treatments-xml/data/03/8C/34/038C3420FF82FFF8AEC9FECF0D01FBAC.xml

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<mods:title id="07D6CF61309DF6C8479D7FF4544EB56A">Marine crabs new to Singapore, with a description of a new species of intertidal xanthid crab of the genus Macromedaeus Ward, 1942 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:date id="3C9CABDEAC8B7BA14CA4A6153D7462FC">2021</mods:date>
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<heading id="D0D2325AFF82FFFEAEC9FECF0C39FECC" bold="true" box="[928,1382,317,341]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="465" reason="2">
<taxonomicName id="4C25FEB5FF82FFFEAEC9FECF0C39FECC" ID-CoL="3KW2C" authority="(Zarenkov, 1971)" authorityName="Serene" authorityYear="1984" baseAuthorityName="Zarenkov" baseAuthorityYear="1971" box="[928,1382,317,341]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAEC9FECF0C39FECC" bold="true" box="[928,1382,317,341]" pageId="2" pageNumber="465">
<emphasis id="B9515924FF82FFFEAEC9FECF0DC8FECC" bold="true" box="[928,1175,317,341]" italics="true" pageId="2" pageNumber="465">Hepatoporus guinotae</emphasis>
(
<bibRefCitation id="EFB4F8C7FF82FFFEA9CEFECF0C01FECC" author="Zarenkov NA" box="[1191,1374,317,341]" pageId="2" pageNumber="465" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov, 1971</bibRefCitation>
)
</emphasis>
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</paragraph>
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<paragraph id="8B9A8536FF82FFFEA92FFEAF0DE0FEEC" blockId="2.[928,1382,317,373]" box="[1094,1215,349,373]" pageId="2" pageNumber="465">
(
<figureCitation id="131E99B3FF82FFFEA927FEAF0DC4FEEC" box="[1102,1179,349,373]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="2" pageNumber="465">Figs. 2</figureCitation>
, 
<figureCitation id="131E99B3FF82FFFEA9C0FEAF0DE9FEEC" box="[1193,1206,349,373]" captionStart="Fig" captionStartId="4.[136,171,1727,1749]" captionTargetBox="[138,774,191,1712]" captionTargetId="figure-8@4.[133,777,186,1738]" captionTargetPageId="4" captionText="Fig. 3. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, in situ photograph showing live colouration (photographed by Toh Chay Hoon); B, left G1, pleonal view; C, distal tip of left G1, pleonal view; D, distal tip of left G1, sternal view. Scale bars: B = 0.5 mm; C, D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7174591" httpUri="https://zenodo.org/record/7174591/files/figure.png" pageId="2" pageNumber="465">3</figureCitation>
)
</paragraph>
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<subSubSection id="C33FD6BDFF82FFFEAE2AFE6E0CE0FDE2" pageId="2" pageNumber="465" type="reference_group">
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<treatmentCitation id="0A84A327FF82FFFEAE2AFE6E0DB7FE28" author="Zarenkov NA" box="[835,1256,411,433]" page="191" pageId="2" pageNumber="465" year="1971">
<taxonomicName id="4C25FEB5FF82FFFEAE2AFE6E0DB7FE28" ID-CoL="RGJM" authority="Zarenkov, 1971: 191" authorityName="Zarenkov" authorityPageNumber="191" authorityYear="1971" box="[835,1256,411,433]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAE2AFE6E0D4CFE28" box="[835,1043,412,433]" italics="true" pageId="2" pageNumber="465">Carpoporus guinotae</emphasis>
<materialsCitation id="3B4D8F6BFF82FFFEA973FE690DB7FE28" box="[1050,1256,411,433]" location="Red Sea" pageId="2" pageNumber="465" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<bibRefCitation id="EFB4F8C7FF82FFFEA973FE690DB7FE28" author="Zarenkov NA" box="[1050,1256,411,433]" pageId="2" pageNumber="465" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov, 1971: 191</bibRefCitation>
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, fig. 86 (
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locality: 
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).
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</paragraph>
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<treatmentCitationGroup id="AB35A218FF82FFFEAE2AFE240D67FDC4" pageId="2" pageNumber="465">
<treatmentCitation id="0A84A327FF82FFFEAE2AFE240D95FE72" author="Serene R" box="[835,1226,469,491]" page="75" pageId="2" pageNumber="465" year="1984">
<taxonomicName id="4C25FEB5FF82FFFEAE2AFE240D95FE72" ID-CoL="3KW2C" authority="Serene, 1984: 75" authorityName="Serene" authorityPageNumber="75" authorityYear="1984" baseAuthorityName="Zarenkov" baseAuthorityYear="1971" box="[835,1226,469,491]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAE2AFE240D43FE72" box="[835,1052,470,491]" italics="true" pageId="2" pageNumber="465">Hepatoporus guinotae</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEA94AFE270D95FE72" author="Serene R" box="[1059,1226,469,491]" pageId="2" pageNumber="465" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984: 75</bibRefCitation>
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, fig. 40, pl. 10 figs. D–F. — 
<treatmentCitation id="0A84A327FF82FFFEAEE0FE000DD3FD9E" author="Davie PJF &amp; Turner PA" box="[905,1164,498,520]" page="83" pageId="2" pageNumber="465" year="1994">
<bibRefCitation id="EFB4F8C7FF82FFFEAEE0FE000DD3FD9E" author="Davie PJF &amp; Turner PA" box="[905,1164,498,520]" pageId="2" pageNumber="465" pagination="83 - 86" refId="ref10712" refString="Davie PJF &amp; Turner PA (1994) A new species and a new record of Hepatoporus from north-western Australia (Crustacea: Decapoda: Xanthidae). Memoirs of the Queensland Museum, 37 (1): 83 - 86." type="journal article" year="1994">Davie &amp; Turner, 1994: 83</bibRefCitation>
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, fig. 1A, B. — 
<treatmentCitation id="0A84A327FF82FFFEA841FE000AD1FDBD" author="Ng PKL &amp; Guinot D &amp; Davie PJF" page="199" pageId="2" pageNumber="465" year="2008">
<bibRefCitation id="EFB4F8C7FF82FFFEA841FE000AD1FDBD" author="Ng PKL &amp; Guinot D &amp; Davie PJF" pageId="2" pageNumber="465" pagination="1 - 286" refId="ref12359" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 199</bibRefCitation>
</treatmentCitation>
(list). — 
<treatmentCitation id="0A84A327FF82FFFEAE91FDFD0D92FDBC" author="Naderloo R" box="[1016,1229,527,549]" page="252" pageId="2" pageNumber="465" year="2017">
<bibRefCitation id="EFB4F8C7FF82FFFEAE91FDFD0D92FDBC" author="Naderloo R" box="[1016,1229,527,549]" pageId="2" pageNumber="465" refId="ref12278" refString="Naderloo R (2017) Atlas of the Crabs of the Persian Gulf. Springer International Publishing, Cham, Switzerland, xxii + 444 pp." type="book" year="2017">Naderloo, 2017: 252</bibRefCitation>
</treatmentCitation>
, figs. 21.20d, 21.25. — 
<treatmentCitation id="0A84A327FF82FFFEAE00FDD90DC4FDD8" author="Takeda M &amp; Komatsu H" box="[873,1179,555,577]" page="161" pageId="2" pageNumber="465" year="2018">
<bibRefCitation id="EFB4F8C7FF82FFFEAE00FDD90DC4FDD8" author="Takeda M &amp; Komatsu H" box="[873,1179,555,577]" pageId="2" pageNumber="465" pagination="153 - 189" refId="ref13265" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018: 161</bibRefCitation>
</treatmentCitation>
, figs. 1D, 2B. — 
<treatmentCitation id="0A84A327FF82FFFEA825FDDE0A9DFDC7" author="Maenosono T" page="25" pageId="2" pageNumber="465" year="2021">
<bibRefCitation id="EFB4F8C7FF82FFFEA825FDDE0A9DFDC7" author="Maenosono T" pageId="2" pageNumber="465" pagination="21 - 33" refId="ref11999" refString="Maenosono T (2021) Notes on some euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) collected from the Ryukyu Islands, including a new record from Japan. Cancer, 30: 21 - 33." type="journal article" year="2021">Maenosono, 2021: 25</bibRefCitation>
</treatmentCitation>
, figs. 1E, 3.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9A8536FF82FFFEAE2AFD970CE0FDE2" blockId="2.[835,1475,411,635]" box="[835,1471,613,635]" pageId="2" pageNumber="465">
<treatmentCitationGroup id="AB35A218FF82FFFEAE2AFD970CE0FDE2" box="[835,1471,613,635]" pageId="2" pageNumber="465">
<treatmentCitation id="0A84A327FF82FFFEAE2AFD970C0AFDE2" author="Takeda M &amp; Nagai S" box="[835,1365,613,635]" page="549" pageId="2" pageNumber="465" year="1986">
<taxonomicName id="4C25FEB5FF82FFFEAE2AFD970C0AFDE2" ID-CoL="RGJL" authority="Takeda &amp; Nagai, 1986: 549" authorityName="Takeda &amp; Nagai" authorityPageNumber="549" authorityYear="1986" baseAuthorityName="Takeda &amp; Nagai" baseAuthorityYear="1986" box="[835,1365,613,635]" class="Malacostraca" family="Xanthidae" genus="Carpoporoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B9515924FF82FFFEAE2AFD970D67FDE3" box="[835,1080,613,634]" italics="true" pageId="2" pageNumber="465">Carpoporoides distinctus</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEA929FD970C0AFDE2" author="Takeda M &amp; Nagai S" box="[1088,1365,613,635]" pageId="2" pageNumber="465" pagination="547 - 550" refId="ref13315" refString="Takeda M &amp; Nagai S (1986) Establishment of a new genus on two Japanese crabs. Zoological Science, 3: 547 - 550." type="journal article" year="1986">Takeda &amp; Nagai, 1986: 549</bibRefCitation>
</taxonomicName>
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, fig. 1a, b.
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</paragraph>
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<emphasis id="B9515924FF82FFFEAE2AFD510D41FD22" bold="true" box="[835,1054,675,699]" pageId="2" pageNumber="465">Material examined.</emphasis>
<materialsCitation id="3B4D8F6BFF82FFFEA94BFD510DFCFD62" collectingDate="2019-03-16" collectionCode="ZRC" collectorName="C. H. Toh" country="Singapore" elevation="10" location="Pulau Hantu" pageId="2" pageNumber="465" specimenCode="ZRC 2019.0663" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D234EBFFF82FFFEA94BFD510D34FD22" box="[1058,1131,675,699]" pageId="2" pageNumber="465" type="male">1 male</specimenCount>
, 8.5 × 
<quantity id="4CDD28D3FF82FFFEA9C4FD510DA3FD22" box="[1197,1276,675,699]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="465" unit="mm" value="6.6">6.6 mm</quantity>
(
<specimenCode id="DB832D4DFF82FFFEA860FD510CE8FD22" box="[1289,1463,675,699]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="465">ZRC 2019.0663</specimenCode>
), 
<collectingCountry id="F332C5A6FF82FFFEAE2AFD310AEDFD42" box="[835,946,707,731]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
, 
<location id="8EFAD3EDFF82FFFEAED6FD310D15FD42" LSID="urn:lsid:plazi:treatment:038C3420FF82FFF8AEC9FECF0D01FBAC:8EFAD3EDFF82FFFEAED6FD310D15FD42" box="[959,1098,707,731]" country="Singapore" name="Pulau Hantu" pageId="2" pageNumber="465">Pulau Hantu</location>
, seabed, rubble, night dive, 
<quantity id="4CDD28D3FF82FFFEA8EBFD310CE2FD42" box="[1410,1469,707,731]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="465" unit="m" value="10.0">
<elevation id="00086205FF82FFFEA8EBFD310CE2FD42" box="[1410,1469,707,731]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="465" unit="m" value="10.0">10 m</elevation>
</quantity>
, coll. 
<collectorName id="26D0E0E0FF82FFFEAE12FD110AB3FD62" box="[891,1004,739,763]" pageId="2" pageNumber="465">C. H. Toh</collectorName>
, 
<date id="FF9BA3F6FF82FFFEAE90FD110DFCFD62" box="[1017,1187,739,763]" pageId="2" pageNumber="465" value="2019-03-16">
<collectingDate id="EFDF5A1EFF82FFFEAE90FD110DFCFD62" box="[1017,1187,739,763]" pageId="2" pageNumber="465" value="2019-03-16">16 March 2019</collectingDate>
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.
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<paragraph id="8B9A8536FF82FFFEAE2AFCD10C65FA62" blockId="2.[834,1475,803,1531]" pageId="2" pageNumber="465">
<emphasis id="B9515924FF82FFFEAE2AFCD10AEDFCA2" bold="true" box="[835,946,803,827]" pageId="2" pageNumber="465">Remarks.</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEAED1FCD10D34FCA2" author="Zarenkov NA" box="[952,1131,803,827]" pageId="2" pageNumber="465" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov (1971)</bibRefCitation>
described 
<taxonomicName id="4C25FEB5FF82FFFEA9B7FCD60C9DFCA2" authorityName="Zarenkov" authorityYear="1971" box="[1246,1474,804,827]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA9B7FCD60C9DFCA2" box="[1246,1474,804,827]" italics="true" pageId="2" pageNumber="465">Carpoporus guinotae</emphasis>
</taxonomicName>
from the Red Sea. 
<bibRefCitation id="EFB4F8C7FF82FFFEA95BFCB10D82FCC2" author="Serene R" box="[1074,1245,835,859]" pageId="2" pageNumber="465" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène (1984)</bibRefCitation>
erected the genus 
<taxonomicName id="4C25FEB5FF82FFFEAE2CFC960A8FFCE2" box="[837,976,868,891]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="B9515924FF82FFFEAE2CFC960A8FFCE2" box="[837,976,868,891]" italics="true" pageId="2" pageNumber="465">Hepatoporus</emphasis>
</taxonomicName>
to separate the Indo-West Pacific species (
<taxonomicName id="4C25FEB5FF82FFFEA8C3FC960D6AFC02" authority="Sakai, 1935" authorityName="Sakai" authorityYear="1935" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B9515924FF82FFFEA8C3FC960AF6FC02" italics="true" pageId="2" pageNumber="465">C. orientalis</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEAEDBFC710D6AFC02" author="Sakai T" box="[946,1077,899,923]" pageId="2" pageNumber="465" pagination="36 - 86" refId="ref12707" refString="Sakai T (1935) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the zoological survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 36 - 86." type="journal article" year="1935">Sakai, 1935</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C25FEB5FF82FFFEA919FC760DB1FC02" authorityName="Zarenkov" authorityYear="1971" box="[1136,1262,900,923]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA919FC760DB1FC02" box="[1136,1262,900,923]" italics="true" pageId="2" pageNumber="465">C. guinotae</emphasis>
</taxonomicName>
) from the Atlantic, and 
<typeStatus id="549E3B94FF82FFFEAE1AFC560AFBFC25" box="[883,932,932,956]" pageId="2" pageNumber="465">type</typeStatus>
, species, 
<taxonomicName id="4C25FEB5FF82FFFEA966FC560C0BFC22" authority="Stimpson, 1871" authorityName="Stimpson" authorityYear="1871" box="[1039,1364,931,955]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="papulosus">
<emphasis id="B9515924FF82FFFEA966FC560DC2FC22" box="[1039,1181,931,955]" italics="true" pageId="2" pageNumber="465">C. papulosus</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEA9CEFC510C0BFC22" author="Stimpson W" box="[1191,1364,931,955]" pageId="2" pageNumber="465" pagination="92 - 136" refId="ref13153" refString="Stimpson W (1871) Notes on North American Crustacea in the Museum of the Smithsonian Institution. No. III. Annals of the Lyceum of Natural History of New York, 10: 92 - 136." type="journal article" year="1871">Stimpson, 1871</bibRefCitation>
</taxonomicName>
; and with 
<taxonomicName id="4C25FEB5FF82FFFEAE2AFC360A92FC42" authorityName="Sakai" authorityYear="1935" box="[835,973,963,987]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B9515924FF82FFFEAE2AFC360A92FC42" box="[835,973,963,987]" italics="true" pageId="2" pageNumber="465">C. orientalis</emphasis>
</taxonomicName>
designated as 
<typeStatus id="549E3B94FF82FFFEA91CFC360DFCFC45" box="[1141,1187,964,988]" pageId="2" pageNumber="465">type</typeStatus>
species for 
<taxonomicName id="4C25FEB5FF82FFFEA846FC360CE4FC42" box="[1327,1467,964,987]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="genus">
<emphasis id="B9515924FF82FFFEA846FC360CE4FC42" box="[1327,1467,964,987]" italics="true" pageId="2" pageNumber="465">Hepatoporus</emphasis>
</taxonomicName>
. Subsequently, other species were described from 
<collectingCountry id="F332C5A6FF82FFFEA832FC160CC7FC62" box="[1371,1432,996,1019]" name="Japan" pageId="2" pageNumber="465">Japan</collectingCountry>
(
<taxonomicName id="4C25FEB5FF82FFFEA8C1FC160DEFFB82" authority="(Takeda &amp; Nagai, 1986))" authorityName="Takeda &amp; Nagai" authorityYear="1986" baseAuthorityName="Takeda &amp; Nagai" baseAuthorityYear="1986" class="Malacostraca" family="Xanthidae" genus="Carpoporoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B9515924FF82FFFEA8C1FC160AF8FB82" italics="true" pageId="2" pageNumber="465">H. distinctus</emphasis>
(
<bibRefCitation id="EFB4F8C7FF82FFFEAEDBFBF10DFFFB82" author="Takeda M &amp; Nagai S" box="[946,1184,1027,1051]" pageId="2" pageNumber="465" pagination="547 - 550" refId="ref13315" refString="Takeda M &amp; Nagai S (1986) Establishment of a new genus on two Japanese crabs. Zoological Science, 3: 547 - 550." type="journal article" year="1986">Takeda &amp; Nagai, 1986</bibRefCitation>
))
</taxonomicName>
, 
<collectingCountry id="F332C5A6FF82FFFEA9D0FBF10C47FB82" box="[1209,1304,1027,1051]" name="Australia" pageId="2" pageNumber="465">Australia</collectingCountry>
(
<taxonomicName id="4C25FEB5FF82FFFEA84FFBF60AB4FBA2" authority="Davie &amp; Turner, 1994" authorityName="Davie &amp; Turner" authorityYear="1994" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B9515924FF82FFFEA84FFBF60C20FB82" box="[1318,1407,1028,1051]" italics="true" pageId="2" pageNumber="465">H. asper</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEA8EDFBF10AB4FBA2" author="Davie PJF &amp; Turner PA" pageId="2" pageNumber="465" pagination="83 - 86" refId="ref10712" refString="Davie PJF &amp; Turner PA (1994) A new species and a new record of Hepatoporus from north-western Australia (Crustacea: Decapoda: Xanthidae). Memoirs of the Queensland Museum, 37 (1): 83 - 86." type="journal article" year="1994">Davie &amp; Turner, 1994</bibRefCitation>
</taxonomicName>
), and the 
<collectingCountry id="F332C5A6FF82FFFEA939FBD10D97FBA2" box="[1104,1224,1059,1083]" name="Philippines" pageId="2" pageNumber="465">Philippines</collectingCountry>
(
<taxonomicName id="4C25FEB5FF82FFFEA9B1FBD60AF5FBC2" authority="Mendoza &amp; Ng, 2008" authorityName="Mendoza &amp; Ng" authorityYear="2008" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="pumex">
<emphasis id="B9515924FF82FFFEA9B1FBD60C61FBA2" box="[1240,1342,1060,1083]" italics="true" pageId="2" pageNumber="465">H. pumex</emphasis>
<bibRefCitation id="EFB4F8C7FF82FFFEA82DFBD10AF5FBC2" author="Mendoza JCE &amp; Ng PKL" pageId="2" pageNumber="465" pagination="385 - 404" refId="ref12093" refString="Mendoza JCE &amp; Ng PKL (2008) New genera and species of euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, the Philippines. Raffles Bulletin of Zoology, 56 (2): 385 - 404." type="journal article" year="2008">Mendoza &amp; Ng, 2008</bibRefCitation>
</taxonomicName>
). 
<bibRefCitation id="EFB4F8C7FF82FFFEAED2FBB10DEFFBC2" author="Davie PJF &amp; Turner PA" box="[955,1200,1091,1115]" pageId="2" pageNumber="465" pagination="83 - 86" refId="ref10712" refString="Davie PJF &amp; Turner PA (1994) A new species and a new record of Hepatoporus from north-western Australia (Crustacea: Decapoda: Xanthidae). Memoirs of the Queensland Museum, 37 (1): 83 - 86." type="journal article" year="1994">Davie &amp; Turner (1994)</bibRefCitation>
also reported 
<taxonomicName id="4C25FEB5FF82FFFEA82CFBB60C9DFBC2" authorityName="Zarenkov" authorityYear="1971" box="[1349,1474,1092,1115]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA82CFBB60C9DFBC2" box="[1349,1474,1092,1115]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
from northwestern 
<collectingCountry id="F332C5A6FF82FFFEA97DFB910D26FBE2" box="[1044,1145,1123,1147]" name="Australia" pageId="2" pageNumber="465">Australia</collectingCountry>
, and further suggested that 
<taxonomicName id="4C25FEB5FF82FFFEA8C1FB960AF7FB02" authorityName="Takeda &amp; Nagai" authorityYear="1986" baseAuthorityName="Takeda &amp; Nagai" baseAuthorityYear="1986" class="Malacostraca" family="Xanthidae" genus="Carpoporoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B9515924FF82FFFEA8C1FB960AF7FB02" italics="true" pageId="2" pageNumber="465">H. distinctus</emphasis>
</taxonomicName>
is possibly synonymous with 
<taxonomicName id="4C25FEB5FF82FFFEA9B4FB760C07FB02" authorityName="Zarenkov" authorityYear="1971" box="[1245,1368,1156,1179]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA9B4FB760C07FB02" box="[1245,1368,1156,1179]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
due to the lack of morphological distinctions beyond the shape of their subhepatic cavities. Subsequent reports of 
<taxonomicName id="4C25FEB5FF82FFFEA863FB360CD8FB42" authorityName="Zarenkov" authorityYear="1971" box="[1290,1415,1220,1243]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA863FB360CD8FB42" box="[1290,1415,1220,1243]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
from 
<collectingCountry id="F332C5A6FF82FFFEAE2AFB160A20FB62" box="[835,895,1252,1275]" name="Japan" pageId="2" pageNumber="465">Japan</collectingCountry>
(i.e., Ogasawara Islands and 
<collectingRegion id="49E14BD4FF82FFFEA9C3FB110C56FB62" box="[1194,1289,1251,1275]" country="Japan" name="Okinawa" pageId="2" pageNumber="465">Okinawa</collectingRegion>
Island) supported this synonymisation (
<bibRefCitation id="EFB4F8C7FF82FFFEA94AFAF10C69FA82" author="Takeda M &amp; Komatsu H" box="[1059,1334,1283,1307]" pageId="2" pageNumber="465" pagination="153 - 189" refId="ref13265" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
; 
<bibRefCitation id="EFB4F8C7FF82FFFEA856FAF60A23FAA2" author="Maenosono T" pageId="2" pageNumber="465" pagination="21 - 33" refId="ref11999" refString="Maenosono T (2021) Notes on some euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) collected from the Ryukyu Islands, including a new record from Japan. Cancer, 30: 21 - 33." type="journal article" year="2021">Maenosono, 2021</bibRefCitation>
), suggesting that the shape of the subhepatic cavity of 
<taxonomicName id="4C25FEB5FF82FFFEAE2AFAB60A9FFAC2" authorityName="Zarenkov" authorityYear="1971" box="[835,960,1348,1371]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAE2AFAB60A9FFAC2" box="[835,960,1348,1371]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
undergoes size-related variation to some degree. The validity of this synonymisation will have to be evaluated rigorously when all material reported thus far, especially the 
<typeStatus id="549E3B94FF82FFFEAE2AFA560A24FA25" box="[835,891,1444,1468]" pageId="2" pageNumber="465">types</typeStatus>
of 
<taxonomicName id="4C25FEB5FF82FFFEAECDFA560DD0FA22" authority="Zarenkov" authorityName="Zarenkov" authorityYear="1971" box="[932,1167,1443,1467]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAECDFA560D7DFA22" box="[932,1058,1444,1467]" italics="true" pageId="2" pageNumber="465">C. guinotae</emphasis>
Zarenkov
</taxonomicName>
and 
<taxonomicName id="4C25FEB5FF82FFFEA9A0FA560ADAFA42" authority="Takeda &amp; Nagai" authorityName="Takeda &amp; Nagai" authorityYear="1986" baseAuthorityName="Takeda &amp; Nagai" baseAuthorityYear="1986" class="Malacostraca" family="Xanthidae" genus="Carpoporoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B9515924FF82FFFEA9A0FA560C10FA22" box="[1225,1359,1443,1467]" italics="true" pageId="2" pageNumber="465">C. distinctus</emphasis>
Takeda &amp; Nagai
</taxonomicName>
, as well as additional material from the entire range of 
<taxonomicName id="4C25FEB5FF82FFFEAE0BFA160ABDFA62" authorityName="Zarenkov" authorityYear="1971" box="[866,994,1508,1531]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEAE0BFA160ABDFA62" box="[866,994,1508,1531]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
, can be examined side by side.
</paragraph>
<paragraph id="8B9A8536FF82FFF8AE2AF9D10AB8FDEC" blockId="2.[835,1475,1571,2107]" lastBlockId="4.[835,1475,189,629]" lastPageId="4" lastPageNumber="467" pageId="2" pageNumber="465">
The present specimen from 
<collectingCountry id="F332C5A6FF82FFFEA917F9D10DB4F9A2" box="[1150,1259,1571,1595]" name="Singapore" pageId="2" pageNumber="465">Singapore</collectingCountry>
(
<figureCitation id="131E99B3FF82FFFEA994F9D10C12F9A2" box="[1277,1357,1571,1595]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="2" pageNumber="465">Figs. 2</figureCitation>
, 
<figureCitation id="131E99B3FF82FFFEA835F9D10C36F9A2" box="[1372,1385,1571,1595]" captionStart="Fig" captionStartId="4.[136,171,1727,1749]" captionTargetBox="[138,774,191,1712]" captionTargetId="figure-8@4.[133,777,186,1738]" captionTargetPageId="4" captionText="Fig. 3. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, in situ photograph showing live colouration (photographed by Toh Chay Hoon); B, left G1, pleonal view; C, distal tip of left G1, pleonal view; D, distal tip of left G1, sternal view. Scale bars: B = 0.5 mm; C, D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7174591" httpUri="https://zenodo.org/record/7174591/files/figure.png" pageId="2" pageNumber="465">3</figureCitation>
) agrees well with the description and illustrations of 
<taxonomicName id="4C25FEB5FF82FFFEA856F9B60C9DF9C2" authorityName="Zarenkov" authorityYear="1971" box="[1343,1474,1604,1627]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF82FFFEA856F9B60C9DF9C2" box="[1343,1474,1604,1627]" italics="true" pageId="2" pageNumber="465">H. guinotae</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFB4F8C7FF82FFFEAE06F9910D44F9E2" author="Zarenkov NA" box="[879,1051,1635,1659]" pageId="2" pageNumber="465" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov, 1971</bibRefCitation>
: fig. 86; 
<bibRefCitation id="EFB4F8C7FF82FFFEA910F9910C57F9E2" author="Serene R" box="[1145,1288,1635,1659]" pageId="2" pageNumber="465" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
: pl. 10 figs. D–F; 
<bibRefCitation id="EFB4F8C7FF82FFFEAE2AF9710AACF902" author="Naderloo R" box="[835,1011,1667,1691]" pageId="2" pageNumber="465" refId="ref12278" refString="Naderloo R (2017) Atlas of the Crabs of the Persian Gulf. Springer International Publishing, Cham, Switzerland, xxii + 444 pp." type="book" year="2017">Naderloo, 2017</bibRefCitation>
: figs. 21.20d, 21.25; 
<bibRefCitation id="EFB4F8C7FF82FFFEA988F9710A21F922" author="Takeda M &amp; Komatsu H" pageId="2" pageNumber="465" pagination="153 - 189" refId="ref13265" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: figs. 1D, 2B; 
<bibRefCitation id="EFB4F8C7FF82FFFEA942F9560DA7F922" author="Maenosono T" box="[1067,1272,1699,1723]" pageId="2" pageNumber="465" pagination="21 - 33" refId="ref11999" refString="Maenosono T (2021) Notes on some euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) collected from the Ryukyu Islands, including a new record from Japan. Cancer, 30: 21 - 33." type="journal article" year="2021">Maenosono, 2021</bibRefCitation>
: figs. 1E, 3). The reticulate pattern of connected granules and small cavities on the dorsal surface of the carapace is prominent and widespread in the present specimen (
<figureCitation id="131E99B3FF82FFFEA98CF8F10C1FF882" box="[1253,1344,1795,1819]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="2" pageNumber="465">Fig. 2A</figureCitation>
), similar to that seen in a much larger (CW 17.0 mm) specimen from 
<collectingCountry id="F332C5A6FF82FFFEAE2AF8B60AD6F8C5" box="[835,905,1860,1884]" name="Kenya" pageId="2" pageNumber="465">Kenya</collectingCountry>
(cf. 
<bibRefCitation id="EFB4F8C7FF82FFFEAEA9F8B10D0DF8C2" author="Serene R" box="[960,1106,1859,1883]" pageId="2" pageNumber="465" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
: pl. 10 fig. D), whereas similarly sized or smaller specimens tend to have less prominent pits, giving the dorsal carapace a smoother appearance (
<bibRefCitation id="EFB4F8C7FF82FFFEAE22F8510AA7F822" author="Naderloo R" box="[843,1016,1955,1979]" pageId="2" pageNumber="465" refId="ref12278" refString="Naderloo R (2017) Atlas of the Crabs of the Persian Gulf. Springer International Publishing, Cham, Switzerland, xxii + 444 pp." type="book" year="2017">Naderloo, 2017</bibRefCitation>
: fig. 21.25; 
<bibRefCitation id="EFB4F8C7FF82FFFEA910F8510CCBF822" author="Takeda M &amp; Komatsu H" box="[1145,1428,1955,1979]" pageId="2" pageNumber="465" pagination="153 - 189" refId="ref13265" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: fig. 2B). Similarly, this reticulate pattern can also be seen in the external surfaces of the chelae (
<figureCitation id="131E99B3FF82FFFEA9FFF8110DB9F862" box="[1174,1254,2019,2043]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="2" pageNumber="465">Fig. 2E</figureCitation>
) and on the thoracic sternum and pleon (
<figureCitation id="131E99B3FF82FFFEA948F7F10DC1F782" box="[1057,1182,2051,2075]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="2" pageNumber="465">Fig. 2G, H</figureCitation>
). The dorsal border of the subhepatic cavity slopes continuously towards the posterior (
<bibRefCitation id="EFB4F8C7FF84FFF8AE22FF4F0A80FF4C" author="Serene R" box="[843,991,189,213]" pageId="4" pageNumber="467" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
: pl. 10 fig. D). The subhepatic cavity itself (
<figureCitation id="131E99B3FF84FFF8AE22FF2F0AE2FF6C" box="[843,957,221,245]" captionStart="Fig" captionStartId="3.[113,148,2006,2028]" captionTargetBox="[127,1437,189,1991]" captionTargetId="figure-7@3.[127,1437,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="4" pageNumber="467">Fig. 2C, D</figureCitation>
) has a well-developed dorsal roof, which is seen in most specimens of 
<taxonomicName id="4C25FEB5FF84FFF8A951FF0C0DE7FE8C" authorityName="Zarenkov" authorityYear="1971" box="[1080,1208,254,277]" class="Malacostraca" family="Xanthidae" genus="Carpoporus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF84FFF8A951FF0C0DE7FE8C" box="[1080,1208,254,277]" italics="true" pageId="4" pageNumber="467">H. guinotae</emphasis>
</taxonomicName>
except for the 
<typeStatus id="549E3B94FF84FFF8A80CFF0F0C9DFE8C" box="[1381,1474,253,277]" pageId="4" pageNumber="467" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C25FEB5FF84FFF8AE09FEEC0AB8FEAC" authorityName="Takeda &amp; Nagai" authorityYear="1986" baseAuthorityName="Takeda &amp; Nagai" baseAuthorityYear="1986" box="[864,999,285,309]" class="Malacostraca" family="Xanthidae" genus="Carpoporoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="B9515924FF84FFF8AE09FEEC0AB8FEAC" box="[864,999,285,309]" italics="true" pageId="4" pageNumber="467">H. distinctus</emphasis>
</taxonomicName>
, where the dorsal roof is mostly effaced (cf. 
<bibRefCitation id="EFB4F8C7FF84FFF8AE2AFECF0D19FECC" author="Takeda M &amp; Nagai S" box="[835,1094,317,341]" pageId="4" pageNumber="467" pagination="547 - 550" refId="ref13315" refString="Takeda M &amp; Nagai S (1986) Establishment of a new genus on two Japanese crabs. Zoological Science, 3: 547 - 550." type="journal article" year="1986">Takeda &amp; Nagai, 1986</bibRefCitation>
: fig. 1a). The presence of capped, mushroom-like (fungiform) spines on the ventral floor of the subhepatic cavities appears to be a consistent feature for this species; in the 
<collectingCountry id="F332C5A6FF84FFF8A92CFE6F0DEEFE2C" box="[1093,1201,413,437]" name="Singapore" pageId="4" pageNumber="467">Singapore</collectingCountry>
specimen, there are two such spines in each cavity (cf. 
<bibRefCitation id="EFB4F8C7FF84FFF8A9F2FE4F0C06FE4C" author="Serene R" box="[1179,1369,445,469]" pageId="4" pageNumber="467" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984: 75</bibRefCitation>
). The G1 of the present specimen (
<figureCitation id="131E99B3FF84FFF8A93FFE2F0D95FE6C" box="[1110,1226,477,501]" captionStart="Fig" captionStartId="4.[136,171,1727,1749]" captionTargetBox="[138,774,191,1712]" captionTargetId="figure-8@4.[133,777,186,1738]" captionTargetPageId="4" captionText="Fig. 3. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, in situ photograph showing live colouration (photographed by Toh Chay Hoon); B, left G1, pleonal view; C, distal tip of left G1, pleonal view; D, distal tip of left G1, sternal view. Scale bars: B = 0.5 mm; C, D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7174591" httpUri="https://zenodo.org/record/7174591/files/figure.png" pageId="4" pageNumber="467">Fig. 3B–D</figureCitation>
) closely resembles the illustration by 
<bibRefCitation id="EFB4F8C7FF84FFF8AE81FE0F0D20FD8C" author="Serene R" box="[1000,1151,509,533]" pageId="4" pageNumber="467" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène (1984</bibRefCitation>
: fig. 40) for a slightly larger male from 
<collectingCountry id="F332C5A6FF84FFF8AEA9FDEF0D1DFDAC" box="[960,1090,541,565]" name="Madagascar" pageId="4" pageNumber="467">Madagascar</collectingCountry>
in the lobulation of the distal tip, although the former is relatively more slender and has fewer subdistal setae.
</paragraph>
<caption id="DF5AD5BEFF83FFFFAD18F8240A1CF7DB" ID-DOI="http://doi.org/10.5281/zenodo.7174589" ID-Zenodo-Dep="7174589" httpUri="https://zenodo.org/record/7174589/files/figure.png" pageId="3" pageNumber="466" startId="3.[113,148,2006,2028]" targetBox="[127,1437,189,1991]" targetPageId="3">
<paragraph id="8B9A8536FF83FFFFAD18F8240A1CF7DB" blockId="3.[113,1452,2006,2115]" pageId="3" pageNumber="466">
Fig. 2. 
<taxonomicName id="4C25FEB5FF83FFFFADD1F8250B1DF872" authority="(Zarenkov, 1971)" authorityName="Serene" authorityYear="1984" baseAuthorityName="Zarenkov" baseAuthorityYear="1971" box="[184,578,2006,2028]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF83FFFFADD1F82508CFF875" box="[184,400,2007,2028]" italics="true" pageId="3" pageNumber="466">Hepatoporus guinotae</emphasis>
(
<bibRefCitation id="EFB4F8C7FF83FFFFACF7F8240B63F872" author="Zarenkov NA" box="[414,572,2006,2028]" pageId="3" pageNumber="466" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov, 1971</bibRefCitation>
)
</taxonomicName>
, male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, habitus, dorsal view; B, eyes, antennae, and third maxillipeds, anteroventral view; C, cephalothorax, anterior view; D, left subhepatic cavity, anterolateral view; E, right chela, external view; F, right P4 and P5, dorsal view; G, male thoracic sternum and pleon, ventral view; H, male thoracic sternum and pleon, posteroventral view. Scale bars: A = 2.0 mm; B–H = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9A8536FF84FFF8AE2AFD6F0D5FFC8C" blockId="4.[835,1474,669,789]" pageId="4" pageNumber="467">
The live colouration of the 
<collectingCountry id="F332C5A6FF84FFF8A91EFD6F0DBCFD2C" box="[1143,1251,669,693]" name="Singapore" pageId="4" pageNumber="467">Singapore</collectingCountry>
specimen (
<figureCitation id="131E99B3FF84FFF8A80AFD6F0CE5FD2C" box="[1379,1466,669,693]" captionStart="Fig" captionStartId="4.[136,171,1727,1749]" captionTargetBox="[138,774,191,1712]" captionTargetId="figure-8@4.[133,777,186,1738]" captionTargetPageId="4" captionText="Fig. 3. Hepatoporus guinotae (Zarenkov, 1971), male, 8.5 × 6.6 mm (ZRC 2019.0663), Pulau Hantu, Singapore. A, in situ photograph showing live colouration (photographed by Toh Chay Hoon); B, left G1, pleonal view; C, distal tip of left G1, pleonal view; D, distal tip of left G1, sternal view. Scale bars: B = 0.5 mm; C, D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7174591" httpUri="https://zenodo.org/record/7174591/files/figure.png" pageId="4" pageNumber="467">Fig. 3A</figureCitation>
) is orange, with scattered and small patches of white on the carapace and pereopods. In life, the specimen was lightly covered with silt.
</paragraph>
<paragraph id="8B9A8536FF84FFF8AE2AFCCC0D01FBAC" blockId="4.[835,1474,829,1077]" pageId="4" pageNumber="467">
<taxonomicName id="4C25FEB5FF84FFF8AE2AFCCC0D61FCCC" authorityName="Serene" authorityYear="1984" baseAuthorityName="Zarenkov" baseAuthorityYear="1971" box="[835,1086,830,853]" class="Malacostraca" family="Xanthidae" genus="Hepatoporus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="B9515924FF84FFF8AE2AFCCC0D61FCCC" box="[835,1086,830,853]" italics="true" pageId="4" pageNumber="467">Hepatoporus guinotae</emphasis>
</taxonomicName>
is currently known from the Red Sea, the 
<typeStatus id="549E3B94FF84FFF8AECEFCAC0A8AFCEF" box="[935,981,862,886]" pageId="4" pageNumber="467">type</typeStatus>
locality (
<bibRefCitation id="EFB4F8C7FF84FFF8A92AFCAF0DA7FCEC" author="Zarenkov NA" box="[1091,1272,861,885]" pageId="4" pageNumber="467" pagination="155 - 203" refId="ref13681" refString="Zarenkov NA (1971) On the species composition and ecology of the Decapoda in the Red Sea. In: Vodyanitzkii VA (ed.) Benthos of the Continental Shelf of the Red Sea. Naukova Dumka, Kiev, pp. 155 - 203. [In Russian]" type="book chapter" year="1971">Zarenkov, 1971</bibRefCitation>
); 
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and the coast of 
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(
<bibRefCitation id="EFB4F8C7FF84FFF8A95EFC8F0D89FC0C" author="Serene R" box="[1079,1238,893,917]" pageId="4" pageNumber="467" pagination="1 - 400" refId="ref13004" refString="Serene R (1984) Crustaces Decapodes Brachyoures de l'Ocean Indien occidental et de la Mer Rouge. Xanthoidea: Xanthidae et Trapeziidae. Avec un addendum par A. Crosnier: Carpiliidae et Menippidae. Faune tropicale, 24: 1 - 400, pls. 1 - 48." type="journal article" year="1984">Serène, 1984</bibRefCitation>
); the Gulf of 
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(
<bibRefCitation id="EFB4F8C7FF84FFF8AE25FC6F0AA0FC2C" author="Naderloo R" box="[844,1023,925,949]" pageId="4" pageNumber="467" refId="ref12278" refString="Naderloo R (2017) Atlas of the Crabs of the Persian Gulf. Springer International Publishing, Cham, Switzerland, xxii + 444 pp." type="book" year="2017">Naderloo, 2017</bibRefCitation>
); the Northwest Shelf, 
<collectingCountry id="F332C5A6FF84FFF8A863FC6F0C31FC2C" box="[1290,1390,925,949]" name="Australia" pageId="4" pageNumber="467">Australia</collectingCountry>
(
<bibRefCitation id="EFB4F8C7FF84FFF8A8E9FC6F0AA9FC4C" author="Davie PJF &amp; Turner PA" pageId="4" pageNumber="467" pagination="83 - 86" refId="ref10712" refString="Davie PJF &amp; Turner PA (1994) A new species and a new record of Hepatoporus from north-western Australia (Crustacea: Decapoda: Xanthidae). Memoirs of the Queensland Museum, 37 (1): 83 - 86." type="journal article" year="1994">Davie &amp; Turner, 1994</bibRefCitation>
); Kii Peninsula, central 
<collectingCountry id="F332C5A6FF84FFF8A86DFC4C0C1EFC4C" box="[1284,1345,958,981]" name="Japan" pageId="4" pageNumber="467">Japan</collectingCountry>
(
<bibRefCitation id="EFB4F8C7FF84FFF8A83AFC4F0A93FC6C" author="Takeda M &amp; Nagai S" pageId="4" pageNumber="467" pagination="547 - 550" refId="ref13315" refString="Takeda M &amp; Nagai S (1986) Establishment of a new genus on two Japanese crabs. Zoological Science, 3: 547 - 550." type="journal article" year="1986">Takeda &amp; Nagai, 1986</bibRefCitation>
), the Ogasawara Islands (
<bibRefCitation id="EFB4F8C7FF84FFF8A98DFC2F0A22FB8C" author="Takeda M &amp; Komatsu H" pageId="4" pageNumber="467" pagination="153 - 189" refId="ref13265" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Nature and Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
), 
<collectingRegion id="49E14BD4FF84FFF8AEFAFC0F0AA8FB8C" box="[915,1015,1021,1045]" country="Japan" name="Okinawa" pageId="4" pageNumber="467">Okinawa</collectingRegion>
, southern 
<collectingCountry id="F332C5A6FF84FFF8A905FC0C0DF6FB8C" box="[1132,1193,1022,1045]" name="Japan" pageId="4" pageNumber="467">Japan</collectingCountry>
(
<bibRefCitation id="EFB4F8C7FF84FFF8A9D2FC0F0CDAFB8C" author="Maenosono T" box="[1211,1413,1021,1045]" pageId="4" pageNumber="467" pagination="21 - 33" refId="ref11999" refString="Maenosono T (2021) Notes on some euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) collected from the Ryukyu Islands, including a new record from Japan. Cancer, 30: 21 - 33." type="journal article" year="2021">Maenosono, 2021</bibRefCitation>
); and now also from 
<collectingCountry id="F332C5A6FF84FFF8AE83FBEF0D05FBAC" box="[1002,1114,1053,1077]" name="Singapore" pageId="4" pageNumber="467">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>