438 lines
58 KiB
XML
438 lines
58 KiB
XML
<document id="45DA9089419B885BC938614441C5E3E7" ID-CLB-Dataset="2971" ID-DOI="10.11646/zootaxa.5081.4.4" ID-GBIF-Dataset="64dc3da2-ee36-4b47-bad3-5c47763359dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="5778843" ID-ZooBank="38021A9F-DE16-4BE6-A40C-2750C34DC00E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639472157431" checkinUser="plazi" docAuthor="Moravec, Jiří & Trýzna, Miloš" docDate="2021" docId="038987BBFFD6582EFF5DFB00FEF7DC3D" docLanguage="en" docName="zootaxa.5081.4.4.pdf" docOrigin="Zootaxa 5081 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pogonostoma (Microgeniatum) signifemorale Moravec & Trýzna 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="532" masterDocId="FFB0FFC3FFD55826FFCAFFDAFFD2DF60" masterDocTitle="New or rare Madagascar tiger beetles- 24. Pogonostoma (Microgeniatum) signifemorale sp. nov. with revised key to species of the subgenus Microgeniatum Rivalier (Coleoptera: Cicindelidae)" masterLastPageNumber="534" masterPageNumber="524" pageNumber="527" updateTime="1699472477359" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="203BBF075DBF124129EABC2A9539E097">
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<mods:title id="4DF5BFE9DBE1AD913E9ADCD2DBA2E2AE">New or rare Madagascar tiger beetles- 24. Pogonostoma (Microgeniatum) signifemorale sp. nov. with revised key to species of the subgenus Microgeniatum Rivalier (Coleoptera: Cicindelidae)</mods:title>
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<mods:name id="A6FE2D1181F8A1E76A6C175DCA368DA5" type="personal">
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<mods:roleTerm id="E5531F416742691F0B4B549FD09D29A6">Author</mods:roleTerm>
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<mods:namePart id="69CB350AF3C605E67F8E84E5FE65EAC3">Moravec, Jiří</mods:namePart>
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<mods:nameIdentifier id="77EF5263E4C1398E8AB774B3CBD8A6F5" type="ORCID">0000-0001-5294-6410</mods:nameIdentifier>
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<mods:affiliation id="81C4C7F3B4A79F40AB81243CA62499CE">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. & jirmor @ quick. cz; https: // orcid. org / 0000 - 0001 - 5294 - 6410</mods:affiliation>
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<mods:nameIdentifier id="A75D779A04D91E61160579641510AF05" type="email">jirmor@quick.cz</mods:nameIdentifier>
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<mods:namePart id="F4FF2AF470B0D9C1EB3DD12E851CB19B">Trýzna, Miloš</mods:namePart>
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<mods:affiliation id="6A65B13DEA293DBF9EBD2366CCF84E53">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, CZ- 613 00 Brno, Czech Republic. & anthribidae @ gmail. com; https: // orcid. org / 0000 - 0002 - 0426 - 3348</mods:affiliation>
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<mods:nameIdentifier id="C818166AB04F16055D9014903AD3F674" type="email">anthribidae@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="24A5598E2F413A8F2D100C92A667ADC6">text</mods:typeOfResource>
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<mods:title id="1AA58915EF7811E8CEF6E5ED322FE6D9">Zootaxa</mods:title>
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<mods:part id="FEC5AB267865C05E4AEBF7BC09EEE4A8">
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<mods:date id="C6C850A892B44332DCD377EAFAA4D5AA">2021</mods:date>
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<mods:number id="965986D4FF11E8FD285113A182B644F1">2021-12-14</mods:number>
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<mods:number id="D589B0BC655DA51DAA0822DDFF1216BD">5081</mods:number>
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<treatment id="038987BBFFD6582EFF5DFB00FEF7DC3D" ID-DOI="http://doi.org/10.5281/zenodo.5779894" ID-GBIF-Taxon="191561777" ID-Zenodo-Dep="5779894" LSID="urn:lsid:plazi:treatment:038987BBFFD6582EFF5DFB00FEF7DC3D" httpUri="http://treatment.plazi.org/id/038987BBFFD6582EFF5DFB00FEF7DC3D" lastPageId="8" lastPageNumber="532" pageId="3" pageNumber="527">
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<subSubSection id="C33A6526FFD65825FF5DFB00FCF7DB93" box="[151,805,1241,1268]" pageId="3" pageNumber="527" type="nomenclature">
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<paragraph id="8B9F36ADFFD65825FF5DFB00FCF7DB93" blockId="3.[151,805,1241,1303]" box="[151,805,1241,1268]" pageId="3" pageNumber="527">
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<heading id="D0D781C1FFD65825FF5DFB00FCF7DB93" bold="true" box="[151,805,1241,1268]" fontSize="11" level="1" pageId="3" pageNumber="527" reason="1">
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<emphasis id="B954EABFFFD65825FF5DFB00FCF7DB93" bold="true" box="[151,805,1241,1268]" pageId="3" pageNumber="527">
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<taxonomicName id="4C204D2EFFD65825FF5DFB00FD6DDB94" authority="Moravec & Trýzna, 2021" authorityName="Moravec & Trýzna" authorityYear="2021" box="[151,703,1241,1268]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
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<emphasis id="B954EABFFFD65825FF5DFB00FEEEDB94" bold="true" box="[151,316,1242,1268]" italics="true" pageId="3" pageNumber="527">Pogonostoma</emphasis>
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(
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<emphasis id="B954EABFFFD65825FE87FB03FDDADB93" bold="true" box="[333,520,1241,1267]" italics="true" pageId="3" pageNumber="527">Microgeniatum</emphasis>
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)
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<emphasis id="B954EABFFFD65825FDD3FB03FD6DDB94" bold="true" box="[537,703,1241,1268]" italics="true" pageId="3" pageNumber="527">signifemorale</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26757C4FFD65825FD0DFB03FCF7DB93" box="[711,805,1241,1267]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A6526FFD65825FF5DFB27FE95DA77" box="[151,327,1277,1303]" pageId="3" pageNumber="527" type="description">
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<paragraph id="8B9F36ADFFD65825FF5DFB27FE95DA77" blockId="3.[151,805,1241,1303]" box="[151,327,1277,1303]" pageId="3" pageNumber="527">
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(
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<figureCitation id="131B2A28FFD65825FF55FB27FEC6DA77" box="[159,276,1277,1303]" captionStart-0="FIGURES 1–8" captionStart-1="FIGURE 9–10" captionStartId-0="5.[151,264,1959,1983]" captionStartId-1="6.[151,250,1867,1891]" captionTargetBox-0="[161,1431,278,1929]" captionTargetBox-1="[151,1435,181,1844]" captionTargetId-0="figure-72@5.[151,1435,270,1933]" captionTargetId-1="figure-17@6.[151,1435,181,1844]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." captionText-1="FIGURE 9–10. Pogonostoma (Microgeniatum) signifemorale sp. nov., Ankarafantsika National Park. 9—collecting site; 10— habitat of the new species." figureDoi-0="http://doi.org/10.5281/zenodo.5778845" figureDoi-1="http://doi.org/10.5281/zenodo.5778847" httpUri-0="https://zenodo.org/record/5778845/files/figure.png" httpUri-1="https://zenodo.org/record/5778847/files/figure.png" pageId="3" pageNumber="527">Figs 1–10</figureCitation>
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,
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<figureCitation id="131B2A28FFD65825FEEAFB27FEEFDA77" box="[288,317,1277,1303]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[233,1354,193,1908]" captionTargetId="figure-29@7.[218,1369,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Distribution of Pogonostoma (Microgeniatum) in Madagascar. 11—P. (M.) pseudominimum W. Horn; 12— P. (M.) infimum Rivalier; 13—P. (M.) flavopalpale Jeannel; 14—P. (M.) signifemorale sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5778849" httpUri="https://zenodo.org/record/5778849/files/figure.png" pageId="3" pageNumber="527">14</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A6526FFD65825FF5DFA9FFA8BDA90" pageId="3" pageNumber="527" type="materials_examined">
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<paragraph id="8B9F36ADFFD65825FF5DFA9FFCDBDAE4" blockId="3.[151,1437,1349,2024]" pageId="3" pageNumber="527">
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<materialsCitation id="3B483CF0FFD65825FF5DFA9FFCD7DAE4" ID-GBIF-Occurrence="3418275301" collectorName="Province" country="Madagascar" location="Ambato Boeni Region" municipality="Ankarafantsika National Park" pageId="3" pageNumber="527" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
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||
<emphasis id="B954EABFFFD65825FF5DFA9FFEE7DA3F" bold="true" box="[151,309,1349,1375]" pageId="3" pageNumber="527">
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||
<typeStatus id="549B880FFFD65825FF5DFA9FFF01DA3F" box="[151,211,1349,1375]" pageId="3" pageNumber="527">Type</typeStatus>
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locality
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</emphasis>
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||
. Northwestern
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||
<collectingCountry id="F337763DFFD65825FE21FA9FFDABDA3F" box="[491,633,1349,1375]" name="Madagascar" pageId="3" pageNumber="527">Madagascar</collectingCountry>
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||
: dry deciduous forest of the
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<collectingMunicipality id="6BFBACD7FFD65825FC01FA9FFAF4DA3F" box="[971,1318,1349,1375]" pageId="3" pageNumber="527">Ankarafantsika National Park</collectingMunicipality>
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, “Tour du Lac” (
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<location id="8EFF6076FFD65825FF17FAB3FE30DAE3" LSID="urn:lsid:plazi:treatment:038987BBFFD6582EFF5DFB00FEF7DC3D:8EFF6076FFD65825FF17FAB3FE30DAE3" box="[221,482,1385,1411]" country="Madagascar" municipality="Ankarafantsika National Park" name="Ambato Boeni Region" pageId="3" pageNumber="527" stateProvince="Mahajanga">Ambato Boeni Region</location>
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||
,
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<collectorName id="26D5537BFFD65825FE24FAB3FD81DAE3" box="[494,595,1385,1411]" pageId="3" pageNumber="527">Province</collectorName>
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of
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||
<collectingRegion id="49E4F84FFFD65825FDB0FAB3FD2CDAE4" box="[634,766,1385,1412]" country="Madagascar" name="Mahajanga" pageId="3" pageNumber="527">Mahajanga</collectingRegion>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B9F36ADFFD65825FF0DFA57FA8BDA90" blockId="3.[151,1437,1349,2024]" pageId="3" pageNumber="527">
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<emphasis id="B954EABFFFD65825FF0DFA57FEA0DAC7" bold="true" box="[199,370,1421,1447]" pageId="3" pageNumber="527">Type material</emphasis>
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||
.
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<materialsCitation id="3B483CF0FFD65825FE4BFA57FA8BDA90" ID-GBIF-Occurrence="3418275302" collectingDate="2015-01-23" collectingDateMax="2015-01-26" collectingDateMin="2015-01-23" collectionCode="NMPC, JVCB" country="Madagascar" elevation="71" latitude="-16.30361" location="Madagascar" longLatPrecision="1" longitude="46.813637" pageId="3" pageNumber="527" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549B880FFFD65825FE4BFA57FE39DAC7" box="[385,491,1421,1447]" pageId="3" pageNumber="527" type="holotype">Holotype</typeStatus>
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||
in
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<collectionCode id="ED31AE68FFD65825FDDBFA57FDB3DAC7" box="[529,609,1421,1447]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="527">NMPC</collectionCode>
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(temporarily in
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<collectionCode id="ED31AE68FFD65825FCD5FA57FCBBDAC7" box="[799,873,1421,1447]" pageId="3" pageNumber="527">JVCB</collectionCode>
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), labelled: “NW
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<collectingCountry id="F337763DFFD65825FBE1FA57FB64DAC7" box="[1067,1206,1421,1447]" name="Madagascar" pageId="3" pageNumber="527">Madagascar</collectingCountry>
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,
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<quantity id="4CD89B48FFD65825FB08FA57FAD2DAC8" box="[1218,1280,1421,1448]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="527" unit="m" value="71.0">
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<elevation id="000DD19EFFD65825FB08FA57FAD2DAC8" box="[1218,1280,1421,1448]" metricMagnitude="1" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="527" unit="m" value="71.0">71 m</elevation>
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</quantity>
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, / Ankarafan- tsika N.P., Tour du Lac /
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<geoCoordinate id="EE14506AFFD65825FE7EFA6BFDB3DAAB" box="[436,609,1457,1483]" degrees="16" direction="south" minutes="18" orientation="latitude" pageId="3" pageNumber="527" precision="1" seconds="13.0" value="-16.30361">S 16°18´13.0´´</geoCoordinate>
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||
;
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<geoCoordinate id="EE14506AFFD65825FDA7FA68FCC9DAAB" box="[621,795,1457,1484]" degrees="46" direction="east" minutes="48" orientation="longitude" pageId="3" pageNumber="527" precision="1" seconds="49.1" value="46.813637">E 46°48´49.1´´</geoCoordinate>
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, /
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<date id="FF9E106DFFD65825FCFCFA6BFC08DAAC" box="[822,986,1457,1484]" pageId="3" pageNumber="527" value="2015-01-23" valueMax="2015-01-26" valueMin="2015-01-23">
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||
<collectingDate id="EFDAE985FFD65825FCFCFA6BFC08DAAC" box="[822,986,1457,1484]" pageId="3" pageNumber="527" value="2015-01-23" valueMax="2015-01-26" valueMin="2015-01-23">23.- 26.i.2015</collectingDate>
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||
</date>
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||
,
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||
<collectorName id="26D5537BFFD65825FC2FFA68FB8EDAAB" box="[997,1116,1457,1484]" pageId="3" pageNumber="527">M. Trýzna</collectorName>
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leg.” [printed] // “Pogonos- toma / (
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||
<taxonomicName id="4C204D2EFFD65825FF24FA0FFE70DA90" box="[238,418,1493,1520]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">Microgeniatum</taxonomicName>
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||
) / signifemorale
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<taxonomicNameLabel id="A26757C4FFD65825FDAEFA0CFD6CDA8F" box="[612,702,1493,1520]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
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||
” / det
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<collectorName id="26D5537BFFD65825FCC3FA0FFC48DA90" box="[777,922,1493,1520]" pageId="3" pageNumber="527">Jiří Moravec</collectorName>
|
||
/ &
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||
<collectorName id="26D5537BFFD65825FC04FA0FFBC2DA8F" box="[974,1040,1493,1519]" pageId="3" pageNumber="527">Miloš</collectorName>
|
||
<bibRefCitation id="EFB14B5CFFD65825FBDDFA0FFB7BDA8F" author="Tryzna, M. & Horak, J. & Holusa, J." box="[1047,1193,1493,1519]" pageId="3" pageNumber="527" pagination="1 - 17" refId="ref7200" refString="Tryzna, M., Horak, J. & Holusa, J. (2021) Open canopy increases the species richness of fungus weevils in Madagascar forests. Forest Ecology and Management, 480, 1 - 17. https: // doi. org / 10.1016 / j. foreco. 2020.118661" type="journal article" year="2021">Trýzna 2021</bibRefCitation>
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||
” [red, printed].
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||
</materialsCitation>
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C33A6526FFD65822FF0DFA23FBABDE7D" lastPageId="4" lastPageNumber="528" pageId="3" pageNumber="527" type="diagnosis">
|
||
<paragraph id="8B9F36ADFFD65825FF0DFA23FDB8D887" blockId="3.[151,1437,1349,2024]" pageId="3" pageNumber="527">
|
||
<emphasis id="B954EABFFFD65825FF0DFA23FE1CD973" bold="true" box="[199,462,1529,1555]" pageId="3" pageNumber="527">Differential diagnosis</emphasis>
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. By its extremely small body and boomerang-bent aedeagus with similarly shaped apex,
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<emphasis id="B954EABFFFD65825FF11F9C4FF3ED957" box="[219,236,1566,1591]" italics="true" pageId="3" pageNumber="527">P</emphasis>
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||
.
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||
<taxonomicName id="4C204D2EFFD65825FF30F9C7FE19D957" authority="Moravec & Trýzna, 2021" authorityName="Moravec & Trýzna" authorityYear="2021" box="[250,459,1565,1592]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD65825FEC9F9C4FEC8D957" box="[259,282,1566,1591]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD65825FEFBF9C7FE19D957" box="[305,459,1565,1591]" italics="true" pageId="3" pageNumber="527">signifemorale</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EABFFFD65825FE18F9C7FDF9D957" bold="true" box="[466,555,1565,1591]" pageId="3" pageNumber="527">
|
||
<taxonomicNameLabel id="A26757C4FFD65825FE18F9C7FDF9D957" box="[466,555,1565,1591]" pageId="3" pageNumber="527" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
may resemble
|
||
<emphasis id="B954EABFFFD65825FD1DF9C4FD3AD957" box="[727,744,1566,1591]" italics="true" pageId="3" pageNumber="527">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD65825FD3CF9C7FC58D957" authorityName="Rivalier" authorityYear="1970" box="[758,906,1565,1592]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD65825FCCAF9C4FCC5D957" box="[768,791,1566,1591]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD65825FCE4F9C7FC58D957" box="[814,906,1565,1591]" italics="true" pageId="3" pageNumber="527">infimum</emphasis>
|
||
</taxonomicName>
|
||
. The new species is immediately distinguished from it, however, and from other three species of the subgenus by its markedly bicoloured meso- and metafemora (
|
||
<figureCitation id="131B2A28FFD65825FF6AF9BFFF30D91F" box="[160,226,1637,1663]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="3" pageNumber="527">Fig. 1</figureCitation>
|
||
) with their basal third yellow, while their remaining, markedly dilated portions are black; moreover, also the basal area of all tibiae is yellow. Antennomeres of the new species are uniquely coloured as well: the scape is black dorsally, yellow-testaceous apically; antennomeres 2–4 ivory to yellow-testaceous (
|
||
<figureCitation id="131B2A28FFD65825FB4CF977FAD8D9A7" box="[1158,1290,1709,1735]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="3" pageNumber="527">Figs 1, 3–4</figureCitation>
|
||
). In contrast, the femora and tibiae, as well as the antennae of
|
||
<emphasis id="B954EABFFFD65825FD04F908FD0DD98B" box="[718,735,1746,1771]" italics="true" pageId="3" pageNumber="527">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD65825FD25F90BFC57D98B" authorityName="Rivalier" authorityYear="1970" box="[751,901,1745,1772]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD65825FD32F908FCDDD98B" box="[760,783,1746,1771]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD65825FCE3F90BFC57D98B" box="[809,901,1745,1771]" italics="true" pageId="3" pageNumber="527">infimum</emphasis>
|
||
</taxonomicName>
|
||
are uniformly black or black-brown (
|
||
<bibRefCitation id="EFB14B5CFFD65825FAFDF90BFF00D86F" author="Moravec, J." pageId="3" pageNumber="527" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
, fig. 1819); the other two species also possess black femora, tibiae and antennae. In addition, the surface of the pronotal disc of the new species is covered with finer rugae (
|
||
<figureCitation id="131B2A28FFD65825FCF1F8C3FCAFD853" box="[827,893,1817,1843]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="3" pageNumber="527">Fig. 8</figureCitation>
|
||
) than the coarser and more almost subreticulateanastomosing ridges on the pronotum of
|
||
<emphasis id="B954EABFFFD65825FD94F8E4FDBDD837" box="[606,623,1854,1879]" italics="true" pageId="3" pageNumber="527">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD65825FDB7F8E7FCC2D837" authorityName="Rivalier" authorityYear="1970" box="[637,784,1853,1880]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD65825FD4CF8E4FD4FD837" box="[646,669,1854,1879]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD65825FD7EF8E7FCC2D837" box="[692,784,1853,1879]" italics="true" pageId="3" pageNumber="527">infimum</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB14B5CFFD65825FC80F8E4FC3DD837" author="Moravec, J." box="[842,1007,1853,1880]" pageId="3" pageNumber="527" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
, fig. 1621). Moreover, the aedeagus of the new species is more abruptly bent above the middle and wider in the point of the angle (
|
||
<figureCitation id="131B2A28FFD65825FB52F8BBFB09D81C" box="[1176,1243,1889,1916]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="3" pageNumber="527">Fig. 5</figureCitation>
|
||
), while the angle is arcuate in
|
||
<emphasis id="B954EABFFFD65825FEE3F85CFEE8D8FF" box="[297,314,1926,1951]" italics="true" pageId="3" pageNumber="527">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD65825FE83F85FFE0CD8FF" authorityName="Rivalier" authorityYear="1970" box="[329,478,1925,1952]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="527" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD65825FE99F85CFEB8D8FF" box="[339,362,1926,1951]" italics="true" pageId="3" pageNumber="527">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD65825FE48F85FFE0CD8FF" box="[386,478,1925,1951]" italics="true" pageId="3" pageNumber="527">infimum</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFB14B5CFFD65825FDD6F85CFD10D8FF" author="Moravec, J." box="[540,706,1925,1952]" pageId="3" pageNumber="527" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
, fig. 1622); the aedeagi of other two species are very differently shaped. For other differences see “Key to species” above, and the detailed redescriptions and illustrations in the monograph of the genus (
|
||
<bibRefCitation id="EFB14B5CFFD65825FE7FF817FD8CD887" author="Moravec, J." box="[437,606,1997,2023]" pageId="3" pageNumber="527" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFF4DFBABDE7D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
It should be mentioned here that also
|
||
<taxonomicName id="4C204D2EFFD15822FDBFFF4DFBB2DFD1" authority="W. Horn, 1927" authorityName="W. Horn" authorityYear="1927" box="[629,1120,151,177]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="sicardi" subGenus="Microstenocera">
|
||
<emphasis id="B954EABFFFD15822FDBFFF4DFD54DFD0" box="[629,646,151,176]" italics="true" pageId="4" pageNumber="528">P</emphasis>
|
||
. (
|
||
<emphasis id="B954EABFFFD15822FD54FF4DFC83DFD0" box="[670,849,151,176]" italics="true" pageId="4" pageNumber="528">Microstenocera</emphasis>
|
||
)
|
||
<emphasis id="B954EABFFFD15822FCA8FF4DFC62DFD1" box="[866,944,151,177]" italics="true" pageId="4" pageNumber="528">sicardi</emphasis>
|
||
W. Horn, 1927
|
||
</taxonomicName>
|
||
may superficially resemble the new species by its yellow femoral bases; however, the yellow area is much less expanded and apart from the irregular elytral setal vesture (diagnostic for
|
||
<taxonomicName id="4C204D2EFFD15822FD5AFF05FC91DF98" authorityName="Jeannel" authorityYear="1946" box="[656,835,223,248]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="526" phylum="Arthropoda" rank="subGenus" subGenus="Microstenocera">
|
||
<emphasis id="B954EABFFFD15822FD5AFF05FC91DF98" box="[656,835,223,248]" italics="true" pageId="4" pageNumber="528">Microstenocera</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C204D2EFFD15822FC91FF05FB47DF99" authorityName="W. Horn" authorityYear="1927" box="[859,1173,223,249]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="528" phylum="Arthropoda" rank="species" species="sicardi" subGenus="Microstenocera">
|
||
<emphasis id="B954EABFFFD15822FC91FF05FCBEDF98" box="[859,876,223,248]" italics="true" pageId="4" pageNumber="528">P</emphasis>
|
||
. (
|
||
<emphasis id="B954EABFFFD15822FC4EFF05FBE5DF98" box="[900,1079,223,248]" italics="true" pageId="4" pageNumber="528">Microstenocera</emphasis>
|
||
)
|
||
<emphasis id="B954EABFFFD15822FB8DFF05FB47DF99" box="[1095,1173,223,249]" italics="true" pageId="4" pageNumber="528">sicardi</emphasis>
|
||
</taxonomicName>
|
||
differs in a complex of other characters including the very different shape of its aedeagus (see
|
||
<bibRefCitation id="EFB14B5CFFD15822FC08FED9FBBFDE7D" author="Moravec, J." box="[962,1133,259,285]" pageId="4" pageNumber="528" refId="ref5728" refString="Moravec, J. (2007) Tiger beetles of Madagascar 1. A monograph of the genus Pogonostoma (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, 499 pp." type="book" year="2007">Moravec 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6526FFD1582EFF0DFEFCFCCFDE95" lastPageId="8" lastPageNumber="532" pageId="4" pageNumber="528" type="description">
|
||
<paragraph id="8B9F36ADFFD15822FF0DFEFCFEE2DE05" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
<emphasis id="B954EABFFFD15822FF0DFEFCFE86DE20" bold="true" box="[199,340,294,320]" pageId="4" pageNumber="528">Description</emphasis>
|
||
(male
|
||
<typeStatus id="549B880FFFD15822FE6DFEFDFDC2DE22" box="[423,528,295,322]" pageId="4" pageNumber="528" type="holotype">holotype</typeStatus>
|
||
). Body (
|
||
<figureCitation id="131B2A28FFD15822FDB9FEFDFD67DE21" box="[627,693,295,321]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 1</figureCitation>
|
||
) extremely small, length
|
||
<quantity id="4CD89B48FFD15822FC14FEFCFB94DE21" box="[990,1094,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="528" unit="mm" value="5.6">5.60 mm</quantity>
|
||
, width
|
||
<quantity id="4CD89B48FFD15822FB56FEFCFAD6DE21" box="[1180,1284,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="528" unit="mm" value="1.5">1.50 mm</quantity>
|
||
, shiny-black; setae whitish.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFEB5FE67DECD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Head (
|
||
<figureCitation id="131B2A28FFD15822FED8FEB5FE86DEE9" box="[274,340,367,393]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 3</figureCitation>
|
||
) slightly narrower than body, width
|
||
<quantity id="4CD89B48FFD15822FD24FEB5FC87DEE9" box="[750,853,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="528" unit="mm" value="1.2">1.20 mm</quantity>
|
||
; temples obliquely sloped, only indistinctly arcuate, 2 times shorter than eyes.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFE6DFEA3DDAD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">Frons anteriad-prolonged, merging with clypeus in middle and continuously confluent with vertex, surface rather finely, irregularly scabrous-rugulose and granulate to foveolate, much finer on anterior area; supra-anten- nal keels indistinct, merging with coarse surface sculpture. Vertex with indistinct anterior-sublateral impressions, median area moderately convex, occipital impression shallow, median area coarsely, irregularly scabrous-rugulose, passing posteriad into irregularly subreticulate-scabrous sculpture; limited occipital area with several very coarse, transverse-wavy rugae; temporal area irregularly sculptured by short, irregularly wavy ridges; whole surface ap- pears glabrous as covered with indistinct, sparse and extremely short, white microtrichia, which are barely notice- able in dorsal view.</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFD0DFED8DC75" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">Genae coarsely rugose nearly throughout their surface, glabrous or with few inconspicuous, short, white mi- crotrichia.</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFCC5FA45DC59" blockId="4.[151,1437,151,2013]" box="[199,1431,799,825]" pageId="4" pageNumber="528">Clypeus black, convex in middle when merging with frons, finely wrinkled, with sparse, whitish microtrichia.</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFC99FAB7DCA9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Labrum (
|
||
<figureCitation id="131B2A28FFD15822FEFBFC99FEA7DC3D" box="[305,373,835,861]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 2</figureCitation>
|
||
) 4–setose, rather short, length
|
||
<quantity id="4CD89B48FFD15822FD1FFC99FCEFDC3D" box="[725,829,835,861]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="528" unit="mm" value="0.32">0.32 mm</quantity>
|
||
, width
|
||
<quantity id="4CD89B48FFD15822FC58FC99FC28DC3D" box="[914,1018,835,861]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="528" unit="mm" value="0.6">0.60 mm</quantity>
|
||
, basolateral margins arcuate towards right-angled lateral indentations, then obliquely narrowed anteriad towards rather distinct, right-angled anterolateral teeth, anterior margin shallowly emarginate between two blunt, yet well developed anterior teeth; surface blackbrown, with moderate basodiscal convexity, glabrous except for a few, barely noticeable whitish microtrichia.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFC09FDF8DB71" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Maxillae. Galea black (except for whitish apical orifice of penultimate galeomere); lacinia black with brown- ish-testaceous setae,
|
||
<quantity id="4CD89B48FFD15822FE4BFC2DFE37DB71" box="[385,485,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="528" unit="mm" value="0.12">0.12 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFBC1FD50DBA5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Palpi (
|
||
<figureCitation id="131B2A28FFD15822FEDBFBC1FE84DB55" box="[273,342,1051,1077]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 3</figureCitation>
|
||
). Maxillary palpi with longest palpomere metallic black (only short basal palpomere testaceous), penultimate and terminal palpomere brown with faint reddish tinge; setae whitish; penultimate (longest) palpomeres of labial palpi notably long, their glabrous side black, while their uneven setose side is pale brownish with ivorytestaceous tinge and ivory-white apices; long setae pure white but irregularly interspaced with darker setae; terminal palpomeres brownish with mahogany tinge.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFB15FD66DA35" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Mandibles (
|
||
<figureCitation id="131B2A28FFD15822FE86FB15FE5CDB89" box="[332,398,1231,1257]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 3</figureCitation>
|
||
) metallic black with mahogany-reddish teeth; with moderately arcuate margins, subsymmetrical, with terminal tooth in left mandible notably shorter than that in right mandible; second tooth of left mandible shorter than third one, while inner teeth of right mandible of approximately equal size; distinct basolateral portions arcuate, with very sparse indistinct microtrichia.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFA85FCC5DA85" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Antennae (
|
||
<figureCitation id="131B2A28FFD15822FE89FA85FE14DA19" box="[323,454,1375,1401]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Figs 1, 3–4</figureCitation>
|
||
) shorter than body, reaching anteapical elytral angles (when completed), notably pale, scape black dorsally, while ventrally and apically yellow-testaceous; antennomeres 2–4 ivory to yellow-testaceous with darkened apices, antennomeres 5–6 with ivory base, then gradually testaceous-darkened towards brownish apex, remaining antennomeres pale brownish-testaceous.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15822FF0DFA35FA43D82D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="528">
|
||
Thorax. Pronotum (
|
||
<figureCitation id="131B2A28FFD15822FE63FA35FE3CD969" box="[425,494,1519,1545]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="4" pageNumber="528">Fig. 8</figureCitation>
|
||
) elongate, length
|
||
<quantity id="4CD89B48FFD15822FD75FA34FCFAD969" box="[703,808,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="528" unit="mm" value="1.35">1.35 mm</quantity>
|
||
, width
|
||
<quantity id="4CD89B48FFD15822FCB5FA34FC3AD969" box="[895,1000,1518,1545]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="528" unit="mm" value="0.85">0.85 mm</quantity>
|
||
; anterior lobe narrower then posterior one, irregularly rugulose and with coarse, irregularly transverse ridges; disc oblong, lateral margins only moderately arcuate; notopleural sutures invisible from above; sculpture of discal surface consisting of coarse but rather dense transverse-wavy ridges which are irregularly anastomosing on anterior area, while more continuous, wavy and coarser on median area of posterior discal half, and more irregular and commonly anastomosing on lateral areas; rugae passing over notopleural sutures on proepisterna; median line indistinct, partly merging with surface sculpture; posterior lobe rather finely and irregularly transverse rugulose, its posterior half ochre-testaceous; whole pronotal surface appears glabrous; proepisterna covered with coarse but rather dense ridges (passing from lateral discal areas over notopleural sutures), glabrous; prosternum, mesosternum and metasternum rather sparsely covered with long and short, hairlike setae; mesepisterna with uneven surface and sparse microtrichia; metepisterna nearly glabrous.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFD15823FF0DF88DFB8EDF99" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="529" pageId="4" pageNumber="528">
|
||
Elytra elongate,
|
||
<quantity id="4CD89B48FFD15822FE4BF88DFE34D811" box="[385,486,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="528" unit="mm" value="3.3">3.30 mm</quantity>
|
||
long, humeri arcuate, then obliquely sloped towards posterior pronotal suture; outer margins subparallel with slightly arched dilatation above the middle and slightly dilated towards rather distinct anteapical convexity; apex arcuate and shallowly emarginated towards small sutural spine; surface moderately convex, basodiscal convexity and discal impression rather distinct, coarsely punctate throughout, punctures notably large and deep with shiny intervals, often anastomosing with declined lateral intervals, often forming irregularly elongate caverns; setal vesture regular, microsetae confined to punctures, very short and therefore barely visible and easily abraded to the measure that particularly posterior elytral third appears glabrous.
|
||
</paragraph>
|
||
<caption id="DF5F6625FFD05823FF5DF87DFADDD883" ID-DOI="http://doi.org/10.5281/zenodo.5778845" ID-Zenodo-Dep="5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="5" pageNumber="529" startId="5.[151,264,1959,1983]" targetBox="[161,1431,278,1929]" targetPageId="5">
|
||
<paragraph id="8B9F36ADFFD05823FF5DF87DFADDD883" blockId="5.[151,1436,1959,2019]" pageId="5" pageNumber="529">
|
||
<emphasis id="B954EABFFFD05823FF5DF87DFEEED8DF" bold="true" box="[151,316,1959,1983]" pageId="5" pageNumber="529">FIGURES 1–8.</emphasis>
|
||
<taxonomicName id="4C204D2EFFD05823FE8EF872FCC4D8DF" authority="Moravec & Trýzna, 2021" authorityName="Moravec & Trýzna" authorityYear="2021" box="[324,790,1959,1983]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="529" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
|
||
<emphasis id="B954EABFFFD05823FE8EF872FE1DD8DF" box="[324,463,1960,1983]" italics="true" pageId="5" pageNumber="529">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B954EABFFFD05823FE15F872FDA8D8DF" box="[479,634,1960,1983]" italics="true" pageId="5" pageNumber="529">Microgeniatum</emphasis>
|
||
)
|
||
<emphasis id="B954EABFFFD05823FD41F87DFCC4D8DF" box="[651,790,1959,1983]" italics="true" pageId="5" pageNumber="529">signifemorale</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EABFFFD05823FCD4F87DFCA2D8DF" bold="true" box="[798,880,1959,1983]" pageId="5" pageNumber="529">
|
||
<taxonomicNameLabel id="A26757C4FFD05823FCD4F87DFCA2D8DF" box="[798,880,1959,1983]" pageId="5" pageNumber="529" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6625FFD35820FF5DF891FE71D8E7" ID-DOI="http://doi.org/10.5281/zenodo.5778847" ID-Zenodo-Dep="5778847" httpUri="https://zenodo.org/record/5778847/files/figure.png" pageId="6" pageNumber="530" startId="6.[151,250,1867,1891]" targetBox="[151,1435,181,1844]" targetPageId="6">
|
||
<paragraph id="8B9F36ADFFD35820FF5DF891FE71D8E7" blockId="6.[151,1436,1867,1928]" pageId="6" pageNumber="530">
|
||
<emphasis id="B954EABFFFD35820FF5DF891FEEBD804" bold="true" box="[151,313,1867,1892]" pageId="6" pageNumber="530">FIGURE 9–10.</emphasis>
|
||
<taxonomicName id="4C204D2EFFD35820FEF5F896FCDFD804" authority="Moravec & Trýzna, 2021" authorityName="Moravec & Trýzna" authorityYear="2021" box="[319,781,1868,1892]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
|
||
<emphasis id="B954EABFFFD35820FEF5F896FE18D803" box="[319,458,1868,1891]" italics="true" pageId="6" pageNumber="530">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B954EABFFFD35820FE13F896FDA6D803" box="[473,628,1868,1891]" italics="true" pageId="6" pageNumber="530">Microgeniatum</emphasis>
|
||
)
|
||
<emphasis id="B954EABFFFD35820FD48F896FCDFD804" box="[642,781,1868,1892]" italics="true" pageId="6" pageNumber="530">signifemorale</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954EABFFFD35820FCDEF896FCB6D804" bold="true" box="[788,868,1868,1892]" pageId="6" pageNumber="530">
|
||
<taxonomicNameLabel id="A26757C4FFD35820FCDEF896FCB6D804" box="[788,868,1868,1892]" pageId="6" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Ankarafantsika National Park. 9—collecting site; 10— habitat of the new species.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F6625FFD25821FF5DF842FBB0D8B4" ID-DOI="http://doi.org/10.5281/zenodo.5778849" ID-Zenodo-Dep="5778849" httpUri="https://zenodo.org/record/5778849/files/figure.png" pageId="7" pageNumber="531" startId="7.[151,264,1944,1968]" targetBox="[233,1354,193,1908]" targetPageId="7">
|
||
<paragraph id="8B9F36ADFFD25821FF5DF842FBB0D8B4" blockId="7.[151,1436,1944,2004]" pageId="7" pageNumber="531">
|
||
<emphasis id="B954EABFFFD25821FF5DF842FE81D8D0" bold="true" box="[151,339,1944,1968]" pageId="7" pageNumber="531">FIGURES 11–14.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C204D2EFFD25821FE3FF843FCE1D8D0" authority="" box="[501,819,1944,1968]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="531" phylum="Arthropoda" rank="subGenus" subGenus="Microgeniatum">
|
||
<emphasis id="B954EABFFFD25821FE3FF843FD52D8D0" box="[501,640,1945,1968]" italics="true" pageId="7" pageNumber="531">Pogonostoma</emphasis>
|
||
(
|
||
<emphasis id="B954EABFFFD25821FD45F843FCF8D8D0" box="[655,810,1945,1968]" italics="true" pageId="7" pageNumber="531">Microgeniatum</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
in Madagascar. 11—
|
||
<emphasis id="B954EABFFFD25821FBC1F843FBC8D8D0" box="[1035,1050,1945,1968]" italics="true" pageId="7" pageNumber="531">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD25821FBEDF842FA8DD8D0" authority="W. Horn" authorityName="W. Horn" authorityYear="1934" box="[1063,1375,1944,1968]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="531" phylum="Arthropoda" rank="species" species="pseudominimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD25821FBFAF843FB97D8D0" box="[1072,1093,1945,1968]" italics="true" pageId="7" pageNumber="531">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD25821FB90F842FAD2D8D0" box="[1114,1280,1944,1968]" italics="true" pageId="7" pageNumber="531">pseudominimum</emphasis>
|
||
W. Horn
|
||
</taxonomicName>
|
||
; 12—
|
||
<emphasis id="B954EABFFFD25821FF5DF867FF74D8B4" box="[151,166,1981,2004]" italics="true" pageId="7" pageNumber="531">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD25821FF79F866FE46D8B4" authority="Rivalier" authorityName="Rivalier" authorityYear="1970" box="[179,404,1980,2004]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="531" phylum="Arthropoda" rank="species" species="infimum" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD25821FF76F867FF03D8B4" box="[188,209,1981,2004]" italics="true" pageId="7" pageNumber="531">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD25821FF2CF866FEEBD8B4" box="[230,313,1980,2004]" italics="true" pageId="7" pageNumber="531">infimum</emphasis>
|
||
Rivalier
|
||
</taxonomicName>
|
||
; 13—
|
||
<emphasis id="B954EABFFFD25821FE1BF867FE32D8B4" box="[465,480,1981,2004]" italics="true" pageId="7" pageNumber="531">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD25821FE27F866FD26D8B4" authority="Jeannel" authorityName="Jeannel" authorityYear="1946" box="[493,756,1980,2004]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="531" phylum="Arthropoda" rank="species" species="flavopalpale" subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD25821FE3CF867FDD9D8B4" box="[502,523,1981,2004]" italics="true" pageId="7" pageNumber="531">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD25821FDEAF866FD4DD8B4" box="[544,671,1980,2004]" italics="true" pageId="7" pageNumber="531">flavopalpale</emphasis>
|
||
Jeannel
|
||
</taxonomicName>
|
||
; 14—
|
||
<emphasis id="B954EABFFFD25821FCFBF867FC92D8B4" box="[817,832,1981,2004]" italics="true" pageId="7" pageNumber="531">P</emphasis>
|
||
.
|
||
<taxonomicName id="4C204D2EFFD25821FC87F866FBD9D8B4" authority="Moravec & Trýzna, 2021" authorityName="Moravec & Trýzna" authorityYear="2021" box="[845,1035,1980,2004]" class="Insecta" family="Carabidae" genus="Pogonostoma" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="531" phylum="Arthropoda" rank="species" species="signifemorale" status="sp. nov." subGenus="Microgeniatum">
|
||
(
|
||
<emphasis id="B954EABFFFD25821FC9FF867FCB8D8B4" box="[853,874,1981,2004]" italics="true" pageId="7" pageNumber="531">M</emphasis>
|
||
.)
|
||
<emphasis id="B954EABFFFD25821FC4AF866FBD9D8B4" box="[896,1035,1980,2004]" italics="true" pageId="7" pageNumber="531">signifemorale</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26757C4FFD25821FBDBF866FBB0D8B4" box="[1041,1122,1980,2004]" pageId="7" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F36ADFFDD582EFF0DFF4DFDAEDFB5" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="532">Abdomen. Ventrites black, except for last ventrite and pleurite which are testaceous, their surface rather sparse- ly covered with short, whitish microtrichia.</paragraph>
|
||
<paragraph id="8B9F36ADFFDD582EFF0DFF05FB7ADE05" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="532">
|
||
Legs (
|
||
<figureCitation id="131B2A28FFDD582EFEC6FF05FE9EDF99" box="[268,332,223,249]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="8" pageNumber="532">Fig. 1</figureCitation>
|
||
). Coxae black, trochanters testaceous; profemora voluminous, black, except for yellow base; meso- and metafemora with basal third yellow, remaining portion black and subclavate-dilated towards constricted apex; basal quarter or third of all tibiae yellow; tarsi brownish to black-brown, claws testaceous; setal vesture of legs inconspicuous: femora glabrous, tibiae and tarsi covered with usual, rather dense microsetae.
|
||
</paragraph>
|
||
<paragraph id="8B9F36ADFFDD582EFF0DFEB5FCCFDE95" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="532">
|
||
Aedeagus (
|
||
<figureCitation id="131B2A28FFDD582EFE82FEB5FE5FDEE9" box="[328,397,366,393]" captionStart="FIGURES 1–8" captionStartId="5.[151,264,1959,1983]" captionTargetBox="[161,1431,278,1929]" captionTargetId="figure-72@5.[151,1435,270,1933]" captionTargetPageId="5" captionText="FIGURES 1–8. Pogonostoma (Microgeniatum) signifemorale sp. nov., male holotype from type locality. 1—habitus; 2—la- brum; 3—head; 4—antennomeres 1–6; 5—aedeagus; 6—elytral apices; 7—elytron; 8—pronotum. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5778845" httpUri="https://zenodo.org/record/5778845/files/figure.png" pageId="8" pageNumber="532">Fig. 5</figureCitation>
|
||
) abruptly boomerang-bent above middle, length
|
||
<quantity id="4CD89B48FFDD582EFC0FFEB5FBFCDEE9" box="[965,1070,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="532" unit="mm" value="1.1">1.10 mm</quantity>
|
||
, width
|
||
<quantity id="4CD89B48FFDD582EFB4DFEB5FB22DEE9" box="[1159,1264,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="532" unit="mm" value="0.2">0.20 mm</quantity>
|
||
, apical portion rather wide, notably dilated in the dorsal angle, conical-constricted towards blunt apex which is ventrally arcuate, dorsally very slightly emarginate, thus slightly directed dorsad; in its dorsal (and ventral) aspect the apical portion appears gradually conical-constricted towards blunt apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6526FFDD582EFF0DFE24FACCDC75" pageId="8" pageNumber="532" type="distribution">
|
||
<paragraph id="8B9F36ADFFDD582EFF0DFE24FACCDC75" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="532">
|
||
<emphasis id="B954EABFFFDD582EFF0DFE24FE38DD78" bold="true" box="[199,490,510,537]" pageId="8" pageNumber="532">Distribution and habitat</emphasis>
|
||
(
|
||
<figureCitation id="131B2A28FFDD582EFE32FE25FDB9DD79" box="[504,619,511,537]" captionStart="FIGURE 9–10" captionStartId="6.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURE 9–10. Pogonostoma (Microgeniatum) signifemorale sp. nov., Ankarafantsika National Park. 9—collecting site; 10— habitat of the new species." figureDoi="http://doi.org/10.5281/zenodo.5778847" httpUri="https://zenodo.org/record/5778847/files/figure.png" pageId="8" pageNumber="532">Figs 9–10</figureCitation>
|
||
,
|
||
<figureCitation id="131B2A28FFDD582EFDBCFE25FD47DD79" box="[630,661,511,537]" captionStart="FIGURES 11–14" captionStartId="7.[151,264,1944,1968]" captionTargetBox="[233,1354,193,1908]" captionTargetId="figure-29@7.[218,1369,181,1920]" captionTargetPageId="7" captionText="FIGURES 11–14. Distribution of Pogonostoma (Microgeniatum) in Madagascar. 11—P. (M.) pseudominimum W. Horn; 12— P. (M.) infimum Rivalier; 13—P. (M.) flavopalpale Jeannel; 14—P. (M.) signifemorale sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5778849" httpUri="https://zenodo.org/record/5778849/files/figure.png" pageId="8" pageNumber="532">14</figureCitation>
|
||
). Known only from the male
|
||
<typeStatus id="549B880FFFDD582EFC14FE25FB92DD79" box="[990,1088,511,537]" pageId="8" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
caught by the second author in the dense, dry deciduous forest of the Ankarafantsika National Park (previously known as Ampijoroa Forest Station, see
|
||
<bibRefCitation id="EFB14B5CFFDD582EFF0AFD9DFE93DD01" author="Viette, P." box="[192,321,583,609]" pageId="8" pageNumber="532" refId="ref7255" refString="Viette, P. (1991) Principales localites ou des Insectes ont ete recueillis a Madagascar. Chief field stations where Insects were collected in Madagascar. Faune de Madagascar. Supplement 2. Privately published, Paris, 88 pp." type="book" year="1991">Viette 1991</bibRefCitation>
|
||
) near Marovoay, northwestern
|
||
<collectingCountry id="F337763DFFDD582EFD51FD9DFCF0DD01" box="[667,802,583,609]" name="Madagascar" pageId="8" pageNumber="532">Madagascar</collectingCountry>
|
||
(Ambato Boeni Region, district of
|
||
<collectingRegion id="49E4F84FFFDD582EFB6DFD9DFAFBDD02" box="[1191,1321,583,610]" country="Madagascar" name="Mahajanga" pageId="8" pageNumber="532">Mahajanga</collectingRegion>
|
||
). The only male was caught along the tracking path “Tour du Lac” in a margin between primary and secondary forest when running along a surface of an almost horizontal fallen twig of about
|
||
<quantity id="4CD89B48FFDD582EFC6AFD55FC34DDC9" box="[928,998,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="532" unit="cm" value="10.0">10 cm</quantity>
|
||
in diameter (
|
||
<figureCitation id="131B2A28FFDD582EFBB4FD55FB16DDC9" box="[1150,1220,655,681]" captionStart="FIGURE 9–10" captionStartId="6.[151,250,1867,1891]" captionTargetBox="[151,1435,181,1844]" captionTargetId="figure-17@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURE 9–10. Pogonostoma (Microgeniatum) signifemorale sp. nov., Ankarafantsika National Park. 9—collecting site; 10— habitat of the new species." figureDoi="http://doi.org/10.5281/zenodo.5778847" httpUri="https://zenodo.org/record/5778847/files/figure.png" pageId="8" pageNumber="532">Fig. 9</figureCitation>
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, bottom). No other adult was observed in the area despite an intense search. Unfortunately, due to serious bushfires which have recently afflicted some of the Madagascan protected areas, including the Ankarafantsika National Park (see
|
||
<bibRefCitation id="EFB14B5CFFDD582EFACDFD0DFA4ADD91" author="Bezain, L." box="[1287,1432,727,753]" pageId="8" pageNumber="532" refId="ref5287" refString="Bezain, L. (2021) Madagascar: un incendie ravage plus de 100 hectares du parc national d'Ankarafantsika. RFI, 15.9. 2021. Available from: https: // www. rfi. fr / fr / afrique / 20210915 - madagascar-un-incendie-ravage-plus-de- 100 - hectares-du-parc-na- tional-d-ankarafantsika (accessed 20 September 2021)" type="url" year="2021">Bezain 2021</bibRefCitation>
|
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,
|
||
<bibRefCitation id="EFB14B5CFFDD582EFF5DFD21FE85DC75" author="Vyawahare, M." box="[151,343,763,789]" pageId="8" pageNumber="532" refId="ref7298" refString="Vyawahare, M. (2020) In Madagascar's dry forests, COVID- 19 sparks an intense, early fire season. Mongabay, News & Inspira- tion From Nature's Frontline, Mongabay series: Conservation in Madagascar, 25 June 2020. Available from: https: // news. mongabay. com / 2020 / 06 / in-madagascars-dry-forests-covid- 19 - sparks-an-intense-early-fire-season / (accessed 20 September 2021)" type="url" year="2020">Vyawahare 2020</bibRefCitation>
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), the chance of finding other adults of the new species appears even more improbable.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A6526FFDD582EFF0DFCC5FEF7DC3D" pageId="8" pageNumber="532" type="etymology">
|
||
<paragraph id="8B9F36ADFFDD582EFF0DFCC5FEF7DC3D" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="532">
|
||
<emphasis id="B954EABFFFDD582EFF0DFCC5FE9ADC59" bold="true" box="[199,328,799,825]" pageId="8" pageNumber="532">Etymology</emphasis>
|
||
. Derived from Latin
|
||
<emphasis id="B954EABFFFDD582EFDF3FCC5FDA4DC58" box="[569,630,799,824]" italics="true" pageId="8" pageNumber="532">signo</emphasis>
|
||
[
|
||
<emphasis id="B954EABFFFDD582EFD4DFCC5FD0ADC58" box="[647,728,799,824]" italics="true" pageId="8" pageNumber="532">signum</emphasis>
|
||
] = marked, referring to the markedly bicoloured femora of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |