treatments-xml/data/03/8E/C9/038EC92D79787E52FF5B6B4B24CEFAF2.xml
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<document id="0C45A424C6AD623734A8ABA62BFA8D1F" ID-DOI="10.26107/RBZ-2023-0047" ID-ISSN="2345-7600" ID-Zenodo-Dep="10271307" ID-ZooBank="821BC4EC-5AF9-4727-84A3-C44839DFBE28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1701819361795" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, Shane T. &amp; Castro, Peter" docDate="2023" docId="038EC92D79787E52FF5B6B4B24CEFAF2" docLanguage="en" docName="RafflesBZool.71.606-631.pdf" docOrigin="Raffles Bulletin of Zoology 71" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Ectaesthesius bifrons Rathbun 1898" docType="treatment" docVersion="2" lastPageNumber="618" masterDocId="FFB7B15579727E5EFFBA6E582531FFA7" masterDocTitle="Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="631" masterPageNumber="606" pageNumber="616" updateTime="1701819819889" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E60520BA0C1077B4086E4C20F510790A">Re-appraisal of the families and subfamilies of Trapezioidea Miers, 1886, with establishment of a new family, Ectaesthesiidae (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="1EEF2E52C87353F30AA51BADC03CCB88">Ng, Peter K. L.</mods:namePart>
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<taxonomicName id="4C2703B879787E54FF5B6B4B27B1FA8B" ID-CoL="38P4F" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[225,640,1299,1324]" class="Malacostraca" family="Xanthidae" genus="Ectaesthesius" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="616" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B953A42979787E54FF5B6B4B27B1FA8B" bold="true" box="[225,640,1299,1324]" pageId="10" pageNumber="616">
<emphasis id="B953A42979787E54FF5B6B4B24FEFA8C" bold="true" box="[225,463,1299,1323]" italics="true" pageId="10" pageNumber="616">Ectaesthesius bifrons</emphasis>
<bibRefCitation id="EFB605CA79787E54FE6D6B4C27B1FA8B" author="Rathbun MJ" box="[471,640,1300,1324]" pageId="10" pageNumber="616" pagination="567 - 616" refId="ref21481" refString="Rathbun MJ (1898) The Brachyura collected by the U. S. Fish Commission steamer Albatross on the voyage from Norfolk, Virginia, to San Francisco, California, 1887 - 1888. Proceedings of the United States National Museum, 21 (1162): 567 - 616, pls. 41 - 44." type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
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<paragraph id="8B98783B79787E54FECF6B6C24DDFAEB" blockId="10.[225,640,1299,1356]" box="[373,492,1332,1356]" pageId="10" pageNumber="616">
(
<figureCitation id="131C64BE79787E54FEC76B6C24D5FAEB" box="[381,484,1332,1356]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[113,148,1231,1253]" captionStartId-1="4.[113,148,1997,2019]" captionStartId-2="6.[113,148,1720,1742]" captionStartId-3="7.[136,171,1138,1160]" captionTargetBox-0="[113,1451,189,1216]" captionTargetBox-1="[114,1452,228,1981]" captionTargetBox-2="[117,1451,182,1704]" captionTargetBox-3="[138,1476,191,1128]" captionTargetId-0="figure-67@2.[113,1451,189,1216]" captionTargetId-1="figure-14@4.[114,1452,228,1981]" captionTargetId-2="figure-14@6.[105,1473,171,1715]" captionTargetId-3="figure-511@7.[111,1493,182,1137]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." captionText-1="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." captionText-2="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." captionText-3="Fig. 4. Ectaesthesius bifrons. A, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands. Left G1 (pleonal view); B, left G1 (sternal view); C, distal part of left G1 (pleonal view); D, distal part of left G1 (sternal view); E, left G2 (pleonal view). Scales: A, B, E = 0.5 mm; C, D = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10271309" figureDoi-1="http://doi.org/10.5281/zenodo.10271313" figureDoi-2="http://doi.org/10.5281/zenodo.10271315" figureDoi-3="http://doi.org/10.5281/zenodo.10271317" httpUri-0="https://zenodo.org/record/10271309/files/figure.png" httpUri-1="https://zenodo.org/record/10271313/files/figure.png" httpUri-2="https://zenodo.org/record/10271315/files/figure.png" httpUri-3="https://zenodo.org/record/10271317/files/figure.png" pageId="10" pageNumber="616">Figs. 14</figureCitation>
)
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<emphasis id="B953A42979787E54FFCB6B2A240CFA20" box="[113,317,1394,1415]" italics="true" pageId="10" pageNumber="616">Ectaesthesius bifrons</emphasis>
<bibRefCitation id="EFB605CA79787E54FEF86B2A2730FA2F" author="Rathbun MJ" box="[322,513,1394,1416]" pageId="10" pageNumber="616" pagination="567 - 616" refId="ref21481" refString="Rathbun MJ (1898) The Brachyura collected by the U. S. Fish Commission steamer Albatross on the voyage from Norfolk, Virginia, to San Francisco, California, 1887 - 1888. Proceedings of the United States National Museum, 21 (1162): 567 - 616, pls. 41 - 44." type="journal article" year="1898">Rathbun, 1898: 591</bibRefCitation>
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, pl. 42 figs. 1214, 1930: 460, text-fig. 76ac;
<treatmentCitation id="0A865E2A79787E54FEED6BD724C6FA02" author="Garth JS" box="[343,503,1423,1445]" page="466" pageId="10" pageNumber="616" year="1946">
<bibRefCitation id="EFB605CA79787E54FEED6BD724C6FA02" author="Garth JS" box="[343,503,1423,1445]" pageId="10" pageNumber="616" pagination="341 - 522" refId="ref19810" refString="Garth JS (1946) Littoral brachyuran fauna of the Galapagos Islands. Allan Hancock Pacific Expeditions, 5 (10): 341 - 522, fig. 1, pls. 49 - 87." type="journal article" year="1946">Garth, 1946: 466</bibRefCitation>
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, pl. 58, figs. 17;
<treatmentCitation id="0A865E2A79787E54FD236BD724A2FA65" author="Hickman CP Jr. &amp; Zimmerman TL" page="116" pageId="10" pageNumber="616" year="2000">
<bibRefCitation id="EFB605CA79787E54FD236BD724A2FA65" author="Hickman CP Jr. &amp; Zimmerman TL" pageId="10" pageNumber="616" refId="ref20127" refString="Hickman CP Jr. &amp; Zimmerman TL (2000) A field guide to crustaceans of Galapagos: an illustrated guidebook to the barnacles, shrimps, lobsters, and crabs of the Galapagos Islands. Sugar Spring Press, Lexington, VA, USA, 156 pp." type="book" year="2000">Hickman &amp; Zimmerman, 2000: 116</bibRefCitation>
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, unnumbered figure;
<treatmentCitation id="0A865E2A79787E54FDE76BF4258CFA79" author="Ng PKL &amp; Guinot D &amp; Davie PJF" page="202" pageId="10" pageNumber="616" year="2008">
<bibRefCitation id="EFB605CA79787E54FDE76BF4258CFA79" author="Ng PKL &amp; Guinot D &amp; Davie PJF" pageId="10" pageNumber="616" pagination="1 - 286" refId="ref21055" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 202</bibRefCitation>
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;
<treatmentCitation id="0A865E2A79787E54FF736B9124D2FA79" author="Poore GCB &amp; Ahyong ST" box="[201,483,1481,1503]" page="728" pageId="10" pageNumber="616" year="2023">
<bibRefCitation id="EFB605CA79787E54FF736B9124D2FA79" author="Poore GCB &amp; Ahyong ST" box="[201,483,1481,1503]" pageId="10" pageNumber="616" refId="ref21418" refString="Poore GCB &amp; Ahyong ST (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, Australia, xii + 916 pp." type="book" year="2023">Poore &amp; Ahyong, 2023: 728</bibRefCitation>
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, 729, 730, fig. 14.139b.
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<emphasis id="B953A42979787E54FFCB685F2457F9B8" bold="true" box="[113,358,1543,1567]" pageId="10" pageNumber="616">Material examined.</emphasis>
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(5.4 × 4.0 mm) (
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), station 346-35, mud and shell bottom, between
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and
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,
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,
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, coll.
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III
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, 1934
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;
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(larger 12.8 × 9.0 mm),
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(11.2 × 8.0 mm),
<materialsCitation id="3B4F726679787E54FD5968DF2749F8B8" collectingDate="1938-01-21" collectionCode="R, V" collectorName="Velero" country="Iceland" county="Galapagos Islands" elevation="70" latitude="-0.27" location="Bahia Sullivan" longLatPrecision="130" longitude="90.58" municipality="James" pageId="10" pageNumber="616" specimenCode="LACM 1938.56" specimenCount="2" specimenCount-juvenile="2">
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(
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), station 795-38, rock and sand substrate,
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(=
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),
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(=
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, coll.
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III
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<collectingDate id="EFDDA71379787E54FE79695F2749F8B8" box="[451,632,1799,1823]" pageId="10" pageNumber="616" value="1938-01-21">21 January 1938</collectingDate>
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female (3.9 ×
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(10.1 × 7.0 mm) (
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), station 795A-38, volcanic sand substrate,
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(=
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(=
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, coll.
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/
<collectionCode id="ED36E0FE79787E54FF3169FF25AFF818" box="[139,158,1959,1983]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="616" type="Museum">V</collectionCode>
<emphasis id="B953A42979787E54FF1F69FF243FF818" box="[165,270,1959,1983]" italics="true" pageId="10" pageNumber="616">
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III
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<collectingDate id="EFDDA71379787E54FEA169FF24FEF818" box="[283,463,1959,1983]" pageId="10" pageNumber="616" value="1938-01-21">21 January 1938</collectingDate>
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;
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1 young female (6.8 × 5.0 mm) (
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), station 814-38, north of
<location id="8EF82EE079787E54FD35699F27C0F878" LSID="urn:lsid:plazi:treatment:038EC92D79787E52FF5B6B4B24CEFAF2:8EF82EE079787E54FD35699F27C0F878" box="[655,753,1991,2015]" country="Iceland" county="Galapagos Islands" latitude="-1.365" longLatPrecision="130" longitude="90.668335" municipality="Hood" name="Espanola" pageId="10" pageNumber="616">Española</location>
(=
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,
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, coll.
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<emphasis id="B953A42979787E54FEF5665F2489F7B8" box="[335,440,2055,2079]" italics="true" pageId="10" pageNumber="616">
<collectorName id="26D21DED79787E54FEF5665F24A7F7B8" box="[335,406,2055,2079]" pageId="10" pageNumber="616">Velero</collectorName>
III
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,
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<collectingDate id="EFDDA71379787E54FE7D665F274EF7B8" box="[455,639,2055,2079]" pageId="10" pageNumber="616" value="1938-01-28">28 January 1938</collectingDate>
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.
</paragraph>
</subSubSection>
<subSubSection id="C33D2BB079787E54FC976EE52174FF72" box="[813,1093,189,214]" pageId="10" pageNumber="616" type="diagnosis">
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<emphasis id="B953A42979787E54FC976EE52690FF72" bold="true" box="[813,929,189,213]" pageId="10" pageNumber="616">Diagnosis.</emphasis>
As for family.
</paragraph>
</subSubSection>
<subSubSection id="C33D2BB079787E52FC976EA527C0FCF2" lastPageId="12" lastPageNumber="618" pageId="10" pageNumber="616" type="description">
<paragraph id="8B98783B79787E54FC976EA521ACFB52" blockId="10.[812,1453,253,1269]" pageId="10" pageNumber="616">
<emphasis id="B953A42979787E54FC976EA52119FEB2" bold="true" box="[813,1064,253,277]" pageId="10" pageNumber="616">Description of male.</emphasis>
Carapace transversely subovate, distinctly wider than long, widest near midlength (
<figureCitation id="131C64BE79787E54FAF26F452097FE92" box="[1352,1446,285,309]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="10" pageNumber="616">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79787E54FC976F652661FEF2" box="[813,848,317,341]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">2A</figureCitation>
,
<figureCitation id="131C64BE79787E54FCE56F6526B3FEF2" box="[863,898,317,341]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
); dorsal surface smooth, gently convex in frontal view, regions poorly demarcated, H-shaped gastrocardiac groove shallow but visible (
<figureCitation id="131C64BE79787E54FBC76F252033FE32" box="[1149,1282,381,405]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Fig. 2A, C</figureCitation>
). Front gently sinuous to almost entire, smooth, weakly rimmed; with very shallow median concavity separating low lobes; lateral lobe low, visible, with rounded margin, barely demarcated from supraorbital margin; supraorbital margin gently concave, without fissure or cleft; outer orbital tooth small but distinct, sharp (
<figureCitation id="131C64BE79787E54FC616C65210FFDF2" box="[987,1086,573,597]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2A</figureCitation>
,
<figureCitation id="131C64BE79787E54FBF76C652140FDF2" box="[1101,1137,573,597]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
). Antero- and posterolateral margins clearly demarcated, margin very finely granulated; anterolateral margin just shorter than posterolateral margin, with 2 triangular teeth, first tooth obtuse, second tooth (marking junction with posterolateral margin) acute, sharp (
<figureCitation id="131C64BE79787E54FC8F6C8526A5FD52" box="[821,916,733,757]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2A</figureCitation>
,
<figureCitation id="131C64BE79787E54FC256C8526F2FD52" box="[927,963,733,757]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
); anterior part of posterolateral margin gently convex before curving to become concave along posterior half (
<figureCitation id="131C64BE79787E54FCDC6D45268CFC92" box="[870,957,797,821]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Fig. 2A</figureCitation>
,
<figureCitation id="131C64BE79787E54FC706D4526DCFC92" box="[970,1005,797,821]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
). Posterior carapace margin short, gently convex to almost straight (
<figureCitation id="131C64BE79787E54FBE36D65218AFCF2" box="[1113,1211,829,853]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="10" pageNumber="616">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79787E54FB716D6521DCFCF2" box="[1227,1261,829,853]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">2A</figureCitation>
,
<figureCitation id="131C64BE79787E54FB466D652011FCF2" box="[1276,1312,829,853]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
). Suborbital, subheptic, pterygostomial, sub-branchial regions smooth (
<figureCitation id="131C64BE79787E54FC8F6D2526A6FC32" box="[821,919,893,917]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2C</figureCitation>
,
<figureCitation id="131C64BE79787E54FC1C6D2526F9FC32" box="[934,968,893,917]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3B</figureCitation>
). Suborbital margin evenly concave, finely granulated, appearing smooth, suborbital tooth low, obtuse, not pronounced (
<figureCitation id="131C64BE79787E54FC4A6DE5217BFC72" box="[1008,1098,957,981]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">Fig. 3B</figureCitation>
). Posterior margin of epistome entire, separated from pterygostomial region by distinct fissure, margin with low median triangular lobe, lateral margin relatively short, entire, gently concave (
<figureCitation id="131C64BE79787E54FAFF6A452096FB92" box="[1349,1447,1053,1077]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2C</figureCitation>
,
<figureCitation id="131C64BE79787E54FC976A65267EFBF2" box="[813,847,1085,1109]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3C</figureCitation>
). Basal antennal article quadrate, mobile, peduncle and flagellum entering hiatus of orbit, flagellum longer than width of orbit (
<figureCitation id="131C64BE79787E54FC296A2526C7FB32" box="[915,1014,1149,1173]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2C</figureCitation>
,
<figureCitation id="131C64BE79787E54FBBF6A252116FB32" box="[1029,1063,1149,1173]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3B</figureCitation>
). Antennules transverse, flagellum folding transversely (
<figureCitation id="131C64BE79787E54FBA86AC52140FB12" box="[1042,1137,1181,1205]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Figs. 2C</figureCitation>
,
<figureCitation id="131C64BE79787E54FBC76AC52190FB12" box="[1149,1185,1181,1205]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
). Eyes short, completely filling orbit, cornea large, black, round; peduncle short, smooth (
<figureCitation id="131C64BE79787E54FC366A8526DDFB52" box="[908,1004,1245,1269]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="10" pageNumber="616">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79787E54FC406A85210FFB52" box="[1018,1086,1245,1269]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">2A, C</figureCitation>
,
<figureCitation id="131C64BE79787E54FBF16A8521BEFB52" box="[1099,1167,1245,1269]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A, B</figureCitation>
).
</paragraph>
<paragraph id="8B98783B79787E54FC976B45209CFA52" blockId="10.[811,1453,1309,1525]" pageId="10" pageNumber="616">
Maxillipeds 3 closing almost completely with only narrow gap between ischia (
<figureCitation id="131C64BE79787E54FC6C6B652118FAF2" box="[982,1065,1341,1365]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">Fig. 2E</figureCitation>
); merus subquadrate, anteroexternal part distinctly auriculiform, wider than subrectangular ischium; ischium rectangular, with oblique submarginal sulcus. Exopod relatively stout, with subdistal triangular lobe on inner margin, distal edge reaching to distal edge of merus; flagellum long, exceeding width of merus (
<figureCitation id="131C64BE79787E54FAF06B852091FA52" box="[1354,1440,1501,1525]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="8B98783B79787E54FC97684520AEF8F2" blockId="10.[811,1452,1565,1877]" pageId="10" pageNumber="616">
Chelipeds heterochelous, relatively long, about 1.7× carapace width; ischiobasis short, smooth, with low protuberance on median part of flexor margin; merus short, quadrate in dorsal view, with sharp granule on subdistal edge of flexor margin, otherwise smooth; extensor margin smooth; carpus with strong tooth on inner angle; propodus outer surface smooth, without tubercles; fingers of major chela shorter than palm, cutting margins unarmed, tips recurved; fingers of minor chela as long as palm, with low teeth along distal third of cutting edges or almost smooth (
<figureCitation id="131C64BE79787E54FB28696521C2F8F2" box="[1170,1267,1853,1877]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="10" pageNumber="616">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79787E54FABB69652015F8F2" box="[1281,1316,1853,1877]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="10" pageNumber="616">2D</figureCitation>
,
<figureCitation id="131C64BE79787E54FA8B696520A3F8F2" box="[1329,1426,1853,1877]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A, E, F</figureCitation>
).
</paragraph>
<paragraph id="8B98783B79787E55FC95692527E0F809" blockId="10.[811,1453,1917,2101]" lastBlockId="11.[135,777,1782,1966]" lastPageId="11" lastPageNumber="617" pageId="10" pageNumber="616">
P25 short; P3 longest, P5 shortest; numerous and short stiff setae on surface and margins of carpus, propodus, dactylus but do not obscure margins, setae on merus more scattered, more concentrated on distal part; merus not elongated, relatively stout, not foliaceous, unarmed on dorsal margin, at most with low angle on subdistal part (
<figureCitation id="131C64BE79787E54FB7666452011F792" box="[1228,1312,2077,2101]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="10" pageNumber="616">Fig. 1A</figureCitation>
,
<figureCitation id="131C64BE79787E54FA9766452061F792" box="[1325,1360,2077,2101]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="10" pageNumber="616">3A</figureCitation>
); carpus short, smooth (
<figureCitation id="131C64BE79797E55FE8E68AE24BEF8A9" box="[308,399,1782,1806]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="11" pageNumber="617">Fig. 1A</figureCitation>
); propodus laterally compressed, dorsal margin uneven but not serrate, ventral margin with spines on distal half (
<figureCitation id="131C64BE79797E55FEC9696E24E2F8E9" box="[371,467,1846,1870]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="11" pageNumber="617">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79797E55FE5B696E2734F8E9" box="[481,517,1846,1870]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">3H</figureCitation>
); dactylo-propodal lock well-developed (
<figureCitation id="131C64BE79797E55FE86690E24A1F8C9" box="[316,400,1878,1902]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">Fig. 3H</figureCitation>
); dactylus margins with ungrouped, robust, movable spines, without comb-like rows of feeding setae, unguis corneous, gently curved (
<figureCitation id="131C64BE79797E55FD8869CE27A3F809" box="[562,658,1942,1966]" captionStart="Fig" captionStartId="2.[113,148,1231,1253]" captionTargetBox="[113,1451,189,1216]" captionTargetId="figure-67@2.[113,1451,189,1216]" captionTargetPageId="2" captionText="Fig. 1. Dorsal habitus. A, Ectaesthesius bifrons, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, Ectaesthesius bifrons, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands; C, Tetralia cinctipes, male (9.5 × 9.5 mm) (ZRC 2019.0701), Japan; D, Tetraloides heterodactylus, male (7.6 × 7.1 mm) (ZRC 2019.0699), Japan." figureDoi="http://doi.org/10.5281/zenodo.10271309" httpUri="https://zenodo.org/record/10271309/files/figure.png" pageId="11" pageNumber="617">Figs. 1A</figureCitation>
,
<figureCitation id="131C64BE79797E55FD1A69CE27F5F809" box="[672,708,1942,1966]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">3H</figureCitation>
).
</paragraph>
<caption id="DF5828B379797E55FF3268412757F918" ID-DOI="http://doi.org/10.5281/zenodo.10271327" ID-Zenodo-Dep="10271327" httpUri="https://zenodo.org/record/10271327/files/figure.png" pageId="11" pageNumber="617" startId="11.[136,171,1561,1583]" targetBox="[136,1474,189,1546]" targetPageId="11" targetType="figure">
<paragraph id="8B98783B79797E55FF3268412757F918" blockId="11.[136,1476,1561,1727]" pageId="11" pageNumber="617">
Fig. 7. Male anterior thoracic sternum and pleon. A,
<taxonomicName id="4C2703B879797E55FDCE6841262AF988" authorityName="Paulson" authorityYear="1875" box="[628,795,1561,1583]" class="Malacostraca" family="Tetraliidae" genus="Tetralia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="B953A42979797E55FDCE6841262AF988" box="[628,795,1561,1583]" italics="true" pageId="11" pageNumber="617">Tetralia cinctipes</emphasis>
</taxonomicName>
(9.5 × 9.5 mm) (ZRC 2019.0701), Japan; B,
<taxonomicName id="4C2703B879797E55FB78684120F0F989" baseAuthorityName="Heller" baseAuthorityYear="1861" box="[1218,1473,1561,1582]" class="Malacostraca" family="Tetraliidae" genus="Tetraloides" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="heterodactylus">
<emphasis id="B953A42979797E55FB78684120F0F989" box="[1218,1473,1561,1582]" italics="true" pageId="11" pageNumber="617">Tetraloides heterodactylus</emphasis>
</taxonomicName>
(7.6 × 7.1 mm) (ZRC 2019.0699), Japan; C,
<taxonomicName id="4C2703B879797E55FDFF686F2633F9EC" baseAuthorityName="Herbst" baseAuthorityYear="1801" box="[581,770,1590,1612]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="cymodoce">
<emphasis id="B953A42979797E55FDFF686F2633F9EC" box="[581,770,1590,1612]" italics="true" pageId="11" pageNumber="617">Trapezia cymodoce</emphasis>
</taxonomicName>
(11.6 × 9.9 mm) (ZRC 1999.2385), Singapore; D,
<taxonomicName id="4C2703B879797E55FB47686E20F5F9EC" authorityName="Alcock" authorityYear="1898" box="[1277,1476,1590,1611]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="maculosa">
<emphasis id="B953A42979797E55FB47686E20F5F9EC" box="[1277,1476,1590,1611]" italics="true" pageId="11" pageNumber="617">Quadrella maculosa</emphasis>
</taxonomicName>
(7.2 × 6.1 mm) (ZRC 2003.0352), Sulawesi, Indonesia; E,
<taxonomicName id="4C2703B879797E55FD69680B26D2F9CF" baseAuthorityName="Wood-Mason in Wood-Mason &amp; Alcock" baseAuthorityYear="1891" box="[723,995,1619,1640]" class="Malacostraca" family="Trapeziidae" genus="Sphenomerides" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="trapezioides">
<emphasis id="B953A42979797E55FD69680B26D2F9CF" box="[723,995,1619,1640]" italics="true" pageId="11" pageNumber="617">Sphenomerides trapezioides</emphasis>
</taxonomicName>
(7.8 × 5.9 mm) (ZRC 2000.2103), Madagascar; F,
<taxonomicName id="4C2703B879797E55FF1E682824BAF922" authorityName=", Calman" authorityYear="1909" box="[164,395,1648,1669]" class="Malacostraca" family="Trapeziidae" genus="Calocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="B953A42979797E55FF1E682824BAF922" box="[164,395,1648,1669]" italics="true" pageId="11" pageNumber="617">Calocarcinus africanus</emphasis>
</taxonomicName>
(9.3 × 7.2 mm) (ZRC 2008.1281), New Caledonia; G,
<taxonomicName id="4C2703B879797E55FC7A682821F4F922" authorityName="Garth &amp; Kim" authorityYear="1983" box="[960,1221,1648,1669]" class="Malacostraca" family="Trapeziidae" genus="Philippicarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="oviformis">
<emphasis id="B953A42979797E55FC7A682821F4F922" box="[960,1221,1648,1669]" italics="true" pageId="11" pageNumber="617">Philippicarcinus oviformis</emphasis>
</taxonomicName>
(16.0 × 11.3 mm) (ZRC 2009.0058), Philippines; H,
<taxonomicName id="4C2703B879797E55FE2F68D52749F906" baseAuthorityName="Desbonne in Desbonne &amp; Schramm" baseAuthorityYear="1867" box="[405,632,1676,1698]" class="Malacostraca" family="Domeciidae" genus="Domecia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="acanthophora">
<emphasis id="B953A42979797E55FE2F68D52749F906" box="[405,632,1676,1698]" italics="true" pageId="11" pageNumber="617">Domecia acanthophora</emphasis>
</taxonomicName>
(7.4 × 5.4 mm) (ZRC 2000.1551), Caribbean Sea, Panama; I,
<taxonomicName id="4C2703B879797E55FB7268D420F3F906" baseAuthorityName="Borradaile" baseAuthorityYear="1902" box="[1224,1474,1676,1697]" class="Malacostraca" family="Domeciidae" genus="Cherusius" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="617" phylum="Arthropoda" rank="species" species="triunguiculatus">
<emphasis id="B953A42979797E55FB7268D420F3F906" box="[1224,1474,1676,1697]" italics="true" pageId="11" pageNumber="617">Cherusius triunguiculatus</emphasis>
</taxonomicName>
(4.7 × 3.9 mm) (ZRC 2016.0121), Easter Island.
</paragraph>
</caption>
<paragraph id="8B98783B79797E55FF32698E204EF7A9" blockId="11.[136,775,2006,2062]" lastBlockId="11.[835,1476,1782,2062]" pageId="11" pageNumber="617">
Thoracic sternum smooth; sternites 1, 2 completely fused to form triangular structure without trace of fissure; sternites 2, 3 fused without trace of lateral fissures; sternites 3, 4 fused without trace of median suture, groove or depression except for short lateral fissure; sternopleonal cavity relatively shallow, poorly demarcated, reaching anteriorly to level of midlength of P2 coxae (
<figureCitation id="131C64BE79797E55FBFD692E21F1F829" box="[1095,1216,1910,1934]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="11" pageNumber="617">Figs. 2E, F</figureCitation>
,
<figureCitation id="131C64BE79797E55FB76692E21C1F829" box="[1228,1264,1910,1934]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">3G</figureCitation>
). Sternite 4 without longitudinal groove; sternites 58 complete with all sutures distinct (
<figureCitation id="131C64BE79797E55FC1269EE211AF869" box="[936,1067,1974,1998]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="11" pageNumber="617">Figs. 2E, F</figureCitation>
,
<figureCitation id="131C64BE79797E55FB8069EE216EF869" box="[1082,1119,1974,1998]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">3G</figureCitation>
). Rounded tubercle of pleonal-locking mechanism small, on posterior edge of thoracic sternite 5, near suture with sternite 6 (
<figureCitation id="131C64BE79797E55FB5F69AE2073F7A9" box="[1253,1346,2038,2062]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="11" pageNumber="617">Figs. 2F</figureCitation>
,
<figureCitation id="131C64BE79797E55FAF469AE2043F7A9" box="[1358,1394,2038,2062]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="11" pageNumber="617">3G</figureCitation>
).
</paragraph>
<paragraph id="8B98783B797E7E52FFCB6EE527BDFED2" blockId="12.[112,754,189,565]" pageId="12" pageNumber="618">
Pleon narrow; all somites and telson free, without trace of fusion; somite 1 trapezoidal, reaching to coxae of P5; somite 2 subrectangular; somite 3 widest, wider than somite 1; lateral margins distinctly convex; somites 4, 5 decreasing in width; somite 6 quadrate, wider than long, lateral margin gently sinuous; telson semicircular (
<figureCitation id="131C64BE797E7E52FE416F05276BFED2" box="[507,602,349,373]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="12" pageNumber="618">Figs. 2E</figureCitation>
,
<figureCitation id="131C64BE797E7E52FDDD6F0527B1FED2" box="[615,640,349,373]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="12" pageNumber="618">3I</figureCitation>
).
</paragraph>
<paragraph id="8B98783B797E7E52FFC96F252740FD92" blockId="12.[112,754,189,565]" pageId="12" pageNumber="618">
G1 arcuate, relatively slender, gently curving outwards, gradually tapering distally; margins of distal half lined with numerous short spinules; distal part dilated with wide opening, forming low dorsal flap (
<figureCitation id="131C64BE797E7E52FE646F852764FE52" box="[478,597,477,501]" captionStart="Fig" captionStartId="7.[136,171,1138,1160]" captionTargetBox="[138,1476,191,1128]" captionTargetId="figure-511@7.[111,1493,182,1137]" captionTargetPageId="7" captionText="Fig. 4. Ectaesthesius bifrons. A, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands. Left G1 (pleonal view); B, left G1 (sternal view); C, distal part of left G1 (pleonal view); D, distal part of left G1 (sternal view); E, left G2 (pleonal view). Scales: A, B, E = 0.5 mm; C, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10271317" httpUri="https://zenodo.org/record/10271317/files/figure.png" pageId="12" pageNumber="618">Fig. 4AD</figureCitation>
). G2 relatively slender, elongated, about four-fifths length of G1; distal portion prominent but short, filiform (
<figureCitation id="131C64BE797E7E52FDB56C452754FD92" box="[527,613,541,565]" captionStart="Fig" captionStartId="7.[136,171,1138,1160]" captionTargetBox="[138,1476,191,1128]" captionTargetId="figure-511@7.[111,1493,182,1137]" captionTargetPageId="7" captionText="Fig. 4. Ectaesthesius bifrons. A, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands. Left G1 (pleonal view); B, left G1 (sternal view); C, distal part of left G1 (pleonal view); D, distal part of left G1 (sternal view); E, left G2 (pleonal view). Scales: A, B, E = 0.5 mm; C, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10271317" httpUri="https://zenodo.org/record/10271317/files/figure.png" pageId="12" pageNumber="618">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="8B98783B797E7E52FFCB6C0527C0FCF2" blockId="12.[112,753,605,853]" pageId="12" pageNumber="618">
<emphasis id="B953A429797E7E52FFCB6C0525E7FDD2" bold="true" box="[113,214,605,629]" pageId="12" pageNumber="618">Females.</emphasis>
Female pleon longitudinally ovate, covering most of thoracic sternum but not along lateral parts; somite 1 trapezoidal, reaching to coxae of P5; all somites and telson free; telson triangular, wider than long, lateral margins gently convex (
<figureCitation id="131C64BE797E7E52FF756C852403FD52" box="[207,306,733,757]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="12" pageNumber="618">Figs. 2G</figureCitation>
,
<figureCitation id="131C64BE797E7E52FEFB6C85246DFD52" box="[321,348,733,757]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="12" pageNumber="618">3J</figureCitation>
). Sternopleonal cavity shallow (
<figureCitation id="131C64BE797E7E52FD796C852586FCB2" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="12" pageNumber="618">Fig. 2G, H</figureCitation>
). Vulva flat, subcircular, positioned on proximal part of sternite 6, adjacent to suture with sternite 5, positioned close to median line, covered by thin membrane (
<figureCitation id="131C64BE797E7E52FD376D6527D5FCF2" box="[653,740,829,853]" captionStart="Fig" captionStartId="4.[113,148,1997,2019]" captionTargetBox="[114,1452,228,1981]" captionTargetId="figure-14@4.[114,1452,228,1981]" captionTargetPageId="4" captionText="Fig. 2. Ectaesthesius bifrons. A, CF, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; B, G, H, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands: A, B, carapace, dorsal view; C, cephalothorax, anterior view; D, right cheliped (dorsal view); E, G, anterior thoracic sternum and pleon; F, male anterior thoracic sternum; H, sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.10271313" httpUri="https://zenodo.org/record/10271313/files/figure.png" pageId="12" pageNumber="618">Fig. 2H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D2BB0797E7E52FFCB6D2524CEFAF2" pageId="12" pageNumber="618" type="discussion">
<paragraph id="8B98783B797E7E52FFCB6D252724FBD2" blockId="12.[112,752,893,1141]" pageId="12" pageNumber="618">
<emphasis id="B953A429797E7E52FFCB6D2525D3FC32" bold="true" box="[113,226,893,917]" pageId="12" pageNumber="618">Remarks.</emphasis>
<taxonomicName id="4C2703B8797E7E52FF566D2524E9FC32" authorityName="Rathbun" authorityYear="1898" box="[236,472,893,917]" class="Malacostraca" family="Xanthidae" genus="Ectaesthesius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="618" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B953A429797E7E52FF566D2524E9FC32" box="[236,472,893,917]" italics="true" pageId="12" pageNumber="618">Ectaesthesius bifrons</emphasis>
</taxonomicName>
was described from one ovigerous female (9.7 × 7.0 mm; USNM 21586) from off Chatham Island, Galápagos Islands, from a depth of 45 fathoms (
<quantity id="4CDFD5DE797E7E52FF6F6D85243FFC52" box="[213,270,989,1013]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="12" pageNumber="618" unit="m" value="82.0">82 m</quantity>
).
<bibRefCitation id="EFB605CA797E7E52FE9A6D852495FC52" author="Garth JS" box="[288,420,989,1013]" pageId="12" pageNumber="618" pagination="341 - 522" refId="ref19810" refString="Garth JS (1946) Littoral brachyuran fauna of the Galapagos Islands. Allan Hancock Pacific Expeditions, 5 (10): 341 - 522, fig. 1, pls. 49 - 87." type="journal article" year="1946">Garth (1946</bibRefCitation>
: pl. 58) redescribed this species from a good series of specimens from the Galápagos Islands and figured a male and a female, but no indication was given as to which specimens was used, but they seem to match with those in LACM 1938.56 (
<figureCitation id="131C64BE797E7E52FE7F6A052739FBD2" box="[453,520,1117,1141]" captionStart="Fig" captionStartId="6.[113,148,1720,1742]" captionTargetBox="[117,1451,182,1704]" captionTargetId="figure-14@6.[105,1473,171,1715]" captionTargetPageId="6" captionText="Fig. 3. Ectaesthesius bifrons. AI, male (12.8 × 9.0 mm) (LACM 1938.56), Galápagos Islands; J, female (11.2 × 8.0 mm) (LACM 1938.56), Galápagos Islands. A, dorsal habitus; B, cephalothorax, anterior view; C, epistome posterior margin; D, left maxilliped 3; E, right chela, anterior view; F, left chela, anterior view; G, thoracic sternum; H, right P5 dactylus and propodus, dorsal view; I, J, pleon. Scales: A = 4.0 mm; B, DF, I, J = 2.0 mm; C, G, H = 0.5 mm. (A, B, DF, I, J after Garth, 1946: pl. 58)." figureDoi="http://doi.org/10.5281/zenodo.10271315" httpUri="https://zenodo.org/record/10271315/files/figure.png" pageId="12" pageNumber="618">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B98783B797E7E52FFCB6AC624CEFAF2" blockId="12.[113,753,1181,1365]" pageId="12" pageNumber="618">
Very little is known about the ecology of
<taxonomicName id="4C2703B8797E7E52FDF16AC62788FB12" authorityName="Rathbun" authorityYear="1898" box="[587,697,1181,1205]" class="Malacostraca" family="Xanthidae" genus="Ectaesthesius" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="618" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="B953A429797E7E52FDF16AC62788FB12" box="[587,697,1181,1205]" italics="true" pageId="12" pageNumber="618">E. bifrons</emphasis>
</taxonomicName>
, and it is somewhat surprising that it has not been recollected since the
<emphasis id="B953A429797E7E52FF646A852414FB52" box="[222,293,1245,1269]" italics="true" pageId="12" pageNumber="618">Velero</emphasis>
cruises in the late 1930s, even though it is not a deep-water species. It appears to be endemic to the Galápagos Islands, being known only from three of the islands: Chatham, James, and Hood.
</paragraph>
</subSubSection>
</treatment>
</document>