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<document id="FF3E25237F890A02BA38F63073D6F596" ID-CLB-Dataset="2631" ID-DOI="10.11646/zootaxa.5091.1.5" ID-GBIF-Dataset="b8e67802-3d14-4b63-85c6-6ee00ebc55d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840635" ID-ZooBank="7E744DAE-E9E6-4FB5-9897-1CCF03D373BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641987074997" checkinUser="plazi" docAuthor="Xu, Xin, Yu, Li, Liu, Fengxiang &amp; Li, Daiqin" docDate="2022" docId="038A8789FFAAFFCE16FE5F18C09AFDC2" docLanguage="en" docName="zootaxa.5091.1.5.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luthela dengfeng Xu &amp; Yu &amp; Liu &amp; Li 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="141" masterDocId="FFB3FFF1FFA2FFC416695C67C467FFA3" masterDocTitle="Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)" masterLastPageNumber="154" masterPageNumber="131" pageNumber="139" updateTime="1699301488680" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="690731E5786A7B069F00533DE6BA87C8">Delimitation of the segmented trapdoor spider genus Luthela gen. nov., with comments on the genus Sinothela from northern China (Araneae, Mesothelae, Liphistiidae)</mods:title>
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<mods:name id="EEB061B4592C1E72F43192114EC78FF0" type="personal">
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<mods:roleTerm id="F4162D3E00AD8371157336093D819E6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2CABDDAE50C94349D74F7736CF9E2F93">Xu, Xin</mods:namePart>
<mods:affiliation id="6C7F7586D6FF547FA59F13D3F213011D">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China &amp; School of Life and Environmental Sciences, University of Sydney, Sydney, NSW, Australia &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering &amp; Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
</mods:name>
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<mods:namePart id="6CF39CD6B638E675F9F4E23D97A4F151">Yu, Li</mods:namePart>
<mods:affiliation id="0526CC201AAFD33BDD4E69DB7FB37714">College of Life Sciences, Hunan Normal University, Changsha, Hunan, China</mods:affiliation>
</mods:name>
<mods:name id="CC8EFB7B1B665141576DC26ADBD27C02" type="personal">
<mods:role id="AC54CFAD24C039A6547F7637A3644277">
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</mods:role>
<mods:namePart id="BC58B79FC43F2A78048CA6D87B2F0027">Liu, Fengxiang</mods:namePart>
<mods:affiliation id="A50AC0A26080E4A8960AFA2962DE2182">State Key Laboratory of Biocatalysis and Enzyme Engineering &amp; Centre for Behavioural Ecology and Evolution, School of Life Sciences, Hubei University, Wuhan, Hubei, China</mods:affiliation>
</mods:name>
<mods:name id="0DF2BD6039511D5226FE675F65F1FE9E" type="personal">
<mods:role id="9B779062936DAEE918CAAFD375A6CE7F">
<mods:roleTerm id="AE8A6DCA62DC874ED030031693F5C562">Author</mods:roleTerm>
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<mods:namePart id="6DBA9995D769E1B4A22AB6BC3FADE6D6">Li, Daiqin</mods:namePart>
<mods:affiliation id="15A871EF33C7224D4DF3FD964B480000">Department of Biological Sciences, National University of Singapore, Singapore</mods:affiliation>
</mods:name>
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<mods:title id="9DE252D94E043CC85E67EAD861E8D1A8">Zootaxa</mods:title>
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<mods:part id="74C2C63D8ACFAA85F6BC644C7EEA773C">
<mods:date id="8FB640984AE02A0DD558D0535F3D8448">2022</mods:date>
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<mods:number id="516254E7DBFF80B2F761F3651FB5BCC1">2022-01-12</mods:number>
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<mods:number id="D5C0C45385795501070109424698659C">5091</mods:number>
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<mods:identifier id="0752869C9BF69D272BFCCDC9426CA084" type="DOI">10.11646/zootaxa.5091.1.5</mods:identifier>
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<treatment id="038A8789FFAAFFCE16FE5F18C09AFDC2" ID-DOI="http://doi.org/10.5281/zenodo.5840630" ID-GBIF-Taxon="192044609" ID-Zenodo-Dep="5840630" LSID="urn:lsid:plazi:treatment:038A8789FFAAFFCE16FE5F18C09AFDC2" httpUri="http://treatment.plazi.org/id/038A8789FFAAFFCE16FE5F18C09AFDC2" lastPageId="10" lastPageNumber="141" pageId="8" pageNumber="139">
<subSubSection id="C3396514FFAAFFCC16FE5F18C5B5FC39" box="[151,466,895,922]" pageId="8" pageNumber="139" type="nomenclature">
<paragraph id="8B9C369FFFAAFFCC16FE5F18C5B5FC39" blockId="8.[151,466,895,959]" box="[151,466,895,922]" pageId="8" pageNumber="139">
<heading id="D0D481F3FFAAFFCC16FE5F18C5B5FC39" bold="true" box="[151,466,895,922]" fontSize="11" level="1" pageId="8" pageNumber="139" reason="1">
<emphasis id="B957EA8DFFAAFFCC16FE5F18C5B5FC39" bold="true" box="[151,466,895,922]" pageId="8" pageNumber="139">
<taxonomicName id="4C234D1CFFAAFFCC16FE5F18C50AFC39" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[151,365,895,922]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC16FE5F18C50AFC39" bold="true" box="[151,365,895,922]" italics="true" pageId="8" pageNumber="139">Luthela dengfeng</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26457F6FFAAFFCC171C5FE7C5B5FC39" box="[373,466,896,922]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3396514FFAAFFCC16FE5FC3C540FC1C" box="[151,295,932,959]" pageId="8" pageNumber="139" type="description">
<paragraph id="8B9C369FFFAAFFCC16FE5FC3C540FC1C" blockId="8.[151,466,895,959]" box="[151,295,932,959]" pageId="8" pageNumber="139">
(
<figureCitation id="13182A1AFFAAFFCC16C95FC3C578FC1C" box="[160,287,932,959]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6AM</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFAAFFCC16FE5F8BC0A1FB35" pageId="8" pageNumber="139" type="materials_examined">
<paragraph id="8B9C369FFFAAFFCC16FE5F8BC481FBED" blockId="8.[151,1437,1004,2039]" pageId="8" pageNumber="139">
<materialsCitation id="3B4B3CC2FFAAFFCC16FE5F8BC481FBED" ID-GBIF-Occurrence="3435942322" collectingDate="2012-04-27" collectingDateMax="2012-07-29" collectingDateMin="2012-04-27" collectorName="F. X. Liu &amp; Z. L. Liu &amp; X. Xu" country="China" county="Dengfeng City" elevation="568" latitude="34.511" location="Male" longLatPrecision="71" longitude="112.95" municipality="Shaolin Temple" pageId="8" pageNumber="139" specimenCount="1" stateProvince="Henan" typeStatus="holotype">
<emphasis id="B957EA8DFFAAFFCC16FE5F8BC56CFBA5" bold="true" box="[151,267,1004,1030]" pageId="8" pageNumber="139">
<typeStatus id="5498883DFFAAFFCC16FE5F8BC561FBA5" box="[151,262,1004,1030]" pageId="8" pageNumber="139" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EFC6044FFAAFFCC177F5F8BC537FBA5" LSID="urn:lsid:plazi:treatment:038A8789FFAAFFCE16FE5F18C09AFDC2:8EFC6044FFAAFFCC177F5F8BC537FBA5" box="[278,336,1004,1030]" country="China" county="Dengfeng City" latitude="34.511" longLatPrecision="71" longitude="112.95" municipality="Shaolin Temple" name="Male" pageId="8" pageNumber="139" stateProvince="Henan">Male</location>
(XUX-2012-031, matured on
<date id="FF9D105FFFAAFFCC14DB5F8BC72DFBA5" box="[690,842,1004,1030]" pageId="8" pageNumber="139" value="2012-07-29">
<collectingDate id="EFD9E9B7FFAAFFCC14DB5F8BC72DFBA5" box="[690,842,1004,1030]" pageId="8" pageNumber="139" value="2012-07-29">29 July 2012</collectingDate>
</date>
at CBEE),
<collectingMunicipality id="6BF8ACE5FFAAFFCC15BF5F8BC0EBFBA5" box="[982,1164,1004,1030]" pageId="8" pageNumber="139">Shaolin Temple</collectingMunicipality>
,
<collectingCounty id="62FD4E13FFAAFFCC12F35F8BC123FBA5" box="[1178,1348,1004,1030]" pageId="8" pageNumber="139">Dengfeng City</collectingCounty>
,
<collectingRegion id="49E7F87DFFAAFFCC133A5F8AC49AFB89" country="China" name="Henan" pageId="8" pageNumber="139">Henan Province</collectingRegion>
,
<collectingCountry id="F334760FFFAAFFCC17645877C534FB89" box="[269,339,1040,1066]" name="China" pageId="8" pageNumber="139">China</collectingCountry>
,
<geoCoordinate id="EE175058FFAAFFCC170B5877C5A8FB88" box="[354,463,1040,1067]" degrees="34.511" direction="north" orientation="latitude" pageId="8" pageNumber="139" precision="55" value="34.511">34.511°N</geoCoordinate>
,
<geoCoordinate id="EE175058FFAAFFCC17B65877C63FFB88" box="[479,600,1040,1067]" degrees="112.950" direction="east" orientation="longitude" pageId="8" pageNumber="139" precision="55" value="112.95">112.950°E</geoCoordinate>
,
<quantity id="4CDB9B7AFFAAFFCC14015877C6D2FB88" box="[616,693,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="5.68" pageId="8" pageNumber="139" unit="m" value="568.0">
<elevation id="000ED1ACFFAAFFCC14015877C6D2FB88" box="[616,693,1040,1067]" metricMagnitude="2" metricUnit="m" metricValue="5.68" pageId="8" pageNumber="139" unit="m" value="568.0">568 m</elevation>
</quantity>
, collected on
<date id="FF9D105FFFAAFFCC15355877C798FB89" box="[860,1023,1040,1066]" pageId="8" pageNumber="139" value="2012-04-27">
<collectingDate id="EFD9E9B7FFAAFFCC15355877C798FB89" box="[860,1023,1040,1066]" pageId="8" pageNumber="139" value="2012-04-27">27 April 2012</collectingDate>
</date>
by
<collectorName id="26D65349FFAAFFCC12585876C0F1FB89" box="[1073,1174,1040,1067]" pageId="8" pageNumber="139">F.X. Liu</collectorName>
,
<collectorName id="26D65349FFAAFFCC12CC5876C16DFB89" box="[1189,1290,1040,1067]" pageId="8" pageNumber="139">Z.L. Liu</collectorName>
, and
<collectorName id="26D65349FFAAFFCC13245876C1FFFB88" box="[1357,1432,1041,1067]" pageId="8" pageNumber="139">X. Xu</collectorName>
, CBEE.
</materialsCitation>
</paragraph>
<paragraph id="8B9C369FFFAAFFCC16AE583FC0A1FB35" blockId="8.[151,1437,1004,2039]" pageId="8" pageNumber="139">
<materialsCitation id="3B4B3CC2FFAAFFCC16AE583FC78AFBD1" ID-GBIF-Occurrence="3435942320" box="[199,1005,1112,1139]" collectingDate="2012-04-27" collectingDateMax="2012-07-29" collectingDateMin="2012-04-27" collectorName="F. X. Liu &amp; Z. L. Liu &amp; X. Xu" country="China" county="Dengfeng City" elevation="568" latitude="34.511" location="Male" longLatPrecision="71" longitude="112.95" municipality="Shaolin Temple" pageId="8" pageNumber="139" specimenCount="2" specimenCount-female="2" stateProvince="Henan" typeStatus="paratype">
<emphasis id="B957EA8DFFAAFFCC16AE583FC521FBD1" bold="true" box="[199,326,1112,1138]" pageId="8" pageNumber="139">
<typeStatus id="5498883DFFAAFFCC16AE583FC526FBD1" box="[199,321,1112,1138]" pageId="8" pageNumber="139" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D25FD16FFAAFFCC1727583FC5DCFBD1" box="[334,443,1112,1138]" pageId="8" pageNumber="139" type="female">2 females</specimenCount>
(XUX-2012-023/030), same data as for holotype
</materialsCitation>
;
<materialsCitation id="3B4B3CC2FFAAFFCC1591583FC525FB35" ID-GBIF-Occurrence="3435942324" collectingDate="2012-04-27" collectingDateMax="2012-07-29" collectingDateMin="2012-04-27" collectorName="F. X. Liu &amp; Z. L. Liu &amp; X. Xu" country="China" county="Dengfeng City" elevation="568" latitude="34.511" location="Male" longLatPrecision="71" longitude="112.95" municipality="Shaolin Temple" pageId="8" pageNumber="139" specimenCount="1" specimenCount-female="1" stateProvince="Henan" typeStatus="paratype">
<specimenCount id="9D25FD16FFAAFFCC1591583FC03DFBD1" box="[1016,1114,1112,1138]" pageId="8" pageNumber="139" type="female">1 female</specimenCount>
(XUX-2016-111) same data as for holotype
</materialsCitation>
,
<materialsCitation id="3B4B3CC2FFAAFFCC1724581BC0A6FB35" ID-GBIF-Occurrence="3435942311" box="[333,1217,1148,1175]" collectingDate="2016-08-23" collectionCode="T" collectorName="F. Li &amp; F. X. Liu &amp; H. Liu" country="China" pageId="8" pageNumber="139" specimenCount="1" stateProvince="Henan" typeStatus="paratype">
collected on
<date id="FF9D105FFFAAFFCC17B5581BC6E9FB35" box="[476,654,1148,1174]" pageId="8" pageNumber="139" value="2016-08-23">
<collectingDate id="EFD9E9B7FFAAFFCC17B5581BC6E9FB35" box="[476,654,1148,1174]" pageId="8" pageNumber="139" value="2016-08-23">23 August 2016</collectingDate>
</date>
by
<collectorName id="26D65349FFAAFFCC14D1581AC689FB35" box="[696,750,1148,1175]" pageId="8" pageNumber="139">F. Li</collectorName>
,
<collectorName id="26D65349FFAAFFCC1493581AC73DFB35" box="[762,858,1148,1175]" pageId="8" pageNumber="139">F.X. Liu</collectorName>
,
<collectorName id="26D65349FFAAFFCC150C581AC7D6FB35" box="[869,945,1148,1175]" pageId="8" pageNumber="139">H. Liu</collectorName>
, and Z.
<collectionCode id="ED32AE5AFFAAFFCC126A581AC071FB34" box="[1027,1046,1149,1175]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="139">T</collectionCode>
. Zhang, CBEE
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFAAFFCC16AE58C7C0C3FA14" pageId="8" pageNumber="139" type="diagnosis">
<paragraph id="8B9C369FFFAAFFCC16AE58C7C0C3FA14" blockId="8.[151,1437,1004,2039]" pageId="8" pageNumber="139">
<emphasis id="B957EA8DFFAAFFCC16AE58C7C525FB19" bold="true" box="[199,322,1184,1210]" pageId="8" pageNumber="139">Diagnosis.</emphasis>
The male of
<taxonomicName id="4C234D1CFFAAFFCC17BD58C6C630FB19" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[468,599,1184,1210]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC17BD58C6C630FB19" box="[468,599,1184,1210]" italics="true" pageId="8" pageNumber="139">L. dengfeng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC143458C7C6D1FB19" bold="true" box="[605,694,1184,1210]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC143458C7C6D1FB19" box="[605,694,1184,1210]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from those of
<taxonomicName id="4C234D1CFFAAFFCC122858C6C0D5FB19" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1089,1202,1184,1210]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC122858C6C0D5FB19" box="[1089,1202,1184,1210]" italics="true" pageId="8" pageNumber="139">L. badong</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC12D158C7C176FB19" bold="true" box="[1208,1297,1184,1210]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC12D158C7C176FB19" box="[1208,1297,1184,1210]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C234D1CFFAAFFCC132F58C6C1FAFB19" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1350,1437,1185,1210]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC132F58C6C1FAFB19" box="[1350,1437,1185,1210]" italics="true" pageId="8" pageNumber="139">L. taian</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC16FE58A3C497FB7D" bold="true" box="[151,240,1220,1246]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC16FE58A3C497FB7D" box="[151,240,1220,1246]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the conductor having one inconspicuous middle spine (
<figureCitation id="13182A1AFFAAFFCC15E858A3C79AFB7C" box="[897,1021,1220,1247]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6DG</figureCitation>
), and from that of
<taxonomicName id="4C234D1CFFAAFFCC12A258A2C15BFB7D" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1227,1340,1220,1246]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC12A258A2C15BFB7D" box="[1227,1340,1220,1246]" italics="true" pageId="8" pageNumber="139">L. badong</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC132A58A3C1FBFB7D" bold="true" box="[1347,1436,1220,1246]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC132A58A3C1FBFB7D" box="[1347,1436,1220,1246]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the conductor with two thicker spines; from those of all the other
<taxonomicName id="4C234D1CFFAAFFCC15F7588FC02BFAA1" box="[926,1100,1256,1282]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B957EA8DFFAAFFCC15F7588FC793FAA1" box="[926,1012,1256,1282]" italics="true" pageId="8" pageNumber="139">Luthela</emphasis>
species
</taxonomicName>
by the conductor with a short lower spine, and a smaller angle between lower spine and conductor (
<figureCitation id="13182A1AFFAAFFCC15CD596BC045FA84" box="[932,1058,1292,1319]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6DG</figureCitation>
). Females of
<taxonomicName id="4C234D1CFFAAFFCC12DE596AC15BFA85" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1207,1340,1292,1318]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC12DE596AC15BFA85" box="[1207,1340,1292,1318]" italics="true" pageId="8" pageNumber="139">L. dengfeng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC132A596BC1FBFA85" bold="true" box="[1347,1436,1292,1318]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC132A596BC1FBFA85" box="[1347,1436,1292,1318]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differ from those of
<taxonomicName id="4C234D1CFFAAFFCC171C5956C582FAE9" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[373,485,1328,1354]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="badong" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC171C5956C582FAE9" box="[373,485,1328,1354]" italics="true" pageId="8" pageNumber="139">L. badong</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC17825957C623FAE9" bold="true" box="[491,580,1328,1354]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC17825957C623FAE9" box="[491,580,1328,1354]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C234D1CFFAAFFCC14395956C693FAE9" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Yin, Tang, Zhao &amp; Chen" baseAuthorityYear="2002" box="[592,756,1328,1354]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="luotianensis">
<emphasis id="B957EA8DFFAAFFCC14395956C693FAE9" box="[592,756,1328,1354]" italics="true" pageId="8" pageNumber="139">L. luotianensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C234D1CFFAAFFCC15465956C7D1FAE9" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[815,950,1328,1354]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="yuncheng" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC15465956C7D1FAE9" box="[815,950,1328,1354]" italics="true" pageId="8" pageNumber="139">L. yuncheng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC15D45957C072FAE9" bold="true" box="[957,1045,1328,1354]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC15D45957C072FAE9" box="[957,1045,1328,1354]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the middle receptacular clusters being smaller than lateral ones (
<figureCitation id="13182A1AFFAAFFCC17955933C617FACC" box="[508,624,1364,1391]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6HJ</figureCitation>
); from those of
<taxonomicName id="4C234D1CFFAAFFCC15495932C7A0FACD" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" baseAuthorityName="Schenkel" baseAuthorityYear="1953" box="[800,967,1364,1390]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="schensiensis">
<emphasis id="B957EA8DFFAAFFCC15495932C7A0FACD" box="[800,967,1364,1390]" italics="true" pageId="8" pageNumber="139">L. schensiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C234D1CFFAAFFCC15975932C001FACD" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1022,1126,1365,1390]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="yiyuan" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC15975932C001FACD" box="[1022,1126,1365,1390]" italics="true" pageId="8" pageNumber="139">L. yiyuan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC12045933C0A1FACD" bold="true" box="[1133,1222,1364,1390]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC12045933C0A1FACD" box="[1133,1222,1364,1390]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the bases of the middle receptacular clusters separated from each other (
<figureCitation id="13182A1AFFAAFFCC1560591FC704FA30" box="[777,867,1400,1427]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6H</figureCitation>
); from those of
<taxonomicName id="4C234D1CFFAAFFCC127C591EC0E1FA31" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1045,1158,1400,1426]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="handan" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC127C591EC0E1FA31" box="[1045,1158,1400,1426]" italics="true" pageId="8" pageNumber="139">L. handan</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC12E4591FC080FA31" bold="true" box="[1165,1255,1400,1426]" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC12E4591FC080FA31" box="[1165,1255,1400,1426]" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C234D1CFFAAFFCC1374591EC113FA31" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[1309,1396,1401,1426]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="139" phylum="Arthropoda" rank="species" species="taian" status="sp. nov.">
<emphasis id="B957EA8DFFAAFFCC1374591EC113FA31" box="[1309,1396,1401,1426]" italics="true" pageId="8" pageNumber="139">L. taian</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFAAFFCC1312591FC4AFFA15" bold="true" pageId="8" pageNumber="139">
<taxonomicNameLabel id="A26457F6FFAAFFCC1312591FC4AFFA15" pageId="8" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the middle receptacular clusters with slender and slightly longer stalks (
<figureCitation id="13182A1AFFAAFFCC127C59FBC0FFFA14" box="[1045,1176,1436,1463]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6HM</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFAAFFCE16AE59A7C512FE0D" lastPageId="10" lastPageNumber="141" pageId="8" pageNumber="139" type="description">
<paragraph id="8B9C369FFFAAFFCC16AE59A7C133F92C" blockId="8.[151,1437,1004,2039]" pageId="8" pageNumber="139">
<emphasis id="B957EA8DFFAAFFCC16AE59A7C53EFA79" bold="true" box="[199,345,1472,1498]" pageId="8" pageNumber="139">Description.</emphasis>
Male (
<typeStatus id="5498883DFFAAFFCC17C359A7C67EFA78" box="[426,537,1472,1499]" pageId="8" pageNumber="139" type="holotype">Holotype</typeStatus>
). Carapace brown; opisthosoma light brown, with dark brown tergites; sternum narrow, much longer than wide; a few long pointed hairs running over ocular mound in a longitudinal row; chelicerae robust with promargin of groove with 910 denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, the second to fifth larger than others and the others small; 7 spinnerets. Measurements: BL 12.50, CL 6.00, CW 5.40, OL 6.90, OW 5.00; ALE&gt; PLE&gt; PME&gt; AME; leg I 17.35 (5.10 + 2.35 + 3.55 + 4.05 + 2.30), leg II miss, leg III 18.10 (4.40 + 2.35 + 3.25 + 5.20 + 2.90), leg IV 23.95 (5.55 + 2.45 + 4.45 + 7.65 + 3.85).
</paragraph>
<paragraph id="8B9C369FFFAAFFCC16AE5AFFC657F80C" blockId="8.[151,1437,1004,2039]" pageId="8" pageNumber="139">
Palp: Prolateral paracymbium unpigmented and unsclerotized, numerous setae and spines at tip of paracymbium (
<figureCitation id="13182A1AFFAAFFCC16C95ADBC57AF974" box="[160,285,1724,1751]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6AC</figureCitation>
). Contrategulum with 6 teeth along the margin (
<figureCitation id="13182A1AFFAAFFCC15235ADBC7C1F974" box="[842,934,1724,1751]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6D</figureCitation>
), and distal one with 4 serrations. Marginal apophysis of tegulum with several serrations proximally and smooth distally, dorsal extension of terminal apophysis of tegulum with irregular serrations, and terminal apophysis of tegulum smooth, triangle-shaped (
<figureCitation id="13182A1AFFAAFFCC13065B63C48FF8E0" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6F, 6G</figureCitation>
). Conductor situated ventro-proximally on embolus, with two large spines, upper one parallel to embolus, the angle between lower one and conductor about 30 ventrally (
<figureCitation id="13182A1AFFAAFFCC150F5B2BC7D9F8C5" box="[870,958,1868,1894]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6B</figureCitation>
,
<figureCitation id="13182A1AFFAAFFCC15A35B2BC78BF8C5" box="[970,1004,1868,1894]" captionStart="FIGURE 4" captionStartId="6.[151,250,1251,1275]" captionTargetBox="[164,1423,381,1228]" captionTargetId="figure-115@6.[164,1423,381,1228]" captionTargetPageId="6" captionText="FIGURE 4. Female genital anatomy of Luthela yiyuan sp. nov. AE, KO, vulva dorsal view. FJ, PT, vulva ventral view. A, F, XUX-2012-034. B, G, XUX-2012-035. C, H, XUX-2012-049. D, I, XUX-2016-101. E, J, XUX-2016-107. K, P, XUX-2012- 042. L, Q, XUX-2012-040. M, R, XUX-2012-046. N, S, XUX-2016-103. O, T, XUX-2016-108. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840651" httpUri="https://zenodo.org/record/5840651/files/figure.png" pageId="8" pageNumber="139">4E</figureCitation>
), and one inconspicuous spine located at between two large spines (
<figureCitation id="13182A1AFFAAFFCC17B45B17C63CF828" box="[477,603,1904,1931]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6DG</figureCitation>
). Embolus largely sclerotized, with a flat opening and several longitudinal ribs retrolaterally (
<figureCitation id="13182A1AFFAAFFCC17015BF3C644F80C" box="[360,547,1940,1967]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="8" pageNumber="139">Fig. 6A, 6D, 6G</figureCitation>
).
</paragraph>
<paragraph id="8B9C369FFFAAFFCE16AE5BDFC558FEBE" blockId="8.[151,1437,1004,2039]" lastBlockId="10.[151,1437,151,610]" lastPageId="10" lastPageNumber="141" pageId="8" pageNumber="139">Female (XUX-2012-030). Carapace and opisthosoma similar to male; chelicerae robust with promargin of groove with 1011 strong denticles of variable size; legs with strong hairs and spines; opisthosoma with 12 tergites, similar to male; 7 spinnerets. Measurements: BL 17.80, CL 6.90, CW 6.60, OL 11.30, OW 10.20; ALE&gt; PLE&gt; PME&gt; AME; palp 11.20 (3.90 + 1.80 + 2.60 + 2.90), leg I 13.70 (4.60 + 1.90 + 2.80 + 2.70 + 1.70), leg II 12.90 (4.00 + 2.20 + 2.40 + 2.50 + 1.80), leg III 13.90 (4.30 + 2.20 + 2.40 + 3.10 + 1.90), leg IV 21.00 (5.90 + 2.80 + 3.80 + 5.70 + 2.80).</paragraph>
<caption id="DF5C6617FFABFFCD16FE5BF0C0E2F854" ID-DOI="http://doi.org/10.5281/zenodo.5840657" ID-Zenodo-Dep="5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="9" pageNumber="140" startId="9.[151,250,1943,1967]" targetBox="[203,1385,185,1919]" targetPageId="9">
<paragraph id="8B9C369FFFABFFCD16FE5BF0C0E2F854" blockId="9.[151,1436,1943,2040]" pageId="9" pageNumber="140">
<emphasis id="B957EA8DFFABFFCD16FE5BF0C571F80C" bold="true" box="[151,278,1943,1967]" pageId="9" pageNumber="140">FIGURE 6.</emphasis>
Male and female genital anatomy of
<taxonomicName id="4C234D1CFFABFFCD14CF5BFFC73DF813" authority="Xu &amp; Yu &amp; Liu &amp; Li, 2022" authorityName="Xu &amp; Yu &amp; Liu &amp; Li" authorityYear="2022" box="[678,858,1944,1968]" class="Arachnida" family="Liphistiidae" genus="Luthela" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="140" phylum="Arthropoda" rank="species" species="dengfeng" status="sp. nov.">
<emphasis id="B957EA8DFFABFFCD14CF5BFFC73DF813" box="[678,858,1944,1968]" italics="true" pageId="9" pageNumber="140">Luthela dengfeng</emphasis>
</taxonomicName>
<emphasis id="B957EA8DFFABFFCD150A5BFFC7D1F813" bold="true" box="[867,950,1944,1968]" pageId="9" pageNumber="140">
<taxonomicNameLabel id="A26457F6FFABFFCD150A5BFFC7D1F813" box="[867,950,1944,1968]" pageId="9" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B9C369FFFA8FFCE16AE5D40C512FE0D" blockId="10.[151,1437,151,610]" pageId="10" pageNumber="141">
Genitalia: Posterior margin of genital sternite slightly incurved (
<figureCitation id="13182A1AFFA8FFCE15F35D40C06AFEE1" box="[922,1037,295,322]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="10" pageNumber="141">Fig. 6HJ</figureCitation>
), two pairs of receptacular clusters, middle ones smaller than lateral ones with slender stalks, separated from each other basally, and situated at anterior margin of bursa copulatrix, lateral ones situated at slightly dorsolateral position of bursa copulatrix with short thick stalks (
<figureCitation id="13182A1AFFA8FFCE168F5DF4C50EFE0D" box="[230,361,403,430]" captionStart="FIGURE 6" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[203,1385,185,1919]" captionTargetId="figure-19@9.[203,1385,181,1919]" captionTargetPageId="9" captionText="FIGURE 6. Male and female genital anatomy of Luthela dengfeng sp. nov. A, D, palp prolateral view. B, E, palp ventral view. C, F, palp retrolateral view. G, palp distal view. HJ, vulva dorsal view. KM, vulva ventral view. AG, XUX-2012-031 (holotype). H, K, XUX-2012-023. I, L, XUX-2012-030. J, M, XUX-2016-111. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5840657" httpUri="https://zenodo.org/record/5840657/files/figure.png" pageId="10" pageNumber="141">Fig. 6HM</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFA8FFCE16AE5DD0C02FFE72" box="[199,1096,439,465]" pageId="10" pageNumber="141" type="etymology">
<paragraph id="8B9C369FFFA8FFCE16AE5DD0C02FFE72" blockId="10.[151,1437,151,610]" box="[199,1096,439,465]" pageId="10" pageNumber="141">
<emphasis id="B957EA8DFFA8FFCE16AE5DD0C529FE72" bold="true" box="[199,334,439,465]" pageId="10" pageNumber="141">Etymology.</emphasis>
The species epithet, a noun in apposition, refers to the
<typeStatus id="5498883DFFA8FFCE15DE5DD0C780FE72" box="[951,999,439,465]" pageId="10" pageNumber="141">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFA8FFCE16AE5DBDC6C3FE55" box="[199,676,474,502]" pageId="10" pageNumber="141" type="distribution">
<paragraph id="8B9C369FFFA8FFCE16AE5DBDC6C3FE55" blockId="10.[151,1437,151,610]" box="[199,676,474,502]" pageId="10" pageNumber="141">
<emphasis id="B957EA8DFFA8FFCE16AE5DBDC506FE57" bold="true" box="[199,353,474,500]" pageId="10" pageNumber="141">Distribution.</emphasis>
<collectingRegion id="49E7F87DFFA8FFCE170E5DBCC67BFE56" box="[359,540,475,501]" country="China" name="Henan" pageId="10" pageNumber="141">Henan Province</collectingRegion>
(Dengfeng)
</paragraph>
</subSubSection>
<subSubSection id="C3396514FFA8FFCE16AE5D98C09AFDC2" pageId="10" pageNumber="141" type="discussion">
<paragraph id="8B9C369FFFA8FFCE16AE5D98C09AFDC2" blockId="10.[151,1437,151,610]" pageId="10" pageNumber="141">
<emphasis id="B957EA8DFFA8FFCE16AE5D98C55CFDBA" bold="true" box="[199,315,511,537]" pageId="10" pageNumber="141">Remarks.</emphasis>
Female variation (
<emphasis id="B957EA8DFFA8FFCE14715D98C64CFDBB" box="[536,555,511,536]" italics="true" pageId="10" pageNumber="141">N</emphasis>
= 3): BL 17.8021.30, CL 6.908.40, CW 6.607.80, OL 11.3012.60, OW 10.2011.20. No intraspecific variation was found among the
<emphasis id="B957EA8DFFA8FFCE15135E44C7CDFD9E" box="[890,938,547,573]" italics="true" pageId="10" pageNumber="141">COI</emphasis>
nucleotide sequences of
<specimenCount id="9D25FD16FFA8FFCE12B25E44C1FAFD9E" box="[1243,1437,547,573]" pageId="10" pageNumber="141" type="generic">seven specimens</specimenCount>
(including juveniles). The GenBank accession code of the
<typeStatus id="5498883DFFA8FFCE154F5E20C7EFFDC2" box="[806,904,583,609]" pageId="10" pageNumber="141" type="holotype">holotype</typeStatus>
(XUX-2012-031) is
<accessionNumber id="9470AB7CFFA8FFCE121D5E20C09EFDC2" box="[1140,1273,583,609]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH172686" pageId="10" pageNumber="141">MH172686</accessionNumber>
.
</paragraph>
</subSubSection>
</treatment>
</document>