pageNumber="279">40.6</td>
</tr>
<tr id="350934B63175BED2FF4F7125705FFA98" box="[159,1401,1358,1380]" gridrow="8" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F712575CEFA98" box="[159,232,1358,1380]" gridcol="0" gridrow="8" pageId="2" pageNumber="279">THL</th>
<td id="76D85DCA3175BED2FEEF7125748FFA98" box="[319,425,1358,1380]" gridcol="1" gridrow="8" pageId="2" pageNumber="279">11.4</td>
<td id="76D85DCA3175BED2FE337125776BFA98" box="[483,589,1358,1380]" gridcol="2" gridrow="8" pageId="2" pageNumber="279">11.7</td>
<td id="76D85DCA3175BED2FD58712577DEFA98" box="[648,760,1358,1380]" gridcol="3" gridrow="8" pageId="2" pageNumber="279">11.4</td>
<td id="76D85DCA3175BED2FCF8712576BFFA98" box="[808,921,1358,1380]" gridcol="4" gridrow="8" pageId="2" pageNumber="279">12.7</td>
<td id="76D85DCA3175BED2FC187125711FFA98" box="[968,1081,1358,1380]" gridcol="5" gridrow="8" pageId="2" pageNumber="279">12.6</td>
<td id="76D85DCA3175BED2FBB8712571FFFA98" box="[1128,1241,1358,1380]" gridcol="6" gridrow="8" pageId="2" pageNumber="279">12.2</td>
<td id="76D85DCA3175BED2FAD87125705FFA98" box="[1288,1401,1358,1380]" gridcol="7" gridrow="8" pageId="2" pageNumber="279">12.4</td>
</tr>
<tr id="350934B63175BED2FF4F7113705FFA72" box="[159,1401,1400,1422]" gridrow="9" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F711375CEFA72" box="[159,232,1400,1422]" gridcol="0" gridrow="9" pageId="2" pageNumber="279">ARM</th>
<td id="76D85DCA3175BED2FEEF7113748FFA72" box="[319,425,1400,1422]" gridcol="1" gridrow="9" pageId="2" pageNumber="279">10.7</td>
<td id="76D85DCA3175BED2FE337113776BFA72" box="[483,589,1400,1422]" gridcol="2" gridrow="9" pageId="2" pageNumber="279">11.0</td>
<td id="76D85DCA3175BED2FD58711377DEFA72" box="[648,760,1400,1422]" gridcol="3" gridrow="9" pageId="2" pageNumber="279">10.8</td>
<td id="76D85DCA3175BED2FCF8711376BFFA72" box="[808,921,1400,1422]" gridcol="4" gridrow="9" pageId="2" pageNumber="279">11.4</td>
<td id="76D85DCA3175BED2FC187113711FFA72" box="[968,1081,1400,1422]" gridcol="5" gridrow="9" pageId="2" pageNumber="279">12.5</td>
<td id="76D85DCA3175BED2FBB8711371FFFA72" box="[1128,1241,1400,1422]" gridcol="6" gridrow="9" pageId="2" pageNumber="279">10.9</td>
<td id="76D85DCA3175BED2FAD87113705FFA72" box="[1288,1401,1400,1422]" gridcol="7" gridrow="9" pageId="2" pageNumber="279">11.1</td>
</tr>
<tr id="350934B63175BED2FF4F71CA705FFA4B" box="[159,1401,1441,1463]" gridrow="10" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F71CA75CEFA4B" box="[159,232,1441,1463]" gridcol="0" gridrow="10" pageId="2" pageNumber="279">HND</th>
<td id="76D85DCA3175BED2FEEF71CA748FFA4B" box="[319,425,1441,1463]" gridcol="1" gridrow="10" pageId="2" pageNumber="279">6.2</td>
<td id="76D85DCA3175BED2FE3371CA776BFA4B" box="[483,589,1441,1463]" gridcol="2" gridrow="10" pageId="2" pageNumber="279">6.3</td>
<td id="76D85DCA3175BED2FD5871CA77DEFA4B" box="[648,760,1441,1463]" gridcol="3" gridrow="10" pageId="2" pageNumber="279">6.5</td>
<td id="76D85DCA3175BED2FCF871CA76BFFA4B" box="[808,921,1441,1463]" gridcol="4" gridrow="10" pageId="2" pageNumber="279">7.0</td>
<td id="76D85DCA3175BED2FC1871CA711FFA4B" box="[968,1081,1441,1463]" gridcol="5" gridrow="10" pageId="2" pageNumber="279">7.6</td>
<td id="76D85DCA3175BED2FBB871CA71FFFA4B" box="[1128,1241,1441,1463]" gridcol="6" gridrow="10" pageId="2" pageNumber="279">6.9</td>
<td id="76D85DCA3175BED2FAD871CA705FFA4B" box="[1288,1401,1441,1463]" gridcol="7" gridrow="10" pageId="2" pageNumber="279">6.7</td>
</tr>
<tr id="350934B63175BED2FF4F71A1705FFA1D" box="[159,1401,1482,1505]" gridrow="11" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F71A175CEFA1D" box="[159,232,1482,1505]" gridcol="0" gridrow="11" pageId="2" pageNumber="279">HW</th>
<td id="76D85DCA3175BED2FEEF71A1748FFA1D" box="[319,425,1482,1505]" gridcol="1" gridrow="11" pageId="2" pageNumber="279">8.8</td>
<td id="76D85DCA3175BED2FE3371A1776BFA1D" box="[483,589,1482,1505]" gridcol="2" gridrow="11" pageId="2" pageNumber="279">9.3</td>
<td id="76D85DCA3175BED2FD5871A177DEFA1D" box="[648,760,1482,1505]" gridcol="3" gridrow="11" pageId="2" pageNumber="279">8.3</td>
<td id="76D85DCA3175BED2FCF871A176BFFA1D" box="[808,921,1482,1505]" gridcol="4" gridrow="11" pageId="2" pageNumber="279">8.9</td>
<td id="76D85DCA3175BED2FC1871A1711FFA1D" box="[968,1081,1482,1505]" gridcol="5" gridrow="11" pageId="2" pageNumber="279">9.7</td>
<td id="76D85DCA3175BED2FBB871A171FFFA1D" box="[1128,1241,1482,1505]" gridcol="6" gridrow="11" pageId="2" pageNumber="279">8.5</td>
<td id="76D85DCA3175BED2FAD871A1705FFA1D" box="[1288,1401,1482,1505]" gridcol="7" gridrow="11" pageId="2" pageNumber="279">8.7</td>
</tr>
<tr id="350934B63175BED2FF4F719F705FF9F6" box="[159,1401,1524,1546]" gridrow="12" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F719F75CEF9F6" box="[159,232,1524,1546]" gridcol="0" gridrow="12" pageId="2" pageNumber="279">HL</th>
<td id="76D85DCA3175BED2FEEF719F748FF9F6" box="[319,425,1524,1546]" gridcol="1" gridrow="12" pageId="2" pageNumber="279">7.6</td>
<td id="76D85DCA3175BED2FE33719F776BF9F6" box="[483,589,1524,1546]" gridcol="2" gridrow="12" pageId="2" pageNumber="279">8.0</td>
<td id="76D85DCA3175BED2FD58719F77DEF9F6" box="[648,760,1524,1546]" gridcol="3" gridrow="12" pageId="2" pageNumber="279">6.9</td>
<td id="76D85DCA3175BED2FCF8719F76BFF9F6" box="[808,921,1524,1546]" gridcol="4" gridrow="12" pageId="2" pageNumber="279">7.4</td>
<td id="76D85DCA3175BED2FC18719F711FF9F6" box="[968,1081,1524,1546]" gridcol="5" gridrow="12" pageId="2" pageNumber="279">8.5</td>
<td id="76D85DCA3175BED2FBB8719F71FFF9F6" box="[1128,1241,1524,1546]" gridcol="6" gridrow="12" pageId="2" pageNumber="279">7.4</td>
<td id="76D85DCA3175BED2FAD8719F705FF9F6" box="[1288,1401,1524,1546]" gridcol="7" gridrow="12" pageId="2" pageNumber="279">7.2</td>
</tr>
<tr id="350934B63175BED2FF4F7276705FF9CF" box="[159,1401,1565,1587]" gridrow="13" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F727675CEF9CF" box="[159,232,1565,1587]" gridcol="0" gridrow="13" pageId="2" pageNumber="279">IO</th>
<td id="76D85DCA3175BED2FEEF7276748FF9CF" box="[319,425,1565,1587]" gridcol="1" gridrow="13" pageId="2" pageNumber="279">2.8</td>
<td id="76D85DCA3175BED2FE337276776BF9CF" box="[483,589,1565,1587]" gridcol="2" gridrow="13" pageId="2" pageNumber="279">2.7</td>
<td id="76D85DCA3175BED2FD58727677DEF9CF" box="[648,760,1565,1587]" gridcol="3" gridrow="13" pageId="2" pageNumber="279">2.5</td>
<td id="76D85DCA3175BED2FCF8727676BFF9CF" box="[808,921,1565,1587]" gridcol="4" gridrow="13" pageId="2" pageNumber="279">2.4</td>
<td id="76D85DCA3175BED2FC187276711FF9CF" box="[968,1081,1565,1587]" gridcol="5" gridrow="13" pageId="2" pageNumber="279">2.8</td>
<td id="76D85DCA3175BED2FBB8727671FFF9CF" box="[1128,1241,1565,1587]" gridcol="6" gridrow="13" pageId="2" pageNumber="279">2.4</td>
<td id="76D85DCA3175BED2FAD87276705FF9CF" box="[1288,1401,1565,1587]" gridcol="7" gridrow="13" pageId="2" pageNumber="279">2.7</td>
</tr>
<tr id="350934B63175BED2FF4F722D705FF9A0" box="[159,1401,1606,1628]" gridrow="14" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F722D75CEF9A0" box="[159,232,1606,1628]" gridcol="0" gridrow="14" pageId="2" pageNumber="279">EW</th>
<td id="76D85DCA3175BED2FEEF722D748FF9A0" box="[319,425,1606,1628]" gridcol="1" gridrow="14" pageId="2" pageNumber="279">2.4</td>
<td id="76D85DCA3175BED2FE33722D776BF9A0" box="[483,589,1606,1628]" gridcol="2" gridrow="14" pageId="2" pageNumber="279">2.6</td>
<td id="76D85DCA3175BED2FD58722D77DEF9A0" box="[648,760,1606,1628]" gridcol="3" gridrow="14" pageId="2" pageNumber="279">2.1</td>
<td id="76D85DCA3175BED2FCF8722D76BFF9A0" box="[808,921,1606,1628]" gridcol="4" gridrow="14" pageId="2" pageNumber="279">2.2</td>
<td id="76D85DCA3175BED2FC18722D711FF9A0" box="[968,1081,1606,1628]" gridcol="5" gridrow="14" pageId="2" pageNumber="279">2.4</td>
<td id="76D85DCA3175BED2FBB8722D71FFF9A0" box="[1128,1241,1606,1628]" gridcol="6" gridrow="14" pageId="2" pageNumber="279">2.1</td>
<td id="76D85DCA3175BED2FAD8722D705FF9A0" box="[1288,1401,1606,1628]" gridcol="7" gridrow="14" pageId="2" pageNumber="279">2.2</td>
</tr>
<tr id="350934B63175BED2FF4F721B705FF97A" box="[159,1401,1648,1670]" gridrow="15" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F721B75CEF97A" box="[159,232,1648,1670]" gridcol="0" gridrow="15" pageId="2" pageNumber="279">ED</th>
<td id="76D85DCA3175BED2FEEF721B748FF97A" box="[319,425,1648,1670]" gridcol="1" gridrow="15" pageId="2" pageNumber="279">3.4</td>
<td id="76D85DCA3175BED2FE33721B776BF97A" box="[483,589,1648,1670]" gridcol="2" gridrow="15" pageId="2" pageNumber="279">3.6</td>
<td id="76D85DCA3175BED2FD58721B77DEF97A" box="[648,760,1648,1670]" gridcol="3" gridrow="15" pageId="2" pageNumber="279">2.9</td>
<td id="76D85DCA3175BED2FCF8721B76BFF97A" box="[808,921,1648,1670]" gridcol="4" gridrow="15" pageId="2" pageNumber="279">3.2</td>
<td id="76D85DCA3175BED2FC18721B711FF97A" box="[968,1081,1648,1670]" gridcol="5" gridrow="15" pageId="2" pageNumber="279">3.7</td>
<td id="76D85DCA3175BED2FBB8721B71FFF97A" box="[1128,1241,1648,1670]" gridcol="6" gridrow="15" pageId="2" pageNumber="279">3.1</td>
<td id="76D85DCA3175BED2FAD8721B705FF97A" box="[1288,1401,1648,1670]" gridcol="7" gridrow="15" pageId="2" pageNumber="279">3.2</td>
</tr>
<tr id="350934B63175BED2FF4F72F1705FF953" box="[159,1401,1690,1711]" gridrow="16" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F72F175CEF953" box="[159,232,1690,1711]" gridcol="0" gridrow="16" pageId="2" pageNumber="279">TD</th>
<td id="76D85DCA3175BED2FEEF72F1748FF953" box="[319,425,1690,1711]" gridcol="1" gridrow="16" pageId="2" pageNumber="279">1.3</td>
<td id="76D85DCA3175BED2FE3372F1776BF953" box="[483,589,1690,1711]" gridcol="2" gridrow="16" pageId="2" pageNumber="279">1.4</td>
<td id="76D85DCA3175BED2FD5872F177DEF953" box="[648,760,1690,1711]" gridcol="3" gridrow="16" pageId="2" pageNumber="279">1.4</td>
<td id="76D85DCA3175BED2FCF872F176BFF953" box="[808,921,1690,1711]" gridcol="4" gridrow="16" pageId="2" pageNumber="279">1.3</td>
<td id="76D85DCA3175BED2FC1872F1711FF953" box="[968,1081,1690,1711]" gridcol="5" gridrow="16" pageId="2" pageNumber="279">1.4</td>
<td id="76D85DCA3175BED2FBB872F171FFF953" box="[1128,1241,1690,1711]" gridcol="6" gridrow="16" pageId="2" pageNumber="279">1.4</td>
<td id="76D85DCA3175BED2FAD872F1705FF953" box="[1288,1401,1690,1711]" gridcol="7" gridrow="16" pageId="2" pageNumber="279">1.4</td>
</tr>
<tr id="350934B63175BED2FF4F72A8705FF925" box="[159,1401,1731,1753]" gridrow="17" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F72A875CEF925" box="[159,232,1731,1753]" gridcol="0" gridrow="17" pageId="2" pageNumber="279">ET</th>
<td id="76D85DCA3175BED2FEEF72A8748FF925" box="[319,425,1731,1753]" gridcol="1" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FE3372A8776BF925" box="[483,589,1731,1753]" gridcol="2" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FD5872A877DEF925" box="[648,760,1731,1753]" gridcol="3" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FCF872A876BFF925" box="[808,921,1731,1753]" gridcol="4" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FC1872A8711FF925" box="[968,1081,1731,1753]" gridcol="5" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FBB872A871FFF925" box="[1128,1241,1731,1753]" gridcol="6" gridrow="17" pageId="2" pageNumber="279">0.2</td>
<td id="76D85DCA3175BED2FAD872A8705FF925" box="[1288,1401,1731,1753]" gridcol="7" gridrow="17" pageId="2" pageNumber="279">0.2</td>
</tr>
<tr id="350934B63175BED2FF4F7287705FF8FE" box="[159,1401,1772,1794]" gridrow="18" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F728775CEF8FE" box="[159,232,1772,1794]" gridcol="0" gridrow="18" pageId="2" pageNumber="279">EN</th>
<td id="76D85DCA3175BED2FEEF7287748FF8FE" box="[319,425,1772,1794]" gridcol="1" gridrow="18" pageId="2" pageNumber="279">2.8</td>
<td id="76D85DCA3175BED2FE337287776BF8FE" box="[483,589,1772,1794]" gridcol="2" gridrow="18" pageId="2" pageNumber="279">2.5</td>
<td id="76D85DCA3175BED2FD58728777DEF8FE" box="[648,760,1772,1794]" gridcol="3" gridrow="18" pageId="2" pageNumber="279">2.3</td>
<td id="76D85DCA3175BED2FCF8728776BFF8FE" box="[808,921,1772,1794]" gridcol="4" gridrow="18" pageId="2" pageNumber="279">2.4</td>
<td id="76D85DCA3175BED2FC187287711FF8FE" box="[968,1081,1772,1794]" gridcol="5" gridrow="18" pageId="2" pageNumber="279">2.7</td>
<td id="76D85DCA3175BED2FBB8728771FFF8FE" box="[1128,1241,1772,1794]" gridcol="6" gridrow="18" pageId="2" pageNumber="279">2.2</td>
<td id="76D85DCA3175BED2FAD87287705FF8FE" box="[1288,1401,1772,1794]" gridcol="7" gridrow="18" pageId="2" pageNumber="279">2.3</td>
</tr>
<tr id="350934B63175BED2FF4F737E705FF8D7" box="[159,1401,1813,1835]" gridrow="19" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F737E75CEF8D7" box="[159,232,1813,1835]" gridcol="0" gridrow="19" pageId="2" pageNumber="279">EE</th>
<td id="76D85DCA3175BED2FEEF737E748FF8D7" box="[319,425,1813,1835]" gridcol="1" gridrow="19" pageId="2" pageNumber="279">4.4</td>
<td id="76D85DCA3175BED2FE33737E776BF8D7" box="[483,589,1813,1835]" gridcol="2" gridrow="19" pageId="2" pageNumber="279">4.5</td>
<td id="76D85DCA3175BED2FD58737E77DEF8D7" box="[648,760,1813,1835]" gridcol="3" gridrow="19" pageId="2" pageNumber="279">4.2</td>
<td id="76D85DCA3175BED2FCF8737E76BFF8D7" box="[808,921,1813,1835]" gridcol="4" gridrow="19" pageId="2" pageNumber="279">4.3</td>
<td id="76D85DCA3175BED2FC18737E711FF8D7" box="[968,1081,1813,1835]" gridcol="5" gridrow="19" pageId="2" pageNumber="279">4.9</td>
<td id="76D85DCA3175BED2FBB8737E71FFF8D7" box="[1128,1241,1813,1835]" gridcol="6" gridrow="19" pageId="2" pageNumber="279">4.1</td>
<td id="76D85DCA3175BED2FAD8737E705FF8D7" box="[1288,1401,1813,1835]" gridcol="7" gridrow="19" pageId="2" pageNumber="279">4.2</td>
</tr>
<tr id="350934B63175BED2FF4F7355705FF8A8" box="[159,1401,1854,1876]" gridrow="20" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F735575CEF8A8" box="[159,232,1854,1876]" gridcol="0" gridrow="20" pageId="2" pageNumber="279">NS</th>
<td id="76D85DCA3175BED2FEEF7355748FF8A8" box="[319,425,1854,1876]" gridcol="1" gridrow="20" pageId="2" pageNumber="279">1.6</td>
<td id="76D85DCA3175BED2FE337355776BF8A8" box="[483,589,1854,1876]" gridcol="2" gridrow="20" pageId="2" pageNumber="279">1.5</td>
<td id="76D85DCA3175BED2FD58735577DEF8A8" box="[648,760,1854,1876]" gridcol="3" gridrow="20" pageId="2" pageNumber="279">1.5</td>
<td id="76D85DCA3175BED2FCF8735576BFF8A8" box="[808,921,1854,1876]" gridcol="4" gridrow="20" pageId="2" pageNumber="279">1.5</td>
<td id="76D85DCA3175BED2FC187355711FF8A8" box="[968,1081,1854,1876]" gridcol="5" gridrow="20" pageId="2" pageNumber="279">1.7</td>
<td id="76D85DCA3175BED2FBB8735571FFF8A8" box="[1128,1241,1854,1876]" gridcol="6" gridrow="20" pageId="2" pageNumber="279">1.5</td>
<td id="76D85DCA3175BED2FAD87355705FF8A8" box="[1288,1401,1854,1876]" gridcol="7" gridrow="20" pageId="2" pageNumber="279">1.5</td>
</tr>
<tr id="350934B63175BED2FF4F7303705FF882" box="[159,1401,1896,1918]" gridrow="21" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F730375CEF882" box="[159,232,1896,1918]" gridcol="0" gridrow="21" pageId="2" pageNumber="279">SL</th>
<td id="76D85DCA3175BED2FEEF7303748FF882" box="[319,425,1896,1918]" gridcol="1" gridrow="21" pageId="2" pageNumber="279">3.8</td>
<td id="76D85DCA3175BED2FE337303776BF882" box="[483,589,1896,1918]" gridcol="2" gridrow="21" pageId="2" pageNumber="279">4.0</td>
<td id="76D85DCA3175BED2FD58730377DEF882" box="[648,760,1896,1918]" gridcol="3" gridrow="21" pageId="2" pageNumber="279">3.7</td>
<td id="76D85DCA3175BED2FCF8730376BFF882" box="[808,921,1896,1918]" gridcol="4" gridrow="21" pageId="2" pageNumber="279">3.8</td>
<td id="76D85DCA3175BED2FC187303711FF882" box="[968,1081,1896,1918]" gridcol="5" gridrow="21" pageId="2" pageNumber="279">4.3</td>
<td id="76D85DCA3175BED2FBB8730371FFF882" box="[1128,1241,1896,1918]" gridcol="6" gridrow="21" pageId="2" pageNumber="279">3.6</td>
<td id="76D85DCA3175BED2FAD87303705FF882" box="[1288,1401,1896,1918]" gridcol="7" gridrow="21" pageId="2" pageNumber="279">3.6</td>
</tr>
<tr id="350934B63175BED2FF4F73FA705FF85B" box="[159,1401,1937,1959]" gridrow="22" pageId="2" pageNumber="279">
<th id="76D85DCA3175BED2FF4F73FA75CEF85B" box="[159,232,1937,1959]" gridcol="0" gridrow="22" pageId="2" pageNumber="279">NN</th>
<td id="76D85DCA3175BED2FEEF73FA748FF85B" box="[319,425,1937,1959]" gridcol="1" gridrow="22" pageId="2" pageNumber="279">2.2</td>
<td id="76D85DCA3175BED2FE3373FA776BF85B" box="[483,589,1937,1959]" gridcol="2" gridrow="22" pageId="2" pageNumber="279">2.3</td>
<td id="76D85DCA3175BED2FD5873FA77DEF85B" box="[648,760,1937,1959]" gridcol="3" gridrow="22" pageId="2" pageNumber="279">2.1</td>
<td id="76D85DCA3175BED2FCF873FA76BFF85B" box="[808,921,1937,1959]" gridcol="4" gridrow="22" pageId="2" pageNumber="279">2.2</td>
<td id="76D85DCA3175BED2FC1873FA711FF85B" box="[968,1081,1937,1959]" gridcol="5" gridrow="22" pageId="2" pageNumber="279">2.5</td>
<td id="76D85DCA3175BED2FBB873FA71FFF85B" box="[1128,1241,1937,1959]" gridcol="6" gridrow="22" pageId="2" pageNumber="279">2.1</td>
<td id="76D85DCA3175BED2FAD873FA705FF85B" box="[1288,1401,1937,1959]" gridcol="7" gridrow="22" pageId="2" pageNumber="279">2.0</td>
</tr>
</table>
</paragraph>
<caption id="DF46667C3174412EFF4772757180F98E" httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="3" pageNumber="280" targetBox="[151,1436,195,1541]" targetPageId="3">
<paragraph id="8B8636F43174412EFF4772757180F98E" blockId="3.[151,1436,1566,1650]" pageId="3" pageNumber="280">
<emphasis id="B94DEAE63174412EFF477275742DF9C8" bold="true" box="[151,267,1566,1589]" pageId="3" pageNumber="280">FIGURE 1</emphasis>
. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of 
<taxonomicName id="4C394D773174412EFBB072747062F9C9" box="[1120,1348,1567,1589]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="280" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE63174412EFBB072747062F9C9" box="[1120,1348,1567,1589]" italics="true" pageId="3" pageNumber="280">Philautus kakipanjang</emphasis>
</taxonomicName>
(NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of 
<taxonomicName id="4C394D773174412EFEC0723774D2F98E" box="[272,500,1628,1650]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="280" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE63174412EFEC0723774D2F98E" box="[272,500,1628,1650]" italics="true" pageId="3" pageNumber="280">Philautus kakipanjang</emphasis>
</taxonomicName>
(NMBE 1060432) in life, showing the colouration during the night.
</paragraph>
</caption>
<paragraph id="8B8636F43174412EFF1772F47187F8DF" blockId="3.[151,1436,1695,2010]" pageId="3" pageNumber="280">
Dorsal surfaces of head, trunk and limbs smooth with several scattered conical tubercles (
<figureCitation id="13022A713174412EFB0872F4701CF94B" box="[1240,1338,1695,1719]" captionStart="FIGURE 1" captionStartId="3.[151,250,1566,1589]" captionTargetBox="[151,1436,195,1541]" captionTargetId="figure@3.[151,1436,195,1543]" captionTargetPageId="3" captionText="FIGURE 1. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of Philautus kakipanjang (NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of Philautus kakipanjang (NMBE 1060432) in life, showing the colouration during the night." httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="3" pageNumber="280">Figure 1</figureCitation>
); lateral sides of trunk shagreened; ventral side of head, trunk and thighs areolate, of tibia, tarsus and arms smooth; supratympanic fold thick and conspicuous, slightly curved, extending from posterior edge of eye to level of insertion of forelimb; heel without dermal appendages; vent covered with small dermal flap.
</paragraph>
<paragraph id="8B8636F431744129FF177344712CFEE0" blockId="3.[151,1436,1695,2010]" lastBlockId="4.[151,1437,151,575]" lastPageId="4" lastPageNumber="281" pageId="3" pageNumber="280">
Forelimbs slender; hand moderately large (HND/SVL 0.28); tips of fingers enlarged into broad oval disks, those of Fingers III and IV twice as wide as penultimate phalanges (
<figureCitation id="13022A713174412EFC6A73387106F897" box="[954,1056,1875,1899]" captionStart="FIGURE 2" captionStartId="4.[151,250,1779,1802]" captionTargetBox="[151,1436,628,1758]" captionTargetId="figure@4.[151,1436,628,1758]" captionTargetPageId="4" captionText="FIGURE 2. Volar view of right hand (left) and plantar view of right foot (right) of the preserved holotype of Philautus kakipanjang (NMBE 1060433). Scale bar represents 5 mm." httpUri="https://zenodo.org/record/221506/files/figure.png" pageId="3" pageNumber="280">Figure 2</figureCitation>
); each disc with circummarginal groove; relative length of fingers: I &lt;II &lt;IV &lt;III; subarticular tubercles rounded, well developed, numbering one on Fingers I and II, two on Fingers III and IV, with proximal tubercles on Fingers III and IV much smaller than distal ones; rudimentary webbing between fingers, formula being 
<emphasis id="B94DEAE63174412EFCA673A976A7F826" bold="true" box="[886,897,1986,2010]" pageId="3" pageNumber="280">I</emphasis>
2.5-2.75 
<emphasis id="B94DEAE63174412EFC0E73A976D2F826" bold="true" box="[990,1012,1986,2010]" pageId="3" pageNumber="280">II</emphasis>
2.5-3.75 
<date id="FF8710343174412EFB8173A971A6F826" box="[1105,1152,1986,2010]" pageId="3" pageNumber="280">
<emphasis id="B94DEAE63174412EFB8173A97154F826" bold="true" box="[1105,1138,1986,2010]" pageId="3" pageNumber="280">III</emphasis>
3
</date>
- -2.5 
<emphasis id="B94DEAE63174412EFB6173A971F6F826" bold="true" box="[1201,1232,1986,2010]" pageId="3" pageNumber="280">IV</emphasis>
(
<figureCitation id="13022A713174412EFB0F73A97067F826" box="[1247,1345,1986,2010]" captionStart="FIGURE 2" captionStartId="4.[151,250,1779,1802]" captionTargetBox="[151,1436,628,1758]" captionTargetId="figure@4.[151,1436,628,1758]" captionTargetPageId="4" captionText="FIGURE 2. Volar view of right hand (left) and plantar view of right foot (right) of the preserved holotype of Philautus kakipanjang (NMBE 1060433). Scale bar represents 5 mm." httpUri="https://zenodo.org/record/221506/files/figure.png" pageId="3" pageNumber="280">Figure 2</figureCitation>
); thenar tubercle distinct, about one-fourth size of thenar; palmar tubercles absent; nuptial pad on dorsal surface of Fingers I, II and III, most conspicuously developed as prominent, small (about the size of disc of Finger I), smooth pad on proximal phalanx of Finger I, otherwise as callous tissue on penultimate phalanx of Finger I and penultimate and proximal phalanx of Finger II, and as callous, scattered tubercles on Finger III.
</paragraph>
<paragraph id="8B8636F431734129FF17754C715DFDC3" blockId="4.[151,1437,151,575]" pageId="4" pageNumber="281">
Hindlimbs slender, long (LEG/SVL 1.72); tibiotarsal articulation reaching about one ED beyond tip of snout when legs are adpressed forwardly to body; tibiofibula long (TFL/SVL 0.56), longer than thigh (TFL/THL 1.06); heels overlapping each other slightly when knees flexed and thighs held perpendicularly to median plane; foot shorter than tibiofibula (FOT/TFL 0.79); tips of toes enlarged into broad oval disks, each with circummarginal groove; discs of toes smaller than those of fingers; relative length of toes: I &lt;II &lt;III &lt;V &lt;IV; subarticular tubercles numbering one on Toes I and II, two on Toes III and V, and three on Toe IV; pedal webbing formula 
<date id="FF87103431734129FAA475B470ABFE0B" box="[1396,1421,479,503]" pageId="4" pageNumber="281">
<emphasis id="B94DEAE631734129FAA475B47059FE0B" bold="true" box="[1396,1407,479,503]" pageId="4" pageNumber="281">I</emphasis>
2
</date>
- - 2.25 
<emphasis id="B94DEAE631734129FF18766875F8FDE7" bold="true" box="[200,222,515,539]" pageId="4" pageNumber="281">II</emphasis>
1.5-2.75 
<emphasis id="B94DEAE631734129FEEB7668747AFDE7" bold="true" box="[315,348,515,539]" pageId="4" pageNumber="281">III</emphasis>
1.75- 
<date id="FF87103431734129FE4776697482FDE6" box="[407,420,514,538]" pageId="4" pageNumber="281">3</date>
<emphasis id="B94DEAE631734129FE74766874E5FDE7" bold="true" box="[420,451,515,539]" pageId="4" pageNumber="281">IV</emphasis>
2.75-1.5 
<emphasis id="B94DEAE631734129FDF076687712FDE7" bold="true" box="[544,564,515,539]" pageId="4" pageNumber="281">V</emphasis>
(
<figureCitation id="13022A7131734129FD967669778DFDE6" box="[582,683,514,538]" captionStart="FIGURE 2" captionStartId="4.[151,250,1779,1802]" captionTargetBox="[151,1436,628,1758]" captionTargetId="figure@4.[151,1436,628,1758]" captionTargetPageId="4" captionText="FIGURE 2. Volar view of right hand (left) and plantar view of right foot (right) of the preserved holotype of Philautus kakipanjang (NMBE 1060433). Scale bar represents 5 mm." httpUri="https://zenodo.org/record/221506/files/figure.png" pageId="4" pageNumber="281">Figure 2</figureCitation>
); inner metatarsal tubercle relatively large (length 1.0 mm), onethird of length of Toe I (3.0 mm), elongated, prominent; outer metatarsal tubercle absent.
</paragraph>
<caption id="DF46667C31734129FF47729877D4F8D4" httpUri="https://zenodo.org/record/221506/files/figure.png" pageId="4" pageNumber="281" targetBox="[151,1436,628,1758]" targetPageId="4">
<paragraph id="8B8636F431734129FF47729877D4F8D4" blockId="4.[151,1436,1779,1832]" pageId="4" pageNumber="281">
<emphasis id="B94DEAE631734129FF4772987437F8F5" bold="true" box="[151,273,1779,1802]" pageId="4" pageNumber="281">FIGURE 2</emphasis>
. Volar view of right hand (left) and plantar view of right foot (right) of the preserved holotype of 
<taxonomicName id="4C394D7731734129FAEB729F7432F8D4" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="281" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE631734129FAEB729F7432F8D4" italics="true" pageId="4" pageNumber="281">Philautus kakipanjang</emphasis>
</taxonomicName>
(NMBE 1060433). Scale bar represents 5 mm.
</paragraph>
</caption>
<paragraph id="8B8636F431734128FF1773247126FEC3" blockId="4.[151,1436,1871,2004]" lastBlockId="5.[151,1437,151,644]" lastPageId="5" lastPageNumber="282" pageId="4" pageNumber="281">
<emphasis id="B94DEAE631734129FF177324747EF894" bold="true" box="[199,344,1871,1896]" pageId="4" pageNumber="281">Colouration</emphasis>
: Dorsal colouration in life varied strikingly between night and day (
<figureCitation id="13022A7131734129FB8F732471E6F89B" box="[1119,1216,1871,1895]" captionStart="FIGURE 1" captionStartId="3.[151,250,1566,1589]" captionTargetBox="[151,1436,195,1541]" captionTargetId="figure@3.[151,1436,195,1543]" captionTargetPageId="3" captionText="FIGURE 1. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of Philautus kakipanjang (NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of Philautus kakipanjang (NMBE 1060432) in life, showing the colouration during the night." httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="4" pageNumber="281">Figure 1</figureCitation>
). During the night, dorsum reddish-cream-coloured with faint greyish-brown markings; light brown stripes along the canthus rostralis, between the eyes, and parallel to spine on dorsum; limbs with faint crossbars. During the day, dorsum light brown with dark brown, conspicuous markings; sides of the head below canthus rostralis, eye, and supratympanic fold almost uniformly dark brown; tympanum yellowish brown; interorbital stripe, crossbars on limbs, and stripes on the back darker and wider than during the night; additional dark brown flecks on dorsum visible. 
<collectingCountry id="F32E766431724128FADC74D77063FF28" box="[1292,1349,188,212]" name="China" pageId="5" pageNumber="282">Chin</collectingCountry>
, throat, chest, and ventral surfaces of forelimbs whitish, speckled with yellow, flanks yellow. Venter and ventral surfaces of hindlimbs mostly unpigmented. Preaxial side of thigh and groin weakly speckled with yellow markings. Iris golden. In preservative, pattern and colouration similar to diurnal colouration.
</paragraph>
<paragraph id="8B8636F431724128FF17752077A2FE08" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="282">
<emphasis id="B94DEAE631724128FF1775207413FE98" bold="true" box="[199,309,331,356]" pageId="5" pageNumber="282">Variation</emphasis>
: The 
<typeStatus id="5482885631724128FE50752674CDFE99" box="[384,491,333,357]" pageId="5" pageNumber="282" type="paratype">paratypes</typeStatus>
are very similar to the 
<typeStatus id="5482885631724128FCD67527764EFE98" box="[774,872,332,356]" pageId="5" pageNumber="282" type="holotype">holotype</typeStatus>
in measurements and proportions (
<tableCitation id="C6BB034F31724128FAD57527707AFE98" box="[1285,1372,332,356]" captionStart="TABLE 1" captionStartId="2.[151,239,977,1001]" captionTargetBox="[159,1401,1028,1959]" captionTargetPageId="2" captionText="TABLE 1. Measurements of the type series of Philautus kakipanjang sp. n. For abbreviations see Material &amp; Methods." httpUri="http://table.plazi.org/id/DF46667C3175412FFF4777BA70B7FC14" pageId="5" pageNumber="282" tableUuid="DF46667C3175412FFF4777BA70B7FC14">Table 1</tableCitation>
). All 
<typeStatus id="5482885631724128FF47751B7424FE74" box="[151,258,368,392]" pageId="5" pageNumber="282" type="paratype">paratypes</typeStatus>
have a median subgular vocal sac and small, smooth, low nuptial pads. Dorsal pattern varies considerably (
<figureCitation id="13022A7131724128FEE775FF74BEFE50" box="[311,408,404,428]" captionStart="FIGURE 1" captionStartId="3.[151,250,1566,1589]" captionTargetBox="[151,1436,195,1541]" captionTargetId="figure@3.[151,1436,195,1543]" captionTargetPageId="3" captionText="FIGURE 1. Left column: Dorsolateral (A), dorsal (B), and ventral (C) views of the holotype of Philautus kakipanjang (NMBE 1060433) in life, showing the daytime colouration. Right column: Dorsolateral (D), dorsal (E), and ventral (F) views of a paratype of Philautus kakipanjang (NMBE 1060432) in life, showing the colouration during the night." httpUri="https://zenodo.org/record/221505/files/figure.png" pageId="5" pageNumber="282">Figure 1</figureCitation>
). One 
<typeStatus id="5482885631724128FE3B75FE776DFE51" box="[491,587,405,429]" pageId="5" pageNumber="282" type="paratype">paratype</typeStatus>
(NMBE 1060429) has a broad longitudinal mid-dorsal band, another one (NMBE 1060431) a fine mid-dorsal stripe. Mottling on the ventral side and yellow colouration on thigh is variable. So far, females of this species are unknown.
</paragraph>
<paragraph id="8B8636F431724128FF17766B707FFD78" blockId="5.[151,1437,151,644]" pageId="5" pageNumber="282">
<emphasis id="B94DEAE631724128FF17766B748CFDE4" bold="true" box="[199,426,511,536]" pageId="5" pageNumber="282">Molecular genetics</emphasis>
: Comparison of a fragment of the 16S rRNA sequence showed that the species differs considerably from all other species of the genus for which homologous information is available. The new species differs in the uncorrected p-distance from the most closely related Bornean and 
<collectingCountry id="F32E766431724128FBF2762C71B3FDA3" box="[1058,1173,583,607]" name="Philippines" pageId="5" pageNumber="282">Philippine</collectingCountry>
species by 9.9–13.4 % (Table 2). Between the comparative species examined, p-distances ranged between 3.5 and 16.0 % (Table 2).
</paragraph>
<caption id="DF46667C31724128FF477369705CF8CB" httpUri="https://zenodo.org/record/221507/files/figure.png" pageId="5" pageNumber="282" targetBox="[177,1393,737,1751]" targetPageId="5">
<paragraph id="8B8636F431724128FF477369705CF8CB" blockId="5.[151,1436,1794,1847]" pageId="5" pageNumber="282">
<emphasis id="B94DEAE631724128FF4773697437F8E4" bold="true" box="[151,273,1794,1817]" pageId="5" pageNumber="282">FIGURE 3</emphasis>
. Audiospectrogram (top) and corresponding oscillogram (bottom) of an advertisement call of the holotype of 
<taxonomicName id="4C394D7731724128FF47734A745DF8CB" box="[151,379,1825,1847]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="282" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE631724128FF47734A745DF8CB" box="[151,379,1825,1847]" italics="true" pageId="5" pageNumber="282">Philautus kakipanjang</emphasis>
</taxonomicName>
(NMBE 1060433), recorded at 18.5 °C at Gunung Serapi, Kubah National Park, Sarawak, Malaysia.
</paragraph>
</caption>
<paragraph id="8B8636F43172412BFF177335713AFDEB" blockId="5.[151,1436,1886,2018]" lastBlockId="6.[151,1436,151,751]" lastPageId="6" lastPageNumber="283" pageId="5" pageNumber="282">
<emphasis id="B94DEAE631724128FF177335748BF88B" bold="true" box="[199,429,1886,1911]" pageId="5" pageNumber="282">Advertisement call</emphasis>
: 34 advertisement calls from six individuals including the 
<typeStatus id="5482885631724128FBBF733471F7F88B" box="[1135,1233,1887,1911]" pageId="5" pageNumber="282" type="holotype">holotype</typeStatus>
and three of the 
<typeStatus id="5482885631724128FF4773E87424F867" box="[151,258,1923,1947]" pageId="5" pageNumber="282" type="paratype">paratypes</typeStatus>
were recorded in the field (air temperature was 18.5 °C at Gunung Serapi and 18.1°C at Gunung Gading). The advertisement call consists of a single note composed of 9–12 (median 11) pulses and lasting 342– 349 ms (nine pulses), 377–383 ms (ten pulses), 420–425 ms (eleven pulses) or 462–478 ms (twelve pulses). Individual pulses had a length of 4 ms. In one-third of the calls, the note was followed by one or two additional pulses after an interval of 155–220 ms. Pulse repetition rate in the main note ranged between 22.7/s and 26.3/s (mean 24.7/s). In about half of the calls, pulse repetition rate at the beginning of the call was lower than at the end of the note. In the other half of the calls, pulse repetition rate was consistent within the note. In the three cases in which two pulses were added to the note after an interval, the additional two pulses were emitted at a rate of 25.0, 27.0, and 35.7/s. Calls were amplitude-modulated. Amplitude of the pulses was lowest at the beginning of the note, reached maximum at about the middle of the call and remained at that level until the end of the call (
<figureCitation id="13022A713171412BFB297504707DFE7B" box="[1273,1371,367,391]" captionStart="FIGURE 3" captionStartId="5.[151,250,1794,1817]" captionTargetBox="[177,1393,737,1751]" captionTargetId="figure@5.[151,1435,697,1773]" captionTargetPageId="5" captionText="FIGURE 3. Audiospectrogram (top) and corresponding oscillogram (bottom) of an advertisement call of the holotype of Philautus kakipanjang (NMBE 1060433), recorded at 18.5 ° C at Gunung Serapi, Kubah National Park, Sarawak, Malaysia." httpUri="https://zenodo.org/record/221507/files/figure.png" pageId="6" pageNumber="283">Figure 3</figureCitation>
). The note was slightly frequency-modulated and the first two or three pulses had a lower dominant frequency (energy maximum at 2180 ± 150 [2000–2500] Hz) than the rest of the pulses (energy maximum at 2518 ± 121 [2300–2700] Hz). Increase of frequency within a note was 339 ± 200 (100–700) Hz. When additional pulses were added, their frequency was as high as the frequency of the pulses at the end of the main note.
</paragraph>
<paragraph id="8B8636F43171412BFF1776487193FD13" blockId="6.[151,1436,151,751]" pageId="6" pageNumber="283">
<emphasis id="B94DEAE63171412BFF17764874CDFDC0" bold="true" box="[199,491,547,572]" pageId="6" pageNumber="283">Habitat and distribution</emphasis>
: We found all specimens after dusk, calling from low bushes and from tops of small trees, ca. 
<quantity id="4CC19B113171412BFED3762C7446FDA3" box="[259,352,583,607]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="6" pageNumber="283" unit="m" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 m</quantity>
above the ground. We heard the call of the species only in the summit regions of both Gunung Serapi and Gunung Gading from approximately 
<quantity id="4CC19B113171412BFD1676077629FD78" box="[710,783,620,644]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="283" unit="m" value="750.0">750 m</quantity>
upwards, where the lowland mixed dipterocarp forest is replaced by the lower montane forest (
<figureCitation id="13022A713171412BFD8C76E477E5FD5B" box="[604,707,655,679]" captionStart="FIGURE 4" captionStartId="6.[151,250,1758,1781]" captionTargetBox="[156,1433,812,1733]" captionTargetId="figure@6.[151,1436,805,1737]" captionTargetPageId="6" captionText="FIGURE 4. Localities of Philautus kakipanjang in western Sarawak, Borneo at Gunung Gading (1), Gunung Serapi (2), and Gunung Santubong (3)." httpUri="https://zenodo.org/record/221508/files/figure.png" pageId="6" pageNumber="283">Figure 4</figureCitation>
). We recorded calls assignable to the new species at a similar elevation on Gunung Santubong (
<figureCitation id="13022A713171412BFDCB76DF7759FD30" box="[539,639,692,716]" captionStart="FIGURE 4" captionStartId="6.[151,250,1758,1781]" captionTargetBox="[156,1433,812,1733]" captionTargetId="figure@6.[151,1436,805,1737]" captionTargetPageId="6" captionText="FIGURE 4. Localities of Philautus kakipanjang in western Sarawak, Borneo at Gunung Gading (1), Gunung Serapi (2), and Gunung Santubong (3)." httpUri="https://zenodo.org/record/221508/files/figure.png" pageId="6" pageNumber="283">Figure 4</figureCitation>
) but were not able to climb to the calling specimens. The species is probably more widespread in the mountains of western 
<collectingCountry id="F32E76643171412BFCD676BC7640FD13" box="[774,870,727,751]" name="Malaysia" pageId="6" pageNumber="283">Sarawak</collectingCountry>
or even the whole of Borneo.
</paragraph>
</subSubSection>
<caption id="DF46667C3171412BFF4772B574A3F8E8" httpUri="https://zenodo.org/record/221508/files/figure.png" pageId="6" pageNumber="283" targetBox="[156,1433,812,1733]" targetPageId="6">
<paragraph id="8B8636F43171412BFF4772B574A3F8E8" blockId="6.[151,1435,1758,1812]" pageId="6" pageNumber="283">
<emphasis id="B94DEAE63171412BFF4772B57432F908" bold="true" box="[151,276,1758,1781]" pageId="6" pageNumber="283">FIGURE 4.</emphasis>
Localities of 
<taxonomicName id="4C394D773171412BFE7472B477AFF909" box="[420,649,1759,1781]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="283" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE63171412BFE7472B477AFF909" box="[420,649,1759,1781]" italics="true" pageId="6" pageNumber="283">Philautus kakipanjang</emphasis>
</taxonomicName>
in western Sarawak, Borneo at Gunung Gading (1), Gunung Serapi (2), and Gunung Santubong (3).
</paragraph>
</caption>
<subSubSection id="C323657F31714124FF17732A76E2FAA8" lastPageId="9" lastPageNumber="286" pageId="6" pageNumber="283" type="etymology">
<paragraph id="8B8636F43171412BFF17732A7435F85C" blockId="6.[151,1436,1856,1952]" pageId="6" pageNumber="283">
<emphasis id="B94DEAE63171412BFF17732A746EF8A5" bold="true" box="[199,328,1857,1881]" pageId="6" pageNumber="283">Etymology</emphasis>
: The species epithet is composed of the two Malay words 
<emphasis id="B94DEAE63171412BFC01732B7126F8A4" box="[977,1024,1856,1880]" italics="true" pageId="6" pageNumber="283">kaki</emphasis>
, meaning leg, and 
<emphasis id="B94DEAE63171412BFB02732A7008F8A4" box="[1234,1326,1857,1880]" italics="true" pageId="6" pageNumber="283">panjang</emphasis>
, meaning long; in reference to the relatively long hind limbs of the species. The composed word is used as a noun in apposition.
</paragraph>
<paragraph id="8B8636F4317F4125FF1774FC74BDFE98" blockId="8.[151,1437,151,2021]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FF1774FC7440FF4C" bold="true" box="[199,358,151,176]" pageId="8" pageNumber="285">Comparisons</emphasis>
: Morphologically, 
<taxonomicName id="4C394D77317F4125FD9874F3766CFF4C" box="[584,842,152,176]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317F4125FD9874F3766CFF4C" box="[584,842,152,176]" italics="true" pageId="8" pageNumber="285">Philautus kakipanjang</emphasis>
</taxonomicName>
is most similar to species of the 
<taxonomicName id="4C394D77317F4125FB3874F2742CFF28" authority="Schlegel" authorityName="Schlegel" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="aurifasciatus">
<emphasis id="B94DEAE6317F4125FB3874F270BAFF4C" box="[1256,1436,152,176]" italics="true" pageId="8" pageNumber="285">P. aurifasciatus</emphasis>
(Schlegel)
</taxonomicName>
group 
<emphasis id="B94DEAE6317F4125FEB174D674B9FF28" box="[353,415,189,212]" italics="true" pageId="8" pageNumber="285">sensu</emphasis>
<bibRefCitation id="EFA84B05317F4125FE7A74D7771CFF28" author="Dring, J." box="[426,570,188,212]" firstAuthor="Dring" journalOrPublisher="Amphibia-Reptilia" pageId="8" pageNumber="285" pagination="19 - 47" part="8" refId="ref6337" refString="Dring, J. (1987) Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia, 8, 19 - 47. http: // dx. doi. org / 10.1163 / 156853887 x 00036" title="Bornean treefrogs of the genus Philautus (Rhacophoridae)" type="journal article" year="1987">Dring (1987</bibRefCitation>
; see also 
<bibRefCitation id="EFA84B05317F4125FD6174D77665FF28" author="Matsui, M." box="[689,835,188,212]" firstAuthor="Matsui" journalOrPublisher="Zoological Science" pageId="8" pageNumber="285" pagination="437 - 442" part="26" refId="ref6827" refString="Matsui, M. (2009) A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo. Zoological Science, 26, 437 - 442." title="A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo" type="journal article" year="2009">Matsui 2009</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317F4125FC8374D776D5FF28" author="Dehling, J. M." box="[851,1011,188,212]" firstAuthor="Dehling" journalOrPublisher="Salamandra" pageId="8" pageNumber="285" pagination="63 - 72" part="46" refId="ref6193" refString="Dehling, J. M. (2010) A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo). Salamandra, 46, 63 - 72." title="A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo)" type="journal article" year="2010">Dehling 2010</bibRefCitation>
) which are all small-sized, have the musculus cutaneus pectoris and small, smooth nuptial pads (in males), and lack vomerine teeth. It is compared in the following paragraph to all other species that share this combination of characters. All other 
<taxonomicName id="4C394D77317F4125FB6B756F7000FEE0" box="[1211,1318,260,284]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="genus">
<emphasis id="B94DEAE6317F4125FB6B756F7000FEE0" box="[1211,1318,260,284]" italics="true" pageId="8" pageNumber="285">Philautus</emphasis>
</taxonomicName>
species of the Sunda region lack this character combination and therefore can be distinguished morphologically from 
<taxonomicName id="4C394D77317F4125FF47752774B5FE98" box="[151,403,332,356]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317F4125FF47752774B5FE98" box="[151,403,332,356]" italics="true" pageId="8" pageNumber="285">Philautus kakipanjang</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8636F4317F4124FF17751B772AFE08" blockId="8.[151,1437,151,2021]" lastBlockId="9.[151,1436,151,1364]" lastPageId="9" lastPageNumber="286" pageId="8" pageNumber="285">
In the following, 
<taxonomicName id="4C394D77317F4125FE46751A7767FE74" box="[406,577,368,392]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317F4125FE46751A7767FE74" box="[406,577,368,392]" italics="true" pageId="8" pageNumber="285">P. kakipanjang</emphasis>
</taxonomicName>
(characters in parentheses) is first compared to Bornean species, then to species with a 
<typeStatus id="54828856317F4125FEEA75FE744CFE51" box="[314,362,405,429]" pageId="8" pageNumber="285">type</typeStatus>
locality outside Borneo. 
<taxonomicName id="4C394D77317F4125FD5275FF7687FE50" authority="Smith" authorityName="Smith" box="[642,929,404,428]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="amoenus">
<emphasis id="B94DEAE6317F4125FD5275FF767EFE50" box="[642,856,404,428]" italics="true" pageId="8" pageNumber="285">Philautus amoenus</emphasis>
Smith
</taxonomicName>
has shorter legs, with TFL/SVL 
<quantity id="4CC19B11317F4125FAC075FF70BAFE50" box="[1296,1436,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.2573" metricValueMax="1.3716" metricValueMin="1.143" pageId="8" pageNumber="285" unit="in" value="0.495" valueMax="0.54" valueMin="0.45">0.45–0.54 in</quantity>
males (0.56–0.64) and the tibiotarsal articulation reaching to tip of snout or slightly beyond (reaching considerably beyond tip of snout); a tubercle present at the tibiotarsal joint (absent); a wider head, HW/SVL 0.40–0.47 (0.38– 0.42); and Toes III and V reaching distinctly beyond the knee when tarsus is adpressed to tibia (toe tips reaching to knee only). 
<taxonomicName id="4C394D77317F4125FECF764F7758FDC0" authority="Matsui" authorityName="Matsui" box="[287,638,548,572]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="davidlabangi">
<emphasis id="B94DEAE6317F4125FECF764F770EFDC0" box="[287,552,548,572]" italics="true" pageId="8" pageNumber="285">Philautus davidlabangi</emphasis>
Matsui
</taxonomicName>
has a longer head, HL/SVL 0.40–0.43 (0.32–0.37); shorter legs, with TFL/SVL 0.52–0.56 (0.56–0.64), tibiotarsal articulation reaching to tip of snout only (reaching considerably beyond tip of snout) and heels meeting or at most slightly overlapping each other (heels overlapping considerably); snout shorter than the eye diameter (longer than eye diameter); interorbital distance shorter than upper eyelid width with IO/EW 0.78–0.90 (greater than EW with IO/EW 1.02–1.21); more extensive toe webbing with 
<emphasis id="B94DEAE6317F4125FAC876DF7005FD30" bold="true" box="[1304,1315,692,716]" pageId="8" pageNumber="285">I</emphasis>
(1.5-1.75)- 2.25 
<date id="FF871034317F4125FF1876B075C0FD0E" box="[200,230,730,755]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FF1876B075F8FD0F" bold="true" box="[200,222,731,755]" pageId="8" pageNumber="285">II</emphasis>
1
</date>
-2.25 
<emphasis id="B94DEAE6317F4125FEF676B07461FD0F" bold="true" box="[294,327,731,755]" pageId="8" pageNumber="285">III</emphasis>
1- 
<date id="FF871034317F4125FE8B76B1744AFD0E" box="[347,364,730,754]" pageId="8" pageNumber="285">2</date>
<emphasis id="B94DEAE6317F4125FEBC76B074ADFD0F" bold="true" box="[364,395,731,755]" pageId="8" pageNumber="285">IV</emphasis>
2– 
<date id="FF871034317F4125FE7B76B174EFFD0F" box="[427,457,730,755]" pageId="8" pageNumber="285">
1 
<emphasis id="B94DEAE6317F4125FE6576B074EFFD0F" bold="true" box="[437,457,731,755]" pageId="8" pageNumber="285">V</emphasis>
</date>
(vs. 
<date id="FF871034317F4125FDC476B0770BFD0E" box="[532,557,730,755]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FDC476B07739FD0F" bold="true" box="[532,543,731,755]" pageId="8" pageNumber="285">I</emphasis>
2
</date>
- -2.25 
<emphasis id="B94DEAE6317F4125FDBD76B077A5FD0F" bold="true" box="[621,643,731,755]" pageId="8" pageNumber="285">II</emphasis>
1.5-2.75 
<emphasis id="B94DEAE6317F4125FD3076B07627FD0F" bold="true" box="[736,769,731,755]" pageId="8" pageNumber="285">III</emphasis>
1.75- 
<date id="FF871034317F4125FCEC76B1766FFD0E" box="[828,841,730,754]" pageId="8" pageNumber="285">3</date>
<emphasis id="B94DEAE6317F4125FC9976B0764EFD0F" bold="true" box="[841,872,731,755]" pageId="8" pageNumber="285">IV</emphasis>
2.75-1.5 
<emphasis id="B94DEAE6317F4125FC1576B176C5FD0F" bold="true" box="[965,995,730,755]" pageId="8" pageNumber="285">V)</emphasis>
; and nuptial pads absent (present). 
<taxonomicName id="4C394D77317F4125FF47769474D5FCEB" authority="Dehling" authorityName="Dehling" box="[151,499,767,791]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="juliandringi">
<emphasis id="B94DEAE6317F4125FF47769474B7FCEB" box="[151,401,767,791]" italics="true" pageId="8" pageNumber="285">Philautus juliandringi</emphasis>
Dehling
</taxonomicName>
has a shorter snout, subequal in length to eye diameter, SL/ED 0.92–1.05 (longer, SL/ED 1.10–1.25) and rounded in dorsal view (acuminate); a relatively larger eye, ED/HL 0.46–0.52 (0.42–0.45); the tympanum is smaller, TD/ED 0.32–0.39 (0.39–0.48) and is covered by a thick layer of skin (conspicuously visible, free except posterodorsal fourth); and the toe webbing is a little less extensive, 
<date id="FF871034317F4125FB4677057189FC79" box="[1174,1199,877,902]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FB4677057187FC7A" bold="true" box="[1174,1185,878,902]" pageId="8" pageNumber="285">I</emphasis>
2
</date>
-2.5 
<emphasis id="B94DEAE6317F4125FB0B770571D7FC7A" bold="true" box="[1243,1265,878,902]" pageId="8" pageNumber="285">II</emphasis>
(1.5–2-)-3 
<date id="FF871034317F4125FA8F770570ABFC79" box="[1375,1421,877,902]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FA8F770570A6FC7A" bold="true" box="[1375,1408,878,902]" pageId="8" pageNumber="285">III</emphasis>
2
</date>
- - 3+ 
<emphasis id="B94DEAE6317F4125FF7D77FE75EAFC51" bold="true" box="[173,204,917,941]" pageId="8" pageNumber="285">IV</emphasis>
(3+–3)- 
<date id="FF871034317F4125FECA77FF741BFC51" box="[282,317,916,941]" pageId="8" pageNumber="285">
2 
<emphasis id="B94DEAE6317F4125FEF977FE741BFC51" bold="true" box="[297,317,917,941]" pageId="8" pageNumber="285">V</emphasis>
</date>
(vs. 
<date id="FF871034317F4125FEAB77FE74B5FC50" box="[379,403,916,941]" pageId="8" pageNumber="285">
<emphasis id="B94DEAE6317F4125FEAB77FE74A0FC51" bold="true" box="[379,390,917,941]" pageId="8" pageNumber="285">I</emphasis>
2
</date>
- -2.25 
<emphasis id="B94DEAE6317F4125FE0377FE74CFFC51" bold="true" box="[467,489,917,941]" pageId="8" pageNumber="285">II</emphasis>
1.5-2.75 
<emphasis id="B94DEAE6317F4125FD9677FE7741FC51" bold="true" box="[582,615,917,941]" pageId="8" pageNumber="285">III</emphasis>
1.75- 
<date id="FF871034317F4125FD7277FF7789FC50" box="[674,687,916,940]" pageId="8" pageNumber="285">3</date>
<emphasis id="B94DEAE6317F4125FD7F77FE77E8FC51" bold="true" box="[687,718,917,941]" pageId="8" pageNumber="285">IV</emphasis>
2.75-1.5 
<emphasis id="B94DEAE6317F4125FCFB77FF766FFC51" bold="true" box="[811,841,916,941]" pageId="8" pageNumber="285">V)</emphasis>
. 
<taxonomicName id="4C394D77317F4125FCB477FF71ACFC50" authority="Smith" authorityName="Smith" box="[868,1162,916,940]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="mjobergi">
<emphasis id="B94DEAE6317F4125FCB477FF711BFC50" box="[868,1085,916,940]" italics="true" pageId="8" pageNumber="285">Philautus mjobergi</emphasis>
Smith
</taxonomicName>
has a rounded snout in dorsal view (acuminate), which is shorter or subequal to eye diameter (longer than eye diameter); a relatively wider head, HW/HL 1.29–1.34 (1.14–1.21); the tympanum is separated from the eye by a distance equal to TD (almost immediately adjacent to eye, separated by less than one-fifth of TD); and the toe webbing is less extensive, Toes II, III, and V not webbed beyond distal, Toe IV not beyond penultimate subarticular tubercles (toes webbed beyond these tubercles). 
<taxonomicName id="4C394D77317F4125FE8A702377BDFB9C" authority="Boulenger" authorityName="Boulenger" box="[346,667,1096,1120]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B94DEAE6317F4125FE8A7023773AFB9C" box="[346,540,1096,1120]" italics="true" pageId="8" pageNumber="285">Philautus petersi</emphasis>
Boulenger
</taxonomicName>
has shorter legs with the tibiotarsal articulation reaching to tip of snout only (reaches beyond tip of snout); a median lingual process (absent); a conical tubercle at the tibiotarsal joint (absent) and the tympanum is separated from the eye by two-thirds the TD (almost immediately adjacent to eye, separated by less than one-fifth of TD). 
<taxonomicName id="4C394D77317F4125FD4670DF7115FB30" authority="Inger and Stuebing" authorityName="Inger and Stuebing" box="[662,1075,1204,1229]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="refugii">
<emphasis id="B94DEAE6317F4125FD4670DF7672FB30" box="[662,852,1204,1228]" italics="true" pageId="8" pageNumber="285">Philautus refugii</emphasis>
Inger and Stuebing
</taxonomicName>
is smaller, SVL of males 15.3– 18.0 mm (
<quantity id="4CC19B11317F4125FEC270B374E6FB0C" box="[274,448,1240,1264]" metricMagnitude="-2" metricUnit="m" metricValue="2.245" metricValueMax="2.37" metricValueMin="2.12" pageId="8" pageNumber="285" unit="mm" value="22.45" valueMax="23.7" valueMin="21.2">21.2–23.7 mm</quantity>
), has less extensively webbed toes, webbing not reaching beyond distal subarticular tubercles of Toes III and V (reaching beyond these tubercles); and a conical tubercle at the tibiotarsal joint, followed by a row of lower tubercles along the postaxial edge of the tarsus and fifth toe (absent); and the anterior face of the thigh has a barred black and white pattern (unpigmented with bright yellow spots). 
<taxonomicName id="4C394D77317F4125FB31712F74A6FA7C" authority="Malkmus and Riede" authorityName="Malkmus and Riede" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="saueri">
<emphasis id="B94DEAE6317F4125FB31712F70BBFAA0" box="[1249,1437,1348,1372]" italics="true" pageId="8" pageNumber="285">Philautus saueri</emphasis>
Malkmus and Riede
</taxonomicName>
has shorter legs, TFL/SVL 0.54 (0.56–0.64); a wider head, HW/SVL 0.44 (0.39–0.42); a larger interorbital distance with IO/EW 1.90 (0.97–1.21); the nostril situated between the eye and the tip of snout, EN/NS 1.01 (1.45–1.59); and a small tubercle at the tibiotarsal joint (absent). 
<taxonomicName id="4C394D77317F4125FC2771DB71DCFA34" authority="Dring" authorityName="Dring" box="[1015,1274,1456,1480]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="umbra">
<emphasis id="B94DEAE6317F4125FC2771DB7194FA34" box="[1015,1202,1456,1480]" italics="true" pageId="8" pageNumber="285">Philautus umbra</emphasis>
Dring
</taxonomicName>
is larger, SVL of males up to 
<quantity id="4CC19B11317F4125FEEB71BF74B9FA10" box="[315,415,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="8" pageNumber="285" unit="mm" value="35.1">35.1 mm</quantity>
(to 
<quantity id="4CC19B11317F4125FE1B71BF7714FA10" box="[459,562,1492,1516]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="8" pageNumber="285" unit="mm" value="23.7">23.7 mm</quantity>
); has a median lingual process (absent); less developed toe webbing; no finger webbing (rudimentary webbing); broader finger and toe tips; colouration light grey with darker markings during the night and almost uniformly black during the day (light brown to yellowish brown with or without marked dark brown pattern during the night, light brown with darker markings during the day); and has the nuptial pads reduced to glandular patches (nuptial pads small, prominent, rounded). 
<taxonomicName id="4C394D77317F4125FC42720F71D4F980" authority="Peters" authorityName="Peters" box="[914,1266,1636,1660]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="acutirostris">
<emphasis id="B94DEAE6317F4125FC42720F71AAF980" box="[914,1164,1636,1660]" italics="true" pageId="8" pageNumber="285">Philautus acutirostris</emphasis>
(Peters)
</taxonomicName>
(including its synonyms 
<taxonomicName id="4C394D77317F4125FEC672E2773FF95C" authority="Taylor" authorityName="Taylor" box="[278,537,1672,1696]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="basilanensis">
<emphasis id="B94DEAE6317F4125FEC672E274E5F95C" box="[278,451,1672,1696]" italics="true" pageId="8" pageNumber="285">P. basilanensis</emphasis>
Taylor
</taxonomicName>
and 
<taxonomicName id="4C394D77317F4125FD8C72E2761AF95C" authority="Stejneger" authorityName="Stejneger" box="[604,828,1672,1696]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="woodi">
<emphasis id="B94DEAE6317F4125FD8C72E277E5F95C" box="[604,707,1672,1696]" italics="true" pageId="8" pageNumber="285">P. woodi</emphasis>
Stejneger
</taxonomicName>
) from the 
<collectingCountry id="F32E7664317F4125FC1572E37165F95C" box="[965,1091,1672,1696]" name="Philippines" pageId="8" pageNumber="285">Philippines</collectingCountry>
has a sharply pointed snout (acuminate), frequently with a pale-coloured projection at tip (absent at least in males); less extensive toe webbing, Toe IV webbed only to proximal subarticular tubercle (webbed beyond middle tubercle on postaxial side); a median lingual process (absent); and can have vomerine teeth (absent). 
<taxonomicName id="4C394D77317F4125FC34729F71CAF8F0" box="[996,1260,1780,1804]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="aurifasciatus">
<emphasis id="B94DEAE6317F4125FC34729F71CAF8F0" box="[996,1260,1780,1804]" italics="true" pageId="8" pageNumber="285">Philautus aurifasciatus</emphasis>
</taxonomicName>
from Java has shorter legs, TFL/SVL 0.49–0.58 (0.56–0.64), the tibiotarsal articulation reaching to nostril or the tip of snout (reaching beyond tip of snout); and has an NN/EN ratio of 1.11–1.32 (0.87–0.96). 
<taxonomicName id="4C394D77317F4125FBED735770BAF8A8" authority="Boulenger" authorityName="Boulenger" box="[1085,1436,1852,1876]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="leitensis">
<emphasis id="B94DEAE6317F4125FBED7357702BF8A8" box="[1085,1293,1852,1876]" italics="true" pageId="8" pageNumber="285">Philautus leitensis</emphasis>
(Boulenger)
</taxonomicName>
from the 
<collectingCountry id="F32E7664317F4125FED0730B7458F884" box="[256,382,1888,1912]" name="Philippines" pageId="8" pageNumber="285">Philippines</collectingCountry>
has a relatively narrower head with HW/HL 1.03–1.17 (1.14–1.25); an indistinct tympanum (distinct); a prominent tubercle at the tibiotarsal articulation (absent); and lacks finger webbing (rudimentary webbing present). 
<taxonomicName id="4C394D77317F4125FEBA73C377E3F83C" authority="Boulenger" authorityName="Boulenger" box="[362,709,1960,1984]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="285" phylum="Chordata" rank="species" species="longicrus">
<emphasis id="B94DEAE6317F4125FEBA73C37761F83C" box="[362,583,1960,1984]" italics="true" pageId="8" pageNumber="285">Philautus longicrus</emphasis>
Boulenger
</taxonomicName>
has a relatively narrower head, HW/HL 1.08–1.16 (1.14–1.21); the internarial distance is larger than the distance from eye to nostril NN/EN 1.09–1.26 (NN shorter than EN, 0.87– 0.96) and larger in relation to head width, NN/HW 0.26–0.29 (0.23–0.26); the nostril is situated a little closer to the tip of snout than to the eye, EN/NS 1.07–1.18 (much closer to tip of snout than to eye, EN/NS 1.45–1.61); the interorbital distance is larger in relation to the width of the upper eyelid IO/EW 1.25–1.34 (0.96–1.21); and the hand is relatively shorter HND/SVL 0.26–0.29 (0.29–0.32). 
<taxonomicName id="4C394D77317E4124FCA8756F71D3FEE0" authority="Boettger" authorityName="Boettger" box="[888,1269,260,284]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="schmackeri">
<emphasis id="B94DEAE6317E4124FCA8756F7155FEE0" box="[888,1139,260,284]" italics="true" pageId="9" pageNumber="286">Philautus schmackeri</emphasis>
(Boettger)
</taxonomicName>
(including 
<taxonomicName id="4C394D77317E4124FA57756E7439FEBC" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="mindorensis">
<emphasis id="B94DEAE6317E4124FA57756E7439FEBC" italics="true" pageId="9" pageNumber="286">P. mindorensis</emphasis>
</taxonomicName>
[Boulenger]) from the 
<collectingCountry id="F32E7664317E4124FDF3754C7787FEC3" box="[547,673,295,319]" name="Philippines" pageId="9" pageNumber="286">Philippines</collectingCountry>
has a relatively narrower head with HW/HL 1.00–1.11 (1.14–1.25); a smaller tympanum with TD/ED 0.20–0.25 (0.39–0.48); and a pointed snout with a conical tip (acuminate without conical tip). 
<taxonomicName id="4C394D77317E4124FEE3751B7782FE7B" authority="Boulenger" authorityName="Boulenger" box="[307,676,367,392]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="B94DEAE6317E4124FEE3751B7728FE74" box="[307,526,368,392]" italics="true" pageId="9" pageNumber="286">Philautus parvulus</emphasis>
(Boulenger)
</taxonomicName>
from 
<collectingCountry id="F32E7664317E4124FD28751B7641FE74" box="[760,871,368,392]" name="Myanmar" pageId="9" pageNumber="286">Myanmar</collectingCountry>
, 
<collectingCountry id="F32E7664317E4124FCA9750476C6FE7B" box="[889,992,367,391]" name="Thailand" pageId="9" pageNumber="286">Thailand</collectingCountry>
, 
<collectingCountry id="F32E7664317E4124FC227504714EFE7B" box="[1010,1128,367,391]" name="Cambodia" pageId="9" pageNumber="286">Cambodia</collectingCountry>
, 
<collectingCountry id="F32E7664317E4124FBAA750471FAFE7B" box="[1146,1244,367,391]" name="Vietnam" pageId="9" pageNumber="286">Vietnam</collectingCountry>
, and Peninsular 
<collectingCountry id="F32E7664317E4124FF4775FF75D8FE50" box="[151,254,404,428]" name="Malaysia" pageId="9" pageNumber="286">Malaysia</collectingCountry>
has shorter legs with TFL/SVL 0.44–0.51 (0.56–0.64) and the tibiotarsal articulation reaches the posterior margin of eye (beyond tip of snout); the heels barely touch when legs are folded at a right angle to the body (overlapping considerably).
</paragraph>
<paragraph id="8B8636F4317E4124FF1775947755FB80" blockId="9.[151,1436,151,1364]" pageId="9" pageNumber="286">
Bioacoustically, 
<taxonomicName id="4C394D77317E4124FE50766B775AFDE4" box="[384,636,512,536]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317E4124FE50766B775AFDE4" box="[384,636,512,536]" italics="true" pageId="9" pageNumber="286">Philautus kakipanjang</emphasis>
</taxonomicName>
is also most similar to the Bornean species of the 
<taxonomicName id="4C394D77317E4124FB77766A7072FDE4" box="[1191,1364,512,536]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="aurifasciatus">
<emphasis id="B94DEAE6317E4124FB77766A7072FDE4" box="[1191,1364,512,536]" italics="true" pageId="9" pageNumber="286">P. aurifasciatus</emphasis>
</taxonomicName>
group 
<emphasis id="B94DEAE6317E4124FF47764E75F3FDC0" box="[151,213,549,572]" italics="true" pageId="9" pageNumber="286">sensu</emphasis>
<bibRefCitation id="EFA84B05317E4124FF31764F7455FDC0" author="Dring, J." box="[225,371,548,572]" firstAuthor="Dring" journalOrPublisher="Amphibia-Reptilia" pageId="9" pageNumber="286" pagination="19 - 47" part="8" refId="ref6337" refString="Dring, J. (1987) Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia, 8, 19 - 47. http: // dx. doi. org / 10.1163 / 156853887 x 00036" title="Bornean treefrogs of the genus Philautus (Rhacophoridae)" type="journal article" year="1987">Dring (1987</bibRefCitation>
; see also 
<bibRefCitation id="EFA84B05317E4124FE3E764F77A5FDC0" author="Matsui, M." box="[494,643,548,572]" firstAuthor="Matsui" journalOrPublisher="Zoological Science" pageId="9" pageNumber="286" pagination="437 - 442" part="26" refId="ref6827" refString="Matsui, M. (2009) A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo. Zoological Science, 26, 437 - 442." title="A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo" type="journal article" year="2009">Matsui 2009</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FD44764F7613FDC0" author="Dehling, J. M." box="[660,821,548,572]" firstAuthor="Dehling" journalOrPublisher="Salamandra" pageId="9" pageNumber="286" pagination="63 - 72" part="46" refId="ref6193" refString="Dehling, J. M. (2010) A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo). Salamandra, 46, 63 - 72." title="A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo)" type="journal article" year="2010">Dehling 2010</bibRefCitation>
) and is compared to these species in the following paragraph. Calls of these species are regularly pulsed and not or only slightly frequency-modulated. All other Bornean 
<taxonomicName id="4C394D77317E4124FF2D7607744EFD78" box="[253,360,620,644]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="genus">
<emphasis id="B94DEAE6317E4124FF2D7607744EFD78" box="[253,360,620,644]" italics="true" pageId="9" pageNumber="286">Philautus</emphasis>
</taxonomicName>
species, except 
<taxonomicName id="4C394D77317E4124FDCE760677C1FD79" authority="Inger" authorityName="Inger" box="[542,743,620,645]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="disgregus">
<emphasis id="B94DEAE6317E4124FDCE76067783FD78" box="[542,677,620,644]" italics="true" pageId="9" pageNumber="286">P. disgregus</emphasis>
Inger
</taxonomicName>
and 
<taxonomicName id="4C394D77317E4124FCCE760671C3FD79" authority="Stuebing and Wong" authorityName="Stuebing and Wong" box="[798,1253,620,645]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="erythrophthalmus">
<emphasis id="B94DEAE6317E4124FCCE76067126FD78" box="[798,1024,620,644]" italics="true" pageId="9" pageNumber="286">P. erythrophthalmus</emphasis>
Stuebing and Wong
</taxonomicName>
, whose calls are unknown, can be distinguished from 
<taxonomicName id="4C394D77317E4124FD9276FA77CDFD54" box="[578,747,656,680]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317E4124FD9276FA77CDFD54" box="[578,747,656,680]" italics="true" pageId="9" pageNumber="286">P. kakipanjang</emphasis>
</taxonomicName>
by their strongly differing advertisement calls (
<bibRefCitation id="EFA84B05317E4124FAC176E470BEFD5B" author="Dring, J." box="[1297,1432,655,679]" firstAuthor="Dring" journalOrPublisher="Amphibia-Reptilia" pageId="9" pageNumber="286" pagination="19 - 47" part="8" refId="ref6337" refString="Dring, J. (1987) Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia, 8, 19 - 47. http: // dx. doi. org / 10.1163 / 156853887 x 00036" title="Bornean treefrogs of the genus Philautus (Rhacophoridae)" type="journal article" year="1987">Dring 1987</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FF4776DF748EFD30" author="Malkmus, R. &amp; Riede, K." box="[151,424,692,716]" firstAuthor="Malkmus" journalOrPublisher="Sauria" pageId="9" pageNumber="286" pagination="27 - 37" part="18" refId="ref6725" refString="Malkmus, R. &amp; Riede, K. (1996) Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.). Sauria, 18 (1), 27 - 37." title="Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.)" type="journal article" year="1996">Malkmus &amp; Riede 1996</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FE6376DF77B8FD30" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[435,670,692,716]" etAl="et al." firstAuthor="Malkmus" journalOrPublisher="Koeltz Scientific Books, Konigstein" pageId="9" pageNumber="286" pagination="424 pp" refId="ref6780" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo). Koeltz Scientific Books, Konigstein, 424 pp." title="Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo)" type="book" year="2002">
Malkmus 
<emphasis id="B94DEAE6317E4124FDF476DE7773FD30" box="[548,597,692,716]" italics="true" pageId="9" pageNumber="286">et al</emphasis>
. 2002
</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FD7976DF7663FD30" author="Dehling, J. M." box="[681,837,692,716]" firstAuthor="Dehling" journalOrPublisher="Salamandra" pageId="9" pageNumber="286" pagination="63 - 72" part="46" refId="ref6193" refString="Dehling, J. M. (2010) A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo). Salamandra, 46, 63 - 72." title="A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo)" type="journal article" year="2010">Dehling 2010</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FC8076DF710AFD30" author="Dehling, J. M. &amp; Yambun, P. &amp; Bin Lakim, M." box="[848,1068,692,716]" etAl="et al." firstAuthor="Dehling" journalOrPublisher="Herpetozoa" pageId="9" pageNumber="286" pagination="51 - 57" part="23" refId="ref6272" refString="Dehling, J. M., Yambun, P. &amp; Bin Lakim, M. (2010) Advertisement calls of Rhacophorus angulirostris AHL, 1927 and Rhacophorus everetti macroscelis BOULENGER, 1896 (Anura: Rhacophoridae) from Gunung Kinabalu, Sabah, East Malaysia (Borneo). Herpetozoa, 23 (1 / 2), 51 - 57." title="Advertisement calls of Rhacophorus angulirostris AHL, 1927 and Rhacophorus everetti macroscelis BOULENGER, 1896 (Anura: Rhacophoridae) from Gunung Kinabalu, Sabah, East Malaysia (Borneo)" type="journal article" year="2010">
Dehling 
<emphasis id="B94DEAE6317E4124FC6176DE76C4FD30" box="[945,994,692,716]" italics="true" pageId="9" pageNumber="286">et al</emphasis>
. 2010
</bibRefCitation>
, unpubl. data of JMD). The calls of the following species differ in main characteristics (in parentheses: number of notes, note length, number of pulses, dominant frequency, ambient temperature, respectively), and thus can be used to distinguish these species from 
<taxonomicName id="4C394D77317E4124FF06774A745BFCC4" box="[214,381,800,824]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317E4124FF06774A745BFCC4" box="[214,381,800,824]" italics="true" pageId="9" pageNumber="286">P. kakipanjang</emphasis>
</taxonomicName>
(1 note, 342–478 ms, 9–12 pulses, 2.0–2.7 kHz, 18.1–18.5°C): 
<taxonomicName id="4C394D77317E4124FBB3774A71C2FCC4" box="[1123,1252,801,824]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="amoenus">
<emphasis id="B94DEAE6317E4124FBB3774A71C2FCC4" box="[1123,1252,801,824]" italics="true" pageId="9" pageNumber="286">P. amoenus</emphasis>
</taxonomicName>
(2–3 notes, 110 ms, 3–4 pulses, 1.8 kHz, 12.5°C; 
<bibRefCitation id="EFA84B05317E4124FDC7772F760AFCA0" author="Malkmus, R. &amp; Riede, K." box="[535,812,836,860]" firstAuthor="Malkmus" journalOrPublisher="Sauria" pageId="9" pageNumber="286" pagination="27 - 37" part="18" refId="ref6725" refString="Malkmus, R. &amp; Riede, K. (1996) Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.). Sauria, 18 (1), 27 - 37." title="Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.)" type="journal article" year="1996">Malkmus &amp; Riede 1996</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FCE8772F710FFCA0" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[824,1065,836,860]" etAl="et al." firstAuthor="Malkmus" journalOrPublisher="Koeltz Scientific Books, Konigstein" pageId="9" pageNumber="286" pagination="424 pp" refId="ref6780" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo). Koeltz Scientific Books, Konigstein, 424 pp." title="Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo)" type="book" year="2002">
Malkmus 
<emphasis id="B94DEAE6317E4124FC7A772E76FBFCA0" box="[938,989,836,860]" italics="true" pageId="9" pageNumber="286">et al</emphasis>
. 2002
</bibRefCitation>
); 
<taxonomicName id="4C394D77317E4124FBED772E71C8FCA0" box="[1085,1262,836,860]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="davidlabangi">
<emphasis id="B94DEAE6317E4124FBED772E71C8FCA0" box="[1085,1262,836,860]" italics="true" pageId="9" pageNumber="286">P. davidlabangi</emphasis>
</taxonomicName>
(1 note, 50–80 ms, 13–18 pulses, 2.9–3.3 kHz, 23.5–24.8°C; 
<bibRefCitation id="EFA84B05317E4124FD4B770C760CFC83" author="Matsui, M." box="[667,810,871,895]" firstAuthor="Matsui" journalOrPublisher="Zoological Science" pageId="9" pageNumber="286" pagination="437 - 442" part="26" refId="ref6827" refString="Matsui, M. (2009) A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo. Zoological Science, 26, 437 - 442." title="A new species of Philautus (Amphibia, Anura, Rhacophoridae) from the lowlands of Sarawak, Western Borneo" type="journal article" year="2009">Matsui 2009</bibRefCitation>
, unpubl. data of JMD); 
<taxonomicName id="4C394D77317E4124FBE5770271F1FC7C" box="[1077,1239,872,896]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="juliandringi">
<emphasis id="B94DEAE6317E4124FBE5770271F1FC7C" box="[1077,1239,872,896]" italics="true" pageId="9" pageNumber="286">P. juliandringi</emphasis>
</taxonomicName>
(1 note, 234–658 ms, 2–9 pulses, 2.6–3.5 kHz, 18.3–19.1°C; 
<bibRefCitation id="EFA84B05317E4124FD5077E77638FC58" author="Dehling, J. M." box="[640,798,908,932]" firstAuthor="Dehling" journalOrPublisher="Salamandra" pageId="9" pageNumber="286" pagination="63 - 72" part="46" refId="ref6193" refString="Dehling, J. M. (2010) A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo). Salamandra, 46, 63 - 72." title="A new bush frog (Anura: Rhacophoridae: Philautus) from Gunung Mulu National Park, East Malaysia (Borneo)" type="journal article" year="2010">Dehling 2010</bibRefCitation>
); 
<taxonomicName id="4C394D77317E4124FCE977E6769EFC58" box="[825,952,908,932]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="mjobergi">
<emphasis id="B94DEAE6317E4124FCE977E6769EFC58" box="[825,952,908,932]" italics="true" pageId="9" pageNumber="286">P. mjobergi</emphasis>
</taxonomicName>
(call series of 1–4 calls, calls consisting of 1–11 notes, 20–70 ms, 2–7 pulses, 2.0–2.8 kHz, 15 °C; 
<bibRefCitation id="EFA84B05317E4124FCF677C47688FC3B" author="Dring, J." box="[806,942,943,967]" firstAuthor="Dring" journalOrPublisher="Amphibia-Reptilia" pageId="9" pageNumber="286" pagination="19 - 47" part="8" refId="ref6337" refString="Dring, J. (1987) Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia, 8, 19 - 47. http: // dx. doi. org / 10.1163 / 156853887 x 00036" title="Bornean treefrogs of the genus Philautus (Rhacophoridae)" type="journal article" year="1987">Dring 1987</bibRefCitation>
); 
<taxonomicName id="4C394D77317E4124FC1577DA7116FC34" box="[965,1072,945,968]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B94DEAE6317E4124FC1577DA7116FC34" box="[965,1072,945,968]" italics="true" pageId="9" pageNumber="286">P. petersi</emphasis>
</taxonomicName>
(1 note, 1180–1300 ms, 51–56 pulses, 1.6–2.2 kHz, 18°C; 
<bibRefCitation id="EFA84B05317E4124FE1977BF776BFC10" author="Dring, J." box="[457,589,980,1004]" firstAuthor="Dring" journalOrPublisher="Amphibia-Reptilia" pageId="9" pageNumber="286" pagination="19 - 47" part="8" refId="ref6337" refString="Dring, J. (1987) Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia, 8, 19 - 47. http: // dx. doi. org / 10.1163 / 156853887 x 00036" title="Bornean treefrogs of the genus Philautus (Rhacophoridae)" type="journal article" year="1987">Dring 1987</bibRefCitation>
); 
<taxonomicName id="4C394D77317E4124FDB177BE77E0FC10" box="[609,710,980,1004]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="refugii">
<emphasis id="B94DEAE6317E4124FDB177BE77E0FC10" box="[609,710,980,1004]" italics="true" pageId="9" pageNumber="286">P. refugii</emphasis>
</taxonomicName>
(1–3 notes, 10–20 ms, 2–5 pulses, 3.5–4.0 kHz, 18.1°C; Dehling &amp; Dehling, unpubl.); 
<taxonomicName id="4C394D77317E4124FE5B779274C8FBEC" box="[395,494,1017,1040]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="saueri">
<emphasis id="B94DEAE6317E4124FE5B779274C8FBEC" box="[395,494,1017,1040]" italics="true" pageId="9" pageNumber="286">P. saueri</emphasis>
</taxonomicName>
(2–4[rarely 5] notes, 30–40 ms, 7–8 pulses, 1.9–2.6 kHz, 15°C; 
<bibRefCitation id="EFA84B05317E4124FB1B779C75F4FBC8" author="Malkmus, R. &amp; Riede, K." firstAuthor="Malkmus" journalOrPublisher="Sauria" pageId="9" pageNumber="286" pagination="27 - 37" part="18" refId="ref6725" refString="Malkmus, R. &amp; Riede, K. (1996) Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.). Sauria, 18 (1), 27 - 37." title="Die Baumfrosche der Gattung Philautus vom Mount Kinabalu - Teil I: Uberblick und die aurifasciatus - Gruppe mit Beschreibung einer neuen Art (Philautus saueri n. sp.)" type="journal article" year="1996">Malkmus &amp; Riede 1996</bibRefCitation>
, 
<bibRefCitation id="EFA84B05317E4124FF0E707774EBFBC8" author="Malkmus, R. &amp; Manthey, U. &amp; Vogel, G. &amp; Hoffmann, P. &amp; Kosuch, J." box="[222,461,1052,1076]" etAl="et al." firstAuthor="Malkmus" journalOrPublisher="Koeltz Scientific Books, Konigstein" pageId="9" pageNumber="286" pagination="424 pp" refId="ref6780" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. &amp; Kosuch, J. (2002) Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo). Koeltz Scientific Books, Konigstein, 424 pp." title="Amphibians &amp; Reptiles of Mount Kinabalu (North Borneo)" type="book" year="2002">
Malkmus 
<emphasis id="B94DEAE6317E4124FE9F707674A7FBC8" box="[335,385,1052,1076]" italics="true" pageId="9" pageNumber="286">et al</emphasis>
. 2002
</bibRefCitation>
); 
<emphasis id="B94DEAE6317E4124FE3170767764FBC8" box="[481,578,1052,1076]" italics="true" pageId="9" pageNumber="286">P. u m b r a</emphasis>
(1 note, 120–587 ms, 2–11 pulses, 1.4–2.1 kHz, 18.3–19.1°C; unpubl. data of JMD). The call of 
<taxonomicName id="4C394D77317E4124FE54702A7724FBA4" box="[388,514,1088,1112]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="parvulus">
<emphasis id="B94DEAE6317E4124FE54702A7724FBA4" box="[388,514,1088,1112]" italics="true" pageId="9" pageNumber="286">P. parvulus</emphasis>
</taxonomicName>
from 
<collectingCountry id="F32E7664317E4124FD977054778CFBAB" box="[583,682,1087,1111]" name="Thailand" pageId="9" pageNumber="286">Thailand</collectingCountry>
differs in consisting of 1–7 single-pulsed notes, each lasting 20–40 ms at unknown temperature (
<bibRefCitation id="EFA84B05317E4124FE0D700F7740FB80" author="Heyer, W. R." box="[477,614,1124,1148]" firstAuthor="Heyer" journalOrPublisher="Fieldiana: Zoology" pageId="9" pageNumber="286" pagination="61 - 82" part="58" refId="ref6470" refString="Heyer, W. R. (1971) Mating calls of some frogs from Thailand. Fieldiana: Zoology, 58, 61 - 82. http: // dx. doi. org / 10.5962 / bhl. title. 2934" title="Mating calls of some frogs from Thailand" type="journal article" year="1971">Heyer 1971</bibRefCitation>
).
</paragraph>
<paragraph id="8B8636F4317E4124FF1770EC76E2FAA8" blockId="9.[151,1436,151,1364]" pageId="9" pageNumber="286">
Finally, the partial 16S rRNA sequence of 
<taxonomicName id="4C394D77317E4124FD7270E376B8FB5C" box="[674,926,1160,1184]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317E4124FD7270E376B8FB5C" box="[674,926,1160,1184]" italics="true" pageId="9" pageNumber="286">Philautus kakipanjang</emphasis>
</taxonomicName>
differs by at least 9.9 % in the uncorrected pdistance from all available sequences of other species of the genus, including the species which are morphologically and bioacoustically most similar. This value is much higher than the distances between morphologically and bioacoustically well distinguished species pairs like 
<taxonomicName id="4C394D77317E4124FC0C709E71AAFAF0" authority="Dring" authorityName="Dring" box="[988,1164,1268,1292]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="acutus">
<emphasis id="B94DEAE6317E4124FC0C709E7164FAF0" box="[988,1090,1269,1292]" italics="true" pageId="9" pageNumber="286">P. acutus</emphasis>
Dring
</taxonomicName>
/ 
<taxonomicName id="4C394D77317E4124FB44709E7041FAF1" authority="Inger" authorityName="Inger" box="[1172,1383,1269,1293]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="aurantium">
<emphasis id="B94DEAE6317E4124FB44709E7003FAF0" box="[1172,1317,1269,1292]" italics="true" pageId="9" pageNumber="286">P. aurantium</emphasis>
Inger
</taxonomicName>
(3.5 %), 
<taxonomicName id="4C394D77317E4124FF1571727472FACC" box="[197,340,1305,1328]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="aurantium">
<emphasis id="B94DEAE6317E4124FF1571727472FACC" box="[197,340,1305,1328]" italics="true" pageId="9" pageNumber="286">P. aurantium</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4C394D77317E4124FE8C717277FFFACC" authority="Inger, Stuebing and Tan" authorityName="Inger, Stuebing and Tan" box="[348,729,1303,1329]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="bunitus">
<emphasis id="B94DEAE6317E4124FE8C717274EFFACC" box="[348,457,1304,1328]" italics="true" pageId="9" pageNumber="286">P. bunitus</emphasis>
Inger, Stuebing and Tan
</taxonomicName>
(4.8 %), or 
<taxonomicName id="4C394D77317E4124FC8E71727160FAD3" authority="Boulenger" authorityName="Boulenger" box="[862,1094,1303,1328]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="everetti">
<emphasis id="B94DEAE6317E4124FC8E717276EAFACC" box="[862,972,1305,1328]" italics="true" pageId="9" pageNumber="286">P. everetti</emphasis>
Boulenger
</taxonomicName>
/ 
<taxonomicName id="4C394D77317E4124FB9E7172704FFAD3" authority="Boulenger" authorityName="Boulenger" box="[1102,1385,1303,1328]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="macroscelis">
<emphasis id="B94DEAE6317E4124FB9E717271C8FACC" box="[1102,1262,1304,1328]" italics="true" pageId="9" pageNumber="286">P. macroscelis</emphasis>
Boulenger
</taxonomicName>
(5.0 %) (Table 2), corroborating the distinct specific status of 
<taxonomicName id="4C394D77317E4124FCC77156769BFAA8" box="[791,957,1340,1364]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="286" phylum="Chordata" rank="species" species="kakipanjang">
<emphasis id="B94DEAE6317E4124FCC77156769BFAA8" box="[791,957,1340,1364]" italics="true" pageId="9" pageNumber="286">P. kakipanjang</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>