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<document id="019CFA61B917CC728E455BDFDFEDE4C4" ID-DOI="10.11646/zootaxa.5384.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10376440" ID-ZooBank="0211978A-E124-4D9F-9A92-B565AA7B7891" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702549873658" checkinUser="plazi" docAuthor="Yu, Kun, Maddison, Wayne P. &amp; Zhang, Junxia" docDate="2023" docId="038CA47F72772060C6B57DBF9230FE97" docLanguage="en" docName="zootaxa.5384.1.1.pdf" docOrigin="Zootaxa 5384 (1)" docSource="https://mapress.com/zt/article/download/zootaxa.5384.1.1/52479" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Orcevia zabkai Yu, Maddison &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="19" masterDocId="FFB5DC0772662072C6227F449060FFD6" masterDocTitle="Taxonomic revision of Orcevia Thorell, 1890, with description of fifteen new species (Araneae, Salticidae, Euophryini)" masterLastPageNumber="79" masterPageNumber="1" pageNumber="18" updateTime="1702550864234" updateUser="ExternalLinkService">
<mods:mods id="312AB8C0CA564342C44BB46C5C54106E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6FE9D9994CA136AD3400D7237A16B39">
<mods:title id="E3B115767806B5B2E1528092842FCA1C">Taxonomic revision of Orcevia Thorell, 1890, with description of fifteen new species (Araneae, Salticidae, Euophryini)</mods:title>
</mods:titleInfo>
<mods:name id="0FB547CA27BFECEA99B1B5A1EA181C98" type="personal">
<mods:role id="C689A03B7C51F7944666B5E06BF90325">
<mods:roleTerm id="B8EB48FDE994F427E280C3842F5781CC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E83BCCB742E6516D37B1CB4F9F74962A">Yu, Kun</mods:namePart>
<mods:nameIdentifier id="93B64FC9E17A08EBB93CD2A80DC43261" type="ORCID">0000-0002-5517-4484</mods:nameIdentifier>
<mods:affiliation id="94704002202082697636E7DB754F13AB">Key Laboratory of Zoological Systematics and Application of Hebei Province, Hebei Basic Science Center for Biotic Interaction, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China. &amp; 13468690210 @ 163. com; https: // orcid. org / 0000 - 0002 - 5517 - 4484</mods:affiliation>
<mods:nameIdentifier id="D15770FE861908EBED7E655E90BC4881" type="email">13468690210@163.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A89E5ACAE8A7AABFC5E9CC5F04D3FFA7" type="personal">
<mods:role id="A8469785416837766D8915E9FB9B667B">
<mods:roleTerm id="B3C0CC21F5B8CFE26B0E1D5D2E82F252">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C0CADF6AD1A02571BCA4B22308216DE1">Maddison, Wayne P.</mods:namePart>
<mods:nameIdentifier id="AA0BC209070F1789F7D6F1B9388C88D6" type="ORCID">0000-0003-4953-4575</mods:nameIdentifier>
<mods:affiliation id="32111120185ED6ACFE42C693D1427537">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada &amp; wayne. maddison @ ubc. ca; https: // orcid. org / 0000 - 0003 - 4953 - 4575</mods:affiliation>
<mods:nameIdentifier id="35AA8CB1B37116930347CB6CC030532D" type="email">wayne.maddison@ubc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="38CEB1F828DC21884EE35DF94B57C938" type="personal">
<mods:role id="ED7FC96258E5D0ADA2645F2901022C11">
<mods:roleTerm id="105E5ECAA65A85F163C1FA4F6AB006A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAE36A25BD8345E2BC98CFF6B30BA7FE">Zhang, Junxia</mods:namePart>
<mods:nameIdentifier id="976665D2C3A6F8AEAA3F0C36599299CC" type="ORCID">0000-0003-2179-3954</mods:nameIdentifier>
<mods:affiliation id="F7702F1ED1F2AAEA946543BB00DC8845">Key Laboratory of Zoological Systematics and Application of Hebei Province, Hebei Basic Science Center for Biotic Interaction, College of Life Sciences, Hebei University, Baoding, Hebei 071002, P. R. China. &amp; jxzhang 1976 @ 163. com; https: // orcid. org / 0000 - 0003 - 2179 - 3954 Corresponding author</mods:affiliation>
<mods:nameIdentifier id="A904D4C874C46002601C6DE85E90305A" type="email">jxzhang1976@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0BCF66934688A95F1D5FF5C156775F39">text</mods:typeOfResource>
<mods:relatedItem id="537A7020A3C4BEA75D68A687544EEB19" type="host">
<mods:titleInfo id="988308B0E40644B6F85E2C280A28D26B">
<mods:title id="555D488679B24A14893EFAF7D3A55C06">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F1BE83FBD902411B2EFDB552203527FB">
<mods:date id="8A4FE38F0CACE91C3246911B712E9E0B">2023</mods:date>
<mods:detail id="F739CF7B6CF32916075CD23CAA7F0C00" type="pubDate">
<mods:number id="30342C980426D7E7C3F42A8EAF2D1356">2023-12-14</mods:number>
</mods:detail>
<mods:detail id="91129796C9D94F088920B2D784FCBB02" type="volume">
<mods:number id="2DF4A2DE04426E3091903BCE84E19737">5384</mods:number>
</mods:detail>
<mods:detail id="5AA69A9ECD96677C8CC3E9437020FE83" type="issue">
<mods:number id="C4BA79C9F15744E8B00064DD9C3E309B">1</mods:number>
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<mods:start id="54B9A0685C9C381D17589052A7009215">1</mods:start>
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<mods:location id="1E78E525AAED0245852C3DE16C2DF482">
<mods:url id="F920850F1C4FE1A2782CF1455030ABE5">https://mapress.com/zt/article/download/zootaxa.5384.1.1/52479</mods:url>
</mods:location>
<mods:classification id="299DAAACC6C36DCDD8A4BD586EF31939">journal article</mods:classification>
<mods:identifier id="F5EF44CED1A8884AD20ECC4B556B6427" type="DOI">10.11646/zootaxa.5384.1.1</mods:identifier>
<mods:identifier id="BA3FE3026E435A0AE15C5F0C4FEE80D6" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="0BDC8D07E05611AD57E31817D69F0237" type="Zenodo-Dep">10376440</mods:identifier>
<mods:identifier id="FDBF45D76E781073D9FBDEBD49D22391" type="ZooBank">0211978A-E124-4D9F-9A92-B565AA7B7891</mods:identifier>
</mods:mods>
<treatment id="038CA47F72772060C6B57DBF9230FE97" ID-DOI="http://doi.org/10.5281/zenodo.10376344" ID-Zenodo-Dep="10376344" LSID="urn:lsid:plazi:treatment:038CA47F72772060C6B57DBF9230FE97" httpUri="http://treatment.plazi.org/id/038CA47F72772060C6B57DBF9230FE97" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="C33F46E272772063C6B57DBF93F3FCC2" box="[151,915,762,789]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8B9A156972772063C6B57DBF93F3FCC2" blockId="17.[151,915,762,825]" box="[151,915,762,789]" pageId="17" pageNumber="18">
<heading id="D0D2A20572772063C6B57DBF93F3FCC2" bold="true" box="[151,915,762,789]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="4">
<emphasis id="B951C97B72772063C6B57DBF93F3FCC2" bold="true" box="[151,915,762,789]" pageId="17" pageNumber="18">
<taxonomicName id="4C256EEA72772063C6B57DBF921BFCC3" authority="Yu, Maddison &amp; Zhang" authorityName="Yu, Maddison &amp; Zhang" authorityYear="2023" box="[151,635,762,789]" class="Arachnida" family="Salticidae" genus="Orcevia" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="zabkai" status="sp. nov.">
<emphasis id="B951C97B72772063C6B57DBF912BFCC3" bold="true" box="[151,331,762,789]" italics="true" pageId="17" pageNumber="18">Orcevia zabkai</emphasis>
Yu, Maddison &amp; Zhang
</taxonomicName>
,
<taxonomicNameLabel id="A262740072772063C4AA7DBF9285FCC3" box="[648,741,763,789]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
(Ẹ氏ĸ尔¤)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272772063C6B57C5B912DFCEF" box="[151,333,798,825]" pageId="17" pageNumber="18" type="description">
<paragraph id="8B9A156972772063C6B57C5B912DFCEF" blockId="17.[151,915,762,825]" box="[151,333,798,825]" pageId="17" pageNumber="18">
<figureCitation id="131E09EC72772063C6B57C5B90BEFCEF" box="[151,222,799,825]" captionStart="FIGURES 1–12" captionStartId="19.[151,264,1823,1847]" captionTargetBox="[151,1436,181,1803]" captionTargetId="figure-17@19.[151,1436,181,1803]" captionTargetPageId="19" captionText="FIGURES 1–12. Comparison of male palpal bulbs of some euophryines, in ventral view. 1–8. Orcevia spp. [O. bokoblin Yu, Maddison &amp; Zhang sp. nov. (1), O. nietzschei Yu, Maddison &amp; Zhang sp. nov. (2), O. jinping Yu &amp; Zhang, sp. nov. (3), O. mercuryi Yu, Maddison &amp; Zhang, sp. nov. (4), O. shuyuani Yu &amp; Zhang, sp. nov. (5), O. zabkai Yu, Maddison &amp; Zhang, sp. nov. (6), O. yahaha Yu, Maddison &amp; Zhang, sp. nov. (7) and O. wuliang Yu &amp; Zhang, sp. nov. (8)]; 9. Laufeia concava Zhang &amp; Maddison, 2012; 10. Amoenema erhai Yu &amp; Zhang, 2023; 11. Lokina nyuewa Yu &amp; Zhang, 2023; 12. Chalcovietnamicus vietnamensis (Żabka, 1985). Black dotted lines indicate retrolateral edge of membrane between salticid radix and tegulum, white arrows point to the retrolateral extension, and yellow lines indicate the proximal margin of tegulum." figureDoi="http://doi.org/10.5281/zenodo.10376442" httpUri="https://zenodo.org/record/10376442/files/figure.png" pageId="17" pageNumber="18">Figs 6</figureCitation>
,
<figureCitation id="131E09EC72772063C6C87C5A912DFCEF" box="[234,333,798,825]" captionStart-0="FIGURES 332–337" captionStart-1="FIGURES 338–345" captionStart-2="FIGURES 346–352" captionStartId-0="72.[151,264,1521,1545]" captionStartId-1="73.[151,264,1845,1869]" captionStartId-2="74.[151,264,1835,1859]" captionTargetBox-0="[317,1271,181,1497]" captionTargetBox-1="[185,1403,151,1837]" captionTargetBox-2="[208,1384,191,1808]" captionTargetId-0="figure-22@72.[317,1271,181,1497]" captionTargetId-1="figure-17@73.[185,1403,151,1837]" captionTargetId-2="figure-22@74.[204,1384,181,1811]" captionTargetPageId-0="72" captionTargetPageId-1="73" captionTargetPageId-2="74" captionText-0="FIGURES 332–337. Living spiders of Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., male (332–334) and female (335–337). Figures are copyrights W.P. Maddison." captionText-1="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." captionText-2="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi-0="http://doi.org/10.5281/zenodo.10376562" figureDoi-1="http://doi.org/10.5281/zenodo.10376564" figureDoi-2="http://doi.org/10.5281/zenodo.10376566" httpUri-0="https://zenodo.org/record/10376562/files/figure.png" httpUri-1="https://zenodo.org/record/10376564/files/figure.png" httpUri-2="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">332–352</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272772063C6B57C239561FBC7" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B9A156972772063C6B57C239161FC3B" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
<emphasis id="B951C97B72772063C6B57C239126FC56" bold="true" box="[151,326,870,897]" pageId="17" pageNumber="18">Type material.</emphasis>
<materialsCitation id="3B4D1F3472772063C76C7C2393B2FC73" collectingDate="2019-06-01" collectingDateMax="2019-06-02" collectingDateMin="2019-06-01" collectionCode="UBCZ" collectorName="W. P. Maddison &amp; Paul Ng" country="Malaysia" elevation="10" latitude="1.336" location="Adam Road" longLatPrecision="78" longitude="103.816" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B951C97B72772063C76C7C2391A4FC57" bold="true" box="[334,452,871,897]" pageId="17" pageNumber="18">
<typeStatus id="549EABCB72772063C76C7C2391DFFC57" box="[334,447,871,897]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D23DEE072772063C7EE7C239181FC57" box="[460,481,871,897]" count="1" pageId="17" pageNumber="18" type="male">♂</specimenCount>
(
<collectionCode id="ED348DAC72772063C7D37C239221FC57" box="[497,577,871,897]" country="Canada" name="University of British Columbia, Spencer Museum" pageId="17" pageNumber="18">UBCZ</collectionCode>
; KYU-SAL502),
<collectingCountry id="F33255F972772063C5297C2393C1FC57" box="[779,929,871,897]" name="Malaysia" pageId="17" pageNumber="18">MALAYSIA</collectingCountry>
: Sarawak,
<location id="8EFA43B272772063C2387C2394C4FC57" LSID="urn:lsid:plazi:treatment:038CA47F72772060C6B57DBF9230FE97:8EFA43B272772063C2387C2394C4FC57" box="[1050,1188,871,897]" country="Malaysia" latitude="1.336" longLatPrecision="78" longitude="103.816" name="Adam Road" pageId="17" pageNumber="18">Adam Road</location>
,
<geoCoordinate id="EE1173AE72772063C2927C239572FC57" box="[1200,1298,871,897]" degrees="1.336" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="55" value="1.336">1.336°N</geoCoordinate>
,
<geoCoordinate id="EE1173AE72772063C33C7C2395F8FC57" box="[1310,1432,871,897]" degrees="103.816" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="55" value="103.816">103.816°E</geoCoordinate>
,
<quantity id="4CDDB88C72772063C6B57CCF90B0FC73" box="[151,208,907,933]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="10.0">
<elevation id="0008F25A72772063C6B57CCF90B0FC73" box="[151,208,907,933]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="10.0">10 m</elevation>
</quantity>
elev.,
<date id="FF9B33A972772063C73A7CCF91DEFC73" box="[280,446,907,933]" pageId="17" pageNumber="18" value="2019-06-01" valueMax="2019-06-02" valueMin="2019-06-01">
<collectingDate id="EFDFCA4172772063C73A7CCF91DEFC73" box="[280,446,907,933]" pageId="17" pageNumber="18" value="2019-06-01" valueMax="2019-06-02" valueMin="2019-06-01">1–2 June 2019</collectingDate>
</date>
, leg.
<collectorName id="26D070BF72772063C7DB7CCF92C2FC73" box="[505,674,907,933]" pageId="17" pageNumber="18">W.P. Maddison</collectorName>
&amp;
<collectorName id="26D070BF72772063C4E47CCF9343FC73" box="[710,803,907,933]" pageId="17" pageNumber="18">Paul Ng</collectorName>
, WPM#19-030
</materialsCitation>
.
<materialsCitation id="3B4D1F3472772063C5FE7CCF9254FC1F" collectingDate="2019-06-01" collectingDateMax="2019-06-02" collectingDateMin="2019-06-01" collectionCode="UBCZ" collectorName="W. P. Maddison &amp; Paul Ng" country="American Samoa" elevation="10" latitude="1.336" location="Adam Road" longLatPrecision="78" longitude="103.816" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B951C97B72772063C5FE7CCF943DFC73" bold="true" box="[988,1117,907,933]" pageId="17" pageNumber="18">
<typeStatus id="549EABCB72772063C5FE7CCF9438FC73" box="[988,1112,907,933]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D23DEE072772063C2467CCF94E7FC73" box="[1124,1159,907,933]" count="1" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
(
<collectionCode id="ED348DAC72772063C2B47CCF9486FC73" box="[1174,1254,907,933]" country="Canada" name="University of British Columbia, Spencer Museum" pageId="17" pageNumber="18">UBCZ</collectionCode>
; KYU-SAL503, #
<collectingCountry id="F33255F972772063C6847CEB90AAFC1F" box="[166,202,943,969]" name="American Samoa" pageId="17" pageNumber="18">AS</collectingCountry>
19.0072), same data as holotype
</materialsCitation>
;
<materialsCitation id="3B4D1F3472772063C41D7CEB909DFC3B" collectingDate="2019-06-01" collectingDateMax="2019-06-02" collectingDateMin="2019-06-01" collectionCode="UBCZ" collectorName="W. P. Maddison &amp; Paul Ng" country="American Samoa" elevation="10" latitude="1.336" location="Adam Road" longLatPrecision="78" longitude="103.816" pageId="17" pageNumber="18" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D23DEE072772063C41D7CEB9202FC1F" box="[575,610,943,969]" count="1" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
<specimenCount id="9D23DEE072772063C4407CEB92E5FC1F" box="[610,645,943,969]" count="2" pageId="17" pageNumber="18" type="female">2♀</specimenCount>
(
<collectionCode id="ED348DAC72772063C4B17CEB9283FC1F" box="[659,739,943,969]" country="Canada" name="University of British Columbia, Spencer Museum" pageId="17" pageNumber="18">UBCZ</collectionCode>
; KYU-SAL511; KYU-SAL513; KYU-SAL522), same data as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8B9A156972772063C6E57CB29561FBC7" blockId="17.[151,1437,870,2013]" box="[199,1281,1014,1041]" pageId="17" pageNumber="18">
<materialsCitation id="3B4D1F3472772063C6E57CB2949EFBC7" box="[199,1278,1014,1041]" collectingDate="2019-06-01" collectingDateMax="2019-06-02" collectingDateMin="2019-06-01" collectionCode="UBCZ" collectorName="W. P. Maddison &amp; Paul Ng" country="Malaysia" elevation="10" latitude="1.336" location="Adam Road" longLatPrecision="78" longitude="103.816" pageId="17" pageNumber="18" specimenCount="1" specimenCount-juvenile="1">
<emphasis id="B951C97B72772063C6E57CB2919CFBC6" bold="true" box="[199,508,1014,1040]" pageId="17" pageNumber="18">
<typeStatus id="549EABCB72772063C6E57CB29196FBC6" box="[199,502,1014,1040]" isOtherTypeStatus="true" pageId="17" pageNumber="18">Other material examined</typeStatus>
.
</emphasis>
<specimenCount id="9D23DEE072772063C4207CB39211FBC7" box="[514,625,1015,1041]" count="1" pageId="17" pageNumber="18" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED348DAC72772063C4A27CB392B0FBC7" box="[640,720,1015,1041]" country="Canada" name="University of British Columbia, Spencer Museum" pageId="17" pageNumber="18">UBCZ</collectionCode>
; DNA-Voucher KYU098), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272772063C6E57B5F93BAFB8F" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8B9A156972772063C6E57B5F93BAFB8F" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
<emphasis id="B951C97B72772063C6E57B5F912EFBE3" bold="true" box="[199,334,1051,1077]" pageId="17" pageNumber="18">Etymology.</emphasis>
The specific epithet is in honor of Dr. Marek
<taxonomicName id="4C256EEA72772063C57F7B5893C3FBE1" authorityName="Wang, Li &amp; Pham" authorityYear="2023" box="[861,931,1052,1079]" class="Arachnida" family="Salticidae" genus="Zabka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="21" phylum="Arthropoda" rank="genus">Żabka</taxonomicName>
(Siedlce University of Natural Sciences and Humanities), who has contributed greatly to the jumping spider taxonomy.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272772063C6E57B2694FDFADB" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="8B9A156972772063C6E57B2694FDFADB" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
<emphasis id="B951C97B72772063C6E57B269122FBAA" bold="true" box="[199,322,1122,1148]" pageId="17" pageNumber="18">Diagnosis.</emphasis>
Males can be distinguished from
<taxonomicName id="4C256EEA72772063C4947B279496FBAB" authority="(Proszynski &amp; Deeleman-Reinhold, 2013)" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" baseAuthorityName="Proszynski &amp; Deeleman-Reinhold" baseAuthorityYear="2013" box="[694,1270,1123,1149]" class="Arachnida" family="Salticidae" genus="Echeclus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sokoli" status="comb. nov.">
<emphasis id="B951C97B72772063C4947B27937AFBAB" box="[694,794,1123,1149]" italics="true" pageId="17" pageNumber="18">O. sokoli</emphasis>
(
<bibRefCitation id="EFB4689872772063C5057B27948EFBAB" author="Proszynski, J. &amp; Deeleman-Reinhold, C. L." box="[807,1262,1123,1149]" pageId="17" pageNumber="18" pagination="113 - 144" refId="ref26880" refString="Proszynski, J. &amp; Deeleman-Reinhold, C. L. (2013) Description of some Salticidae (Araneae) from the Malay Archipelago. III. Salticidae of Borneo, with comments on adjacent territories. Arthropoda Selecta, 22 (2), 113 - 144. https: // doi. org / 10.15298 / arthsel. 22.2.02" type="journal article" year="2013">Prószyński &amp; Deeleman-Reinhold, 2013</bibRefCitation>
)
</taxonomicName>
<emphasis id="B951C97B72772063C2D97B27951AFBAB" bold="true" box="[1275,1402,1123,1149]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="A262740072772063C2D97B27951AFBAB" box="[1275,1402,1123,1149]" pageId="17" pageNumber="18" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
by the unbranched RTA (
<figureCitation id="131E09EC72772063C7AE7BC3918EFB77" box="[396,494,1159,1185]" captionStart="FIGURES 346–352" captionStartId="74.[151,264,1835,1859]" captionTargetBox="[208,1384,191,1808]" captionTargetId="figure-22@74.[204,1384,181,1811]" captionTargetPageId="74" captionText="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.10376566" httpUri="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">Fig. 347</figureCitation>
; vs. tip of RTA is bicuspid in
<taxonomicName id="4C256EEA72772063C5187BC393FFFB77" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" baseAuthorityName="Proszynski &amp; Deeleman-Reinhold" baseAuthorityYear="2013" box="[826,927,1159,1185]" class="Arachnida" family="Salticidae" genus="Echeclus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sokoli">
<emphasis id="B951C97B72772063C5187BC393FFFB77" box="[826,927,1159,1185]" italics="true" pageId="17" pageNumber="18">O. sokoli</emphasis>
</taxonomicName>
) and the slender embolus (
<figureCitation id="131E09EC72772063C2EC7BC39551FB77" box="[1230,1329,1159,1185]" captionStart="FIGURES 346–352" captionStartId="74.[151,264,1835,1859]" captionTargetBox="[208,1384,191,1808]" captionTargetId="figure-22@74.[204,1384,181,1811]" captionTargetPageId="74" captionText="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.10376566" httpUri="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">Fig. 348</figureCitation>
); females can be distinguished from
<taxonomicName id="4C256EEA72772063C79E7BEF9240FB13" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" baseAuthorityName="Proszynski &amp; Deeleman-Reinhold" baseAuthorityYear="2013" box="[444,544,1195,1221]" class="Arachnida" family="Salticidae" genus="Echeclus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sokoli">
<emphasis id="B951C97B72772063C79E7BEF9240FB13" box="[444,544,1195,1221]" italics="true" pageId="17" pageNumber="18">O. sokoli</emphasis>
</taxonomicName>
by the starting part of copulatory duct that is fused to spermatheca, the relatively short accessory glands, with the ratio of length of AG to diameter of spermatheca as 1:3 ~ 1:2 (
<figureCitation id="131E09EC72772063C2E47B8B9548FB3F" box="[1222,1320,1230,1257]" captionStart="FIGURES 346–352" captionStartId="74.[151,264,1835,1859]" captionTargetBox="[208,1384,191,1808]" captionTargetId="figure-22@74.[204,1384,181,1811]" captionTargetPageId="74" captionText="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.10376566" httpUri="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">Fig. 352</figureCitation>
; vs. AG is obviously long, with the ratio of AG length to diameter of spermatheca ca. 3:
<quantity id="4CDDB88C72772063C5D17BB79441FADB" box="[1011,1057,1267,1293]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="17" pageNumber="18" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C256EEA72772063C2057BB794EDFADB" authorityName="Yu &amp; Maddison &amp; Zhang" authorityYear="2023" baseAuthorityName="Proszynski &amp; Deeleman-Reinhold" baseAuthorityYear="2013" box="[1063,1165,1267,1293]" class="Arachnida" family="Salticidae" genus="Echeclus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sokoli">
<emphasis id="B951C97B72772063C2057BB794EDFADB" box="[1063,1165,1267,1293]" italics="true" pageId="17" pageNumber="18">O. sokoli</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272772060C6E57A5292DAFECB" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8B9A156972772063C6E57A529424F9FB" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
<emphasis id="B951C97B72772063C6E57A529139FAE6" bold="true" box="[199,345,1302,1328]" pageId="17" pageNumber="18">Description.</emphasis>
Male. Habitus of
<typeStatus id="549EABCB72772063C4027A5392E2FAE7" box="[544,642,1303,1329]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
as shown in
<figureCitation id="131E09EC72772063C5327A53930FFAE7" box="[784,879,1303,1329]" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="17" pageNumber="18">Fig. 338</figureCitation>
. Measurements of
<typeStatus id="549EABCB72772063C2627A5394C6FAE7" box="[1088,1190,1303,1329]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
: carapace length 3.23; abdomen length 2.87; measurements of eyes: AME 0.57, ALE 0.36, PME 0.08, PLE 0.29; measurements of legs: I 7.27 (2.12, 1.35, 1.80, 1.27, 0.73), II 5.05 (1.57, 0.92, 0.98, 0.99, 0.59), III 4.98 (1.61, 0.77, 0.96, 1.03, 0.61), IV 5.66 (1.73, 0.79, 1.31, 1.24, 0.59); leg formula 1423. Carapace dark, with rather dense pale setae overhead and almost no setae at flanks and posterior part; with some yellowish-brown setae between anterior lens; dorsal abdomen dark, with dense pale setae anteriorly and gradually thinning to rear, three oblique pale setal bands present on last two thirds of dorsal abdomen each side. Legs I and femora II–IV dark, other segments lighter-colored; pale setae present from distal femora to tibia dorsally, and a little on distal two thirds of metatarsi I–II.
</paragraph>
<paragraph id="8B9A156972772063C6E579739507F96B" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
Palp (
<figureCitation id="131E09EC72772063C72B797391CAF987" box="[265,426,1591,1617]" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="17" pageNumber="18">Figs 340–342</figureCitation>
): palpal bulb with obvious retrolateral constriction between tegular lobe and main part of tegulum; embolus with many small grooves on basal part along direction of embolic extension, one row of saw-like denticles present on median part of embolus; tip of embolus cylindric (
<figureCitation id="131E09EC72772063C5B9793B939AF94F" box="[923,1018,1662,1689]" captionStart="FIGURES 346–352" captionStartId="74.[151,264,1835,1859]" captionTargetBox="[208,1384,191,1808]" captionTargetId="figure-22@74.[204,1384,181,1811]" captionTargetPageId="74" captionText="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.10376566" httpUri="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">Fig. 351</figureCitation>
); RTA basally wide, almost aequilatus in basal half, then gradually narrowed to tip; one small bump present on ventral side of base of RTA (
<figureCitation id="131E09EC72772063C2DB79E7953BF96B" box="[1273,1371,1699,1725]" captionStart="FIGURES 346–352" captionStartId="74.[151,264,1835,1859]" captionTargetBox="[208,1384,191,1808]" captionTargetId="figure-22@74.[204,1384,181,1811]" captionTargetPageId="74" captionText="FIGURES 346–352. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., 346. Male palp; 347. RTA; 348–349. Expanded palpal bulb; 350. Embolus; 351. Tip of embolus; 352. Vulva; in ventral (346, 350–351), retrolateral (347), prolateral (348) and dorsal (349, 352) view. Abbreviations:AG, accessory gland; DH, distal haematodocha; E, embolus; ED, embolic disc; FD, fertilization duct; LPT, lobe-like protrusion of tegulum; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; TL, tegular lobe; ST, subtegulum." figureDoi="http://doi.org/10.5281/zenodo.10376566" httpUri="https://zenodo.org/record/10376566/files/figure.png" pageId="17" pageNumber="18">Fig. 346</figureCitation>
).
</paragraph>
<paragraph id="8B9A156972772063C6E57983913BF843" blockId="17.[151,1437,870,2013]" pageId="17" pageNumber="18">
Female. Habitus as shown in
<figureCitation id="131E09EC72772063C40A798392ECF937" box="[552,652,1735,1761]" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="17" pageNumber="18">Fig. 339</figureCitation>
. Measurements of
<typeStatus id="549EABCB72772063C54C798393AEF937" box="[878,974,1735,1761]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
(KYU-SAL513): carapace length 3.07; abdomen length 3.18; measurements of eyes: AME 0.55, ALE 0.36, PME 0.08, PLE 0.29; measurements of legs: I 5.23 (1.57, 1.10, 1.15, 0.80, 0.61), II 4.68 (1.55, 0.95, 0.93, 0.76, 0.49), III 4.92 (1.60, 0.79, 1.00, 0.96, 0.57), IV 5.85 (1.79, 0.86, 1.34, 1.18, 0.68); leg formula 4132. Distribution of pale setae and body color like male, but pale setae relatively sparse, especially on cephalon; pale setae reduced on face; patellae and tibiae III–IV relatively darker than male.
</paragraph>
<paragraph id="8B9A156972772060C6E578DB9267FF2F" blockId="17.[151,1437,870,2013]" lastBlockId="18.[151,1437,151,321]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
Epigynum (
<figureCitation id="131E09EC72772063C77378DB9193F86F" box="[337,499,1951,1977]" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="17" pageNumber="18">Fig. 343–344</figureCitation>
): atria subcircular, close to each other; outer edges of atria stretching upward, forming one shallow and indistinct concave area above atria. Vulva (
<figureCitation id="131E09EC72772063C5B778879398F80B" box="[917,1016,1986,2013]" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="17" pageNumber="18">Fig. 345</figureCitation>
): copulatory ducts long and relatively wide, downwards extended first, then turning upwards and forming a loop at ventral side of spermathecae (
<figureCitation id="131E09EC72742060C34D7FD390A5FF03" captionStart="FIGURES 338–345" captionStartId="73.[151,264,1845,1869]" captionTargetBox="[185,1403,151,1837]" captionTargetId="figure-17@73.[185,1403,151,1837]" captionTargetPageId="73" captionText="FIGURES 338–345. Orcevia zabkai Yu, Maddison &amp; Zhang, sp. nov., holotype male (338), paratype male (340–342) and female (339, 343–345). 338–339. Habitus; 340–342. Palp, with arrows indicating the row of serrula on embolus, and the small bump near base of RTA; 343–344. Epigynum; 345. Vulva; in dorsal (338–339, 345), prolateral (341), ventral (340, 343–344) and retrolateral (342) view. Abbreviations: AE, apophysis of embolic disc; AG, accessory gland; CD, copulatory duct; E, embolus; ED, embolic disc; LPT, lobe-like protrusion of tegulum; pH, pro-ventral hump of male palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, salticid radix; T, tegulum; TL, tegular lobe; VFB, ventral femoral bump of male palp." figureDoi="http://doi.org/10.5281/zenodo.10376564" httpUri="https://zenodo.org/record/10376564/files/figure.png" pageId="18" pageNumber="19">Fig. 344</figureCitation>
); spermathecae slightly darker than copulatory ducts; accessory glands pointing upwards, partly covered by fertilization ducts in dorsal view.
</paragraph>
<paragraph id="8B9A156972742060C6E57E4792DAFECB" blockId="18.[151,1437,151,321]" box="[199,698,258,285]" pageId="18" pageNumber="19">
<emphasis id="B951C97B72742060C6E57E4791E5FECA" bold="true" box="[199,389,258,285]" pageId="18" pageNumber="19">Natural history.</emphasis>
Tree trunk dwelling spider.
</paragraph>
</subSubSection>
<subSubSection id="C33F46E272742060C6E57E629230FE97" box="[199,592,294,321]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8B9A156972742060C6E57E629230FE97" blockId="18.[151,1437,151,321]" box="[199,592,294,321]" pageId="18" pageNumber="19">
<emphasis id="B951C97B72742060C6E57E629101FE96" bold="true" box="[199,353,294,320]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingCountry id="F33255F972742060C7457E6391AEFE97" box="[359,462,295,321]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
(
<collectingCountry id="F33255F972742060C7FF7E639224FE97" box="[477,580,295,321]" name="Malaysia" pageId="18" pageNumber="19">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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