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<mods:title id="956CC212F424CC155740ADF387C86A75">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<paragraph id="8B9F36B6FFC44B27B384A0B8101CFF65" blockId="62.[914,1350,189,250]" pageId="62" pageNumber="287">
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Key to species and species groups of
<taxonomicName id="4C204D35FFC44B27B3E0A0DA1141FF65" ID-CoL="8VS2M" authorityName="Stephens" authorityYear="1829" box="[1014,1115,223,250]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B954EAA4FFC44B27B3E0A0DA1141FF65" bold="true" box="[1014,1115,223,250]" italics="true" pageId="62" pageNumber="287">Quedius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFC44B27B470A0E5101CFF65" box="[1126,1286,224,250]" pageId="62" pageNumber="287" sensu="stricto">sensu stricto</taxonomicNameLabel>
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<keyStep id="30D42E13FFC44B27B32AA1141086FE57" pageId="62" pageNumber="287">
<paragraph id="8B9F36B6FFC44B27B32AA1141086FE17" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
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1 Presence of 4 or 6 additional setiferous punctures between anterior frontal punctures (
<figureCitation id="131B2A33FFC44B27B50CA134108AFED7" box="[1306,1424,305,328]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="62" pageNumber="287">Fig. K1A</figureCitation>
). Palearctic, two species introduced to the Nearctic.
<emphasis id="B954EAA4FFC44B27B594A155112BFE17" bold="true" pageId="62" pageNumber="287">
<taxonomicName id="4C204D35FFC44B27B594A15516FEFE17" ID-CoL="4R2YF" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFC44B27B594A15516FEFE17" bold="true" italics="true" pageId="62" pageNumber="287">Q. fuliginosus</emphasis>
</taxonomicName>
-group
</emphasis>
.................................................. 5
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC44B27B32AA1941086FE57" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA1941086FE57" pageId="62" pageNumber="287">
Presence of 2 or no additional setiferous punctures between anterior frontal punctures (
<figureCitation id="131B2A33FFC44B27B4F0A1B41054FE57" box="[1254,1358,433,456]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen &amp; Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="62" pageNumber="287">Fig. K1B</figureCitation>
). ..... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC44B27B32AA1D41086FDF7" pageId="62" pageNumber="287">
<paragraph id="8B9F36B6FFC44B27B32AA1D41087FDB7" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA1D41087FDB7" pageId="62" pageNumber="287">
2 Presence of 2 additional setiferous punctures between anterior frontal punctures (
<figureCitation id="131B2A33FFC44B27B4BFA1F41004FD97" box="[1193,1310,497,520]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="62" pageNumber="287">Fig. K2A</figureCitation>
). Nearctic.
<emphasis id="B954EAA4FFC44B27B37FA215116DFDB7" bold="true" box="[873,1143,528,552]" pageId="62" pageNumber="287">
<taxonomicName id="4C204D35FFC44B27B37FA2151130FDB7" ID-CoL="4R2YW" authorityName="Smetana" authorityYear="1965" box="[873,1066,528,552]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="molochinoides">
<emphasis id="B954EAA4FFC44B27B37FA2151130FDB7" bold="true" box="[873,1066,528,552]" italics="true" pageId="62" pageNumber="287">Q. molochinoides</emphasis>
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-group
</emphasis>
..................................... 15
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</paragraph>
<paragraph id="8B9F36B6FFC44B27B32AA2341086FDF7" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA2341086FDF7" pageId="62" pageNumber="287">
No additional setiferous punctures between anterior frontal punctures (
<figureCitation id="131B2A33FFC44B27B43BA254118FFDF7" box="[1069,1173,593,616]" captionStart="Fig" captionStartId="2.[151,182,1953,1972]" captionTargetBox="[199,1388,189,1937]" captionTargetId="figure-13@2.[199,1388,189,1937]" captionTargetPageId="2" captionText="Fig. 2. Habitats of Quedius s. str. A Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti)." figureDoi="http://doi.org/10.5281/zenodo.7399706" httpUri="https://zenodo.org/record/7399706/files/figure.png" pageId="62" pageNumber="287">Fig. K2B</figureCitation>
). ................................. 3
</keyLead>
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</keyStep>
<keyStep id="30D42E13FFC44B27B32AA2741086FC97" pageId="62" pageNumber="287">
<paragraph id="8B9F36B6FFC44B27B32AA2741086FD57" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA2741086FD57" pageId="62" pageNumber="287">
3 Additional 1 to 3 punctures in the posterior half of the sublateral row (
<figureCitation id="131B2A33FFC44B27B419A2941162FD37" box="[1039,1144,657,680]" captionStart="Fig" captionStartId="11.[151,182,2008,2027]" captionTargetBox="[151,1437,191,1879]" captionTargetId="figure-13@11.[151,1437,191,1968]" captionTargetPageId="11" captionText="Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399708" httpUri="https://zenodo.org/record/7399708/files/figure.png" pageId="62" pageNumber="287">Fig. K3A</figureCitation>
). W Palearctic.
<emphasis id="B954EAA4FFC44B27B509A29516B7FD57" bold="true" pageId="62" pageNumber="287">
<taxonomicName id="4C204D35FFC44B27B509A2951089FD37" ID-CoL="4R2YZ" authorityName="Lucas" authorityYear="1846" box="[1311,1427,656,680]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFC44B27B509A2951089FD37" bold="true" box="[1311,1427,656,680]" italics="true" pageId="62" pageNumber="287">Q. pallipes</emphasis>
</taxonomicName>
- group
</emphasis>
...................................................................... 8
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC44B27B32AA2D41086FC97" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA2D41086FC97" pageId="62" pageNumber="287">
No additional puncture in the posterior half of the sublateral row (
<figureCitation id="131B2A33FFC44B27B419A2F4116DFC97" box="[1039,1143,753,776]" captionStart="Fig" captionStartId="11.[151,182,2008,2027]" captionTargetBox="[151,1437,191,1879]" captionTargetId="figure-13@11.[151,1437,191,1968]" captionTargetPageId="11" captionText="Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP &lt;0.90, UFB &lt;95 and SH-aLRT &lt;80." figureDoi="http://doi.org/10.5281/zenodo.7399708" httpUri="https://zenodo.org/record/7399708/files/figure.png" pageId="62" pageNumber="287">Fig. K3B</figureCitation>
). ..................................... 4
</keyLead>
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<keyStep id="30D42E13FFC44B27B32AA3141087FCF7" pageId="62" pageNumber="287">
<paragraph id="8B9F36B6FFC44B27B32AA3141086FCB7" blockId="62.[828,1437,273,1032]" box="[828,1436,784,808]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA3141086FCB7" box="[828,1436,784,808]" pageId="62" pageNumber="287">
4 Palearctic.
<emphasis id="B954EAA4FFC44B27B3F6A31511D4FCB7" bold="true" box="[992,1230,784,808]" pageId="62" pageNumber="287">
<taxonomicName id="4C204D35FFC44B27B3F6A315119BFCB7" ID-CoL="4R2YX" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[992,1153,784,808]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFC44B27B3F6A315119BFCB7" bold="true" box="[992,1153,784,808]" italics="true" pageId="62" pageNumber="287">Q. molochinus</emphasis>
</taxonomicName>
-group
</emphasis>
.......................... 9
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</paragraph>
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<keyLead id="30D19583FFC44B27B32AA3341087FCF7" pageId="62" pageNumber="287">
Nearctic.
<emphasis id="B954EAA4FFC44B27B3CFA33511B7FCD7" bold="true" box="[985,1197,816,840]" pageId="62" pageNumber="287">
<taxonomicName id="4C204D35FFC44B27B3CFA335117AFCD7" ID-CoL="4R2YN" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[985,1120,816,840]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFC44B27B3CFA335117AFCD7" bold="true" box="[985,1120,816,840]" italics="true" pageId="62" pageNumber="287">Q. laticollis</emphasis>
</taxonomicName>
-group
</emphasis>
and
<taxonomicName id="4C204D35FFC44B27B4E4A3351086FCD7" ID-CoL="4R2YX" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[1266,1436,816,840]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFC44B27B4E4A3351086FCD7" bold="true" box="[1266,1436,816,840]" italics="true" pageId="62" pageNumber="287">Q. molochinus</emphasis>
</taxonomicName>
introduced from Palearctic. .................................. 19
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</paragraph>
</keyStep>
<keyStep id="30D42E13FFC44B27B32AA3741087FB97" pageId="62" pageNumber="287">
<paragraph id="8B9F36B6FFC44B27B32AA3741086FC57" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA3741086FC57" pageId="62" pageNumber="287">
5 Scutellum punctured and pubescent (
<figureCitation id="131B2A33FFC44B27B517A3741075FC17" box="[1281,1391,881,904]" captionStart="Fig" captionStartId="15.[151,182,1229,1248]" captionTargetBox="[166,1431,196,1208]" captionTargetId="figure-13@15.[151,1436,186,1218]" captionTargetPageId="15" captionText="Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal)." figureDoi="http://doi.org/10.5281/zenodo.7399714" httpUri="https://zenodo.org/record/7399714/files/figure.png" pageId="62" pageNumber="287">Fig. K5A</figureCitation>
). W Palearctic, usually preferring warmer drier habitats. .................................
<taxonomicName id="4C204D35FFC44B27B45BA3B51086FC57" ID-CoL="4R2YR" authority="Brulle, 1832" authorityName="Brulle" authorityYear="1832" box="[1101,1436,944,968]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="levicollis">
<emphasis id="B954EAA4FFC44B27B45BA3B5100AFC57" bold="true" box="[1101,1296,944,968]" italics="true" pageId="62" pageNumber="287">Quedius levicollis</emphasis>
<bibRefCitation id="EFB14B47FFC44B27B501A3B41086FC57" author="BRULLE G. A." box="[1303,1436,945,968]" pageId="62" pageNumber="287" pagination="289 - 400" refId="ref80797" refString="BRULLE G. A. 1832 - 1833: Expedition scientifique de Moree. Section des Sciences Physiques. Tome III. 1. re Partie. Zoologie. Deuxieme Section. Des animales articules. F. G. Levrault, Paris, 400 pp. [Pp. 1 - 288 published in 1832, pp. 289 - 400 in 1833.]" type="book chapter" year="1832">Brullé, 1832</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC44B27B32AA3D41087FB97" blockId="62.[828,1437,273,1032]" pageId="62" pageNumber="287">
<keyLead id="30D19583FFC44B27B32AA3D41087FB97" pageId="62" pageNumber="287">
Scutellum unpunctured and glabrous (
<figureCitation id="131B2A33FFC44B27B4E9A3D41072FC77" box="[1279,1384,977,1000]" captionStart="Fig" captionStartId="15.[151,182,1229,1248]" captionTargetBox="[166,1431,196,1208]" captionTargetId="figure-13@15.[151,1436,186,1218]" captionTargetPageId="15" captionText="Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal)." figureDoi="http://doi.org/10.5281/zenodo.7399714" httpUri="https://zenodo.org/record/7399714/files/figure.png" pageId="62" pageNumber="287">Fig. K5B</figureCitation>
). ......................................................................................6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A0BB17EDFDAA" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A0BB17EDFECA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A0BB17EDFECA" pageId="63" pageNumber="288">
6 Head with 4 additional punctures between posterior frontal punctures, but without additional puncture between anterior and posterior frontal punctures (
<figureCitation id="131B2A33FFC54B26B2DBA0FB15E6FEAA" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A general scheme; B Q. curtipennis Bernhauer, 1908, C Q. fuliginosus (Gravenhorst, 1802); D Q. levicollis Brullé, 1832; E Q. afrofuliginosus Gusarov, 1991; F Q. molochinus- and Q. laticollis-groups; G Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="63" pageNumber="288">Fig. K6A</figureCitation>
). Median lobe without apical ridge. Paleartic. Introduced to multiple Nearctic localities. ............. 7
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A15B17EDFDAA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A15B17EDFDAA" pageId="63" pageNumber="288">
Presence of 4 additional punctures between posterior frontal punctures and one additional puncture on either side between anterior and posterior frontal punctures (
<figureCitation id="131B2A33FFC54B26B12DA1BB14B9FE4A" box="[315,419,446,469]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A general scheme; B Q. curtipennis Bernhauer, 1908, C Q. fuliginosus (Gravenhorst, 1802); D Q. levicollis Brullé, 1832; E Q. afrofuliginosus Gusarov, 1991; F Q. molochinus- and Q. laticollis-groups; G Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="63" pageNumber="288">Fig. K6B</figureCitation>
). Median lobe with apical ridge (
<figureCitation id="131B2A33FFC54B26B0DAA1DB1435FE6A" box="[204,303,478,501]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="63" pageNumber="288">Fig. 14A</figureCitation>
). Palearctic, known only from North Africa and the islands of Sardinia and
<collectingCountry id="F3377626FFC54B26B205A1FB174FFD8A" box="[531,597,510,533]" name="France" pageId="63" pageNumber="288">Corse</collectingCountry>
. ...................... ..................
<taxonomicName id="4C204D35FFC54B26B15CA21817EDFDAA" ID-CoL="4R2Y8" authority="Gusarov, 1991" authorityName="Gusarov" authorityYear="1991" box="[330,759,541,565]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="afrofuliginosus">
<emphasis id="B954EAA4FFC54B26B15CA218174CFDAA" bold="true" box="[330,598,541,565]" italics="true" pageId="63" pageNumber="288">Quedius afrofuliginosus</emphasis>
<bibRefCitation id="EFB14B47FFC54B26B24AA21B17EDFDAA" author="GUSAROV V. I." box="[604,759,542,565]" pageId="63" pageNumber="288" pagination="3 - 12" refId="ref83976" refString="GUSAROV V. I. 1991: Novye i maloizvestnye palearkticheskie stafilinidy (Coleoptera: Staphylinidae) [New and little-known Palearctic rove beetles (Coleoptera: Staphylinidae)] Soobshchenie 3: 3 - 12 (in Russian)." type="journal article" year="1991">Gusarov, 1991</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A23B17EDFC4A" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A23B17E2FCAA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A23B17E2FCAA" pageId="63" pageNumber="288">
7 Basal three antennomeres at least slightly darkened. Eyes clearly protruding making the head seem more rectangular wider across (
<figureCitation id="131B2A33FFC54B26B1C3A27B1724FD0A" box="[469,574,638,661]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A Q. afrofuliginosus Gusarov, 1991; B Q. curtipennis Bernhauer, 1908; C Q. fuliginosus (Gravenhorst, 1802); D Q. levicollis Brullé, 1832; E Q. unicolor Kiesenwetter, 1847; F Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="63" pageNumber="288">Fig. K7A</figureCitation>
). Apex of median lobe not laterally flattened (
<figureCitation id="131B2A33FFC54B26B213A29B1775FD2A" box="[517,623,670,693]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="63" pageNumber="288">Fig. 14D</figureCitation>
). Palearctic. Introduced to E
<collectingCountry id="F3377626FFC54B26B16DA2BB14D4FD4A" box="[379,462,702,725]" name="Canada" pageId="63" pageNumber="288">Canada</collectingCountry>
. Prefers wetter humus rich habitats. Paleartic. Introduced to multiple Nearctic localities. ................................................................... .................
<taxonomicName id="4C204D35FFC54B26B12DA31817E2FCAA" ID-CoL="4R2YF" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[315,760,797,821]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="B954EAA4FFC54B26B12DA3181703FCAA" bold="true" box="[315,537,797,821]" italics="true" pageId="63" pageNumber="288">Quedius fuliginosus</emphasis>
(
<bibRefCitation id="EFB14B47FFC54B26B231A31B17EAFCAA" author="GRAVENHORST J. L. C." box="[551,752,798,821]" pageId="63" pageNumber="288" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A33B17EDFC4A" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A33B17EDFC4A" pageId="63" pageNumber="288">
Basal three antennomeres completely pale. Eyes not clearly protruding making the head seem round (
<figureCitation id="131B2A33FFC54B26B0DAA37B142DFC0A" box="[204,311,894,917]" captionStart="Fig" captionStartId="17.[151,182,2023,2042]" captionTargetBox="[180,1386,189,1955]" captionTargetId="figure-13@17.[177,1411,186,1980]" captionTargetPageId="17" captionText="Fig. 7. Habitus of Quedius s. str. A Q. afrofuliginosus Gusarov, 1991; B Q. curtipennis Bernhauer, 1908; C Q. fuliginosus (Gravenhorst, 1802); D Q. levicollis Brullé, 1832; E Q. unicolor Kiesenwetter, 1847; F Q. subunicolor Korge, 1961." figureDoi="http://doi.org/10.5281/zenodo.7399720" httpUri="https://zenodo.org/record/7399720/files/figure.png" pageId="63" pageNumber="288">Fig. K7B</figureCitation>
). Apex of median lobe laterally flattened (
<figureCitation id="131B2A33FFC54B26B0DAA39B1434FC2A" box="[204,302,926,949]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="63" pageNumber="288">Fig. 14C</figureCitation>
). Paleartic. Introduced to multiple Nearctic localities. .....
<taxonomicName id="4C204D35FFC54B26B177A3B817EDFC4A" ID-CoL="78PXW" authority="Bernhauer, 1908" authorityName="Bernhauer" authorityYear="1908" box="[353,759,957,981]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="B954EAA4FFC54B26B177A3B8175AFC4A" bold="true" box="[353,576,957,981]" italics="true" pageId="63" pageNumber="288">Quedius curtipennis</emphasis>
<bibRefCitation id="EFB14B47FFC54B26B251A3BB17EDFC4A" author="BERNHAUER M." box="[583,759,958,981]" pageId="63" pageNumber="288" pagination="320 - 335" refId="ref80365" refString="BERNHAUER M. 1908: Beitrage zur Kenntnis der palaarktischen Staphyliniden-Fauna. I. Munchener Koleopterologische Zeitschrift 3: 320 - 335." type="journal article" year="1908">Bernhauer, 1908</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A3DB17EDFAAA" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A3DB17EDFB2A" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A3DB17EDFB2A" pageId="63" pageNumber="288">
8 Sides of median lobe apex folded with distinct tooth situated at lower part (
<figureCitation id="131B2A33FFC54B26B1DFA3FB172AFB8A" box="[457,560,1022,1045]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="63" pageNumber="288">Fig. 15C</figureCitation>
). Western Europe along Atlantic Coast from
<collectingCountry id="F3377626FFC54B26B1E0A41B1743FBAA" box="[502,601,1054,1077]" name="Denmark" pageId="63" pageNumber="288">Denmark</collectingCountry>
to
<collectingCountry id="F3377626FFC54B26B291A41B17DFFBAA" box="[647,709,1054,1077]" name="Georgia" pageId="63" pageNumber="288">Iberia</collectingCountry>
and inland in Southern
<collectingCountry id="F3377626FFC54B26B1B0A43B14F5FBCA" box="[422,495,1086,1109]" name="France" pageId="63" pageNumber="288">France</collectingCountry>
and Iberian Peninsula, also on the islands of Madeira, Azores, Canaries, and Balearic (
<figureCitation id="131B2A33FFC54B26B13AA47B148CFB0A" box="[300,406,1150,1173]" captionStart="Fig" captionStartId="18.[151,182,2027,2046]" captionTargetBox="[189,1393,195,2007]" captionTargetId="figure-13@18.[174,1413,185,2007]" captionTargetPageId="18" captionText="Fig. 8. Habitus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. meridiocarpathicus Smetana, 1958; C Q. molochinus (Gravenhorst, 1806) (red elytra variant); D Q. molochinus (black elytra variant); E Q. balticus Korge, 1960; F Q. sundukovi Smetana, 2003." figureDoi="http://doi.org/10.5281/zenodo.7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="63" pageNumber="288">Fig. K8A</figureCitation>
).
<emphasis id="B954EAA4FFC54B26B1BFA47B147BFB2A" italics="true" pageId="63" pageNumber="288">.................................................. .......................</emphasis>
<taxonomicName id="4C204D35FFC54B26B171A49817EDFB2A" ID-CoL="4R3NY" authority="Bernhauer, 1898" authorityName="Bernhauer" authorityYear="1898" box="[359,759,1181,1205]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="hispanicus">
<emphasis id="B954EAA4FFC54B26B171A498175BFB2A" bold="true" box="[359,577,1181,1205]" italics="true" pageId="63" pageNumber="288">Quedius hispanicus</emphasis>
<bibRefCitation id="EFB14B47FFC54B26B251A49B17EDFB2A" author="BERNHAUER M." box="[583,759,1182,1205]" pageId="63" pageNumber="288" pagination="338 - 341" refId="ref80314" refString="BERNHAUER M. 1898: Neue Staphyliniden aus Europa. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 48: 338 - 341." type="journal article" year="1898">Bernhauer, 1898</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A4BB17EDFAAA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A4BB17EDFAAA" pageId="63" pageNumber="288">
Sides of median lobe apex folded with distinct tooth situated at medial part (
<figureCitation id="131B2A33FFC54B26B1A8A4DB173AFB6A" box="[446,544,1246,1269]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="63" pageNumber="288">Fig. 15D</figureCitation>
). North Africa,
<collectingCountry id="F3377626FFC54B26B2D1A4DB17E2FB6A" box="[711,760,1246,1269]" name="Italy" pageId="63" pageNumber="288">Italy</collectingCountry>
and islands of
<collectingRegion id="49E4F854FFC54B26B149A4FB14A2FA8A" box="[351,440,1278,1301]" country="Italy" name="Sardegna" pageId="63" pageNumber="288">Sardinia</collectingRegion>
and
<collectingCountry id="F3377626FFC54B26B1FAA4FB1730FA8A" box="[492,554,1278,1301]" name="France" pageId="63" pageNumber="288">Corse</collectingCountry>
(
<figureCitation id="131B2A33FFC54B26B22FA4FB17BBFA8A" box="[569,673,1278,1301]" captionStart="Fig" captionStartId="18.[151,182,2027,2046]" captionTargetBox="[189,1393,195,2007]" captionTargetId="figure-13@18.[174,1413,185,2007]" captionTargetPageId="18" captionText="Fig. 8. Habitus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. meridiocarpathicus Smetana, 1958; C Q. molochinus (Gravenhorst, 1806) (red elytra variant); D Q. molochinus (black elytra variant); E Q. balticus Korge, 1960; F Q. sundukovi Smetana, 2003." figureDoi="http://doi.org/10.5281/zenodo.7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="63" pageNumber="288">Fig. K8B</figureCitation>
). .......... ...................................
<taxonomicName id="4C204D35FFC54B26B1AEA51817EDFAAA" ID-CoL="4R2YZ" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[440,759,1309,1333]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B954EAA4FFC54B26B1AEA5181774FAAA" bold="true" box="[440,622,1309,1333]" italics="true" pageId="63" pageNumber="288">Quedius pallipes</emphasis>
<bibRefCitation id="EFB14B47FFC54B26B263A51B17EDFAAA" author="LUCAS P. H." box="[629,759,1310,1333]" pageId="63" pageNumber="288" refId="ref87139" refString="LUCAS P. H. 1846 - 1849: Histoire naturelle des animaux articules. Deuxieme partie. Insectes. In: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842 publiee par ordre du Gouvernement et avec le concours d'une Commission Academique. Sciences physiques. Zoologie II. A. Bertrand, Paris, 590 pp." type="book" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A53B17E2F9AA" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A53B17EDFA4A" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A53B17EDFA4A" pageId="63" pageNumber="288">
9 Elytra shortened, distinctly shorter than pronotum (
<figureCitation id="131B2A33FFC54B26B0DAA55B142FFAEA" box="[204,309,1374,1397]" captionStart="Fig" captionStartId="19.[151,182,2028,2047]" captionTargetBox="[203,1360,198,1958]" captionTargetId="figure-13@19.[183,1405,186,1971]" captionTargetPageId="19" captionText="Fig. 9. Habitus of Quedius s. str. A Q. hispanicus Bernhauer, 1898; B Q. hispanicus (pale sutures variant); C Q. pallipes Lucas, 1846; D Q. laticollis (Gravenhorst, 1802) (black elytra variant); E Q. laticollis (red elytra variant); F Q. strenuus Casey, 1915." figureDoi="http://doi.org/10.5281/zenodo.7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="63" pageNumber="288">Fig. K9A</figureCitation>
), obviously brachypterous species without whitish apical seam on abdominal tergite VII (
<figureCitation id="131B2A33FFC54B26B2DBA57B15EBFA2A" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A Q. afrofuliginosus Gusarov, 1991; B Q. levicollis Brullé, 1832; C Q. curtipennis Bernhauer, 1908; D Q. fuliginosus (Gravenhorst, 1802). I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="63" pageNumber="288">Fig. 11F</figureCitation>
). East Palearctic. ................................................. ..........................
<taxonomicName id="4C204D35FFC54B26B168A5B817EDFA4A" ID-CoL="4R2Z9" authority="Smetana, 2003" authorityName="Smetana" authorityYear="2003" box="[382,759,1469,1493]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="sundukovi">
<emphasis id="B954EAA4FFC54B26B168A5B81748FA4A" bold="true" box="[382,594,1469,1493]" italics="true" pageId="63" pageNumber="288">Quedius sundukovi</emphasis>
<bibRefCitation id="EFB14B47FFC54B26B24FA5BB17EDFA4A" author="SMETANA A." box="[601,759,1470,1493]" pageId="63" pageNumber="288" pagination="189 - 193" refId="ref92269" refString="SMETANA A. 2003: Quedius (Quedius) sundukovi (Coleoptera, Staphylinidae, Staphylinini, Quediina), an interesting new species from the Russian Far East. Elytra 31: 189 - 193." type="journal article" year="2003">Smetana, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A5DB17E2F9AA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A5DB17E2F9AA" pageId="63" pageNumber="288">
Elytra normal, not shortened, about as long as pronotum (
<figureCitation id="131B2A33FFC54B26B0DAA5FB142CF98A" box="[204,310,1534,1557]" captionStart="Fig" captionStartId="19.[151,182,2028,2047]" captionTargetBox="[203,1360,198,1958]" captionTargetId="figure-13@19.[183,1405,186,1971]" captionTargetPageId="19" captionText="Fig. 9. Habitus of Quedius s. str. A Q. hispanicus Bernhauer, 1898; B Q. hispanicus (pale sutures variant); C Q. pallipes Lucas, 1846; D Q. laticollis (Gravenhorst, 1802) (black elytra variant); E Q. laticollis (red elytra variant); F Q. strenuus Casey, 1915." figureDoi="http://doi.org/10.5281/zenodo.7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="63" pageNumber="288">Fig. K9B</figureCitation>
), with whitish apical seam on abdominal tergite VII. ............................................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A63B17E2F90A" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A63B17E2F9EA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A63B17E2F9EA" pageId="63" pageNumber="288">
10 Basal three antennomeres at least slightly darkened (
<figureCitation id="131B2A33FFC54B26B0DAA65B1459F9EA" box="[204,323,1630,1653]" captionStart="Fig" captionStartId="20.[151,182,2030,2049]" captionTargetBox="[202,1343,201,1961]" captionTargetId="figure-13@20.[182,1406,186,1971]" captionTargetPageId="20" captionText="Fig. 10. Habitus of Quedius s. str. A Q. horni Hatch, 1957; B Q. lanei Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. molochinoides Smetana, 1965 (red elytra variant); E Q. molochinoides (black elytra variant); F Q. labradorensis Smetana, 1965." figureDoi="http://doi.org/10.5281/zenodo.7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="63" pageNumber="288">Fig. K10A</figureCitation>
). ......................................................... 11
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A67B17E2F90A" blockId="63.[151,761,190,2069]" box="[151,760,1662,1685]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A67B17E2F90A" box="[151,760,1662,1685]" pageId="63" pageNumber="288">
Basal three antennomeres pale (
<figureCitation id="131B2A33FFC54B26B20EA67B1797F90A" box="[536,653,1662,1685]" captionStart="Fig" captionStartId="20.[151,182,2030,2049]" captionTargetBox="[202,1343,201,1961]" captionTargetId="figure-13@20.[182,1406,186,1971]" captionTargetPageId="20" captionText="Fig. 10. Habitus of Quedius s. str. A Q. horni Hatch, 1957; B Q. lanei Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. molochinoides Smetana, 1965 (red elytra variant); E Q. molochinoides (black elytra variant); F Q. labradorensis Smetana, 1965." figureDoi="http://doi.org/10.5281/zenodo.7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="63" pageNumber="288">Fig. K10B</figureCitation>
). ....... 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B26B081A69B17E2F84A" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B26B081A69B17EDF8CA" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A69B17EDF8CA" pageId="63" pageNumber="288">
11 Elytra most often reddish, very rarely darkened. Appendages only slightly darkened. Tibia, tarsi and apical antennomeres of antennae pale reddish brown (
<figureCitation id="131B2A33FFC54B26B100A6FB14DFF88A" box="[278,453,1790,1813]" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A Q. afrofuliginosus Gusarov, 1991; B Q. levicollis Brullé, 1832; C Q. curtipennis Bernhauer, 1908; D Q. fuliginosus (Gravenhorst, 1802). I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="63" pageNumber="288">Figs K11A, 11E</figureCitation>
). Found near water bodies in Central-Eastern Europe and Middle Asia (Fig. 23). .. ..................................
<taxonomicName id="4C204D35FFC54B26B1A2A73817EDF8CA" ID-CoL="4R2YB" authority="Korge, 1960" authorityName="Korge" authorityYear="1960" box="[436,759,1853,1877]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="288" phylum="Arthropoda" rank="species" species="balticus">
<emphasis id="B954EAA4FFC54B26B1A2A7381776F8CA" bold="true" box="[436,620,1853,1877]" italics="true" pageId="63" pageNumber="288">Quedius balticus</emphasis>
Korge, 1960
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFC54B26B081A75B17E2F84A" blockId="63.[151,761,190,2069]" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B26B081A75B17E2F84A" pageId="63" pageNumber="288">
Elytra always dark. Appendages darkened. Tarsi slightly paler brown to dark brown at most (
<figureCitation id="131B2A33FFC54B26B2DEA77B141FF82A" captionStart="Fig" captionStartId="24.[151,182,1386,1405]" captionTargetBox="[209,1335,203,1358]" captionTargetId="figure-422@24.[193,1395,189,1368]" captionTargetPageId="24" captionText="Fig. 11. Aedeagus ofQuedius s. str. A Q. afrofuliginosus Gusarov, 1991; B Q. levicollis Brullé, 1832; C Q. curtipennis Bernhauer, 1908; D Q. fuliginosus (Gravenhorst, 1802). I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399728" httpUri="https://zenodo.org/record/7399728/files/figure.png" pageId="63" pageNumber="288">Figs K11B</figureCitation>
,
<figureCitation id="131B2A33FFC54B26B108A79B1473F82A" box="[286,361,1950,1973]" captionStart="Fig" captionStartId="20.[151,182,2030,2049]" captionTargetBox="[202,1343,201,1961]" captionTargetId="figure-13@20.[182,1406,186,1971]" captionTargetPageId="20" captionText="Fig. 10. Habitus of Quedius s. str. A Q. horni Hatch, 1957; B Q. lanei Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. molochinoides Smetana, 1965 (red elytra variant); E Q. molochinoides (black elytra variant); F Q. labradorensis Smetana, 1965." figureDoi="http://doi.org/10.5281/zenodo.7399726" httpUri="https://zenodo.org/record/7399726/files/figure.png" pageId="63" pageNumber="288">10EF</figureCitation>
). Found at higher elevations or latitudes (Figs 25, 27). .......................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFC54B59B081A7DB17EDFE4A" lastPageId="64" lastPageNumber="289" pageId="63" pageNumber="288">
<paragraph id="8B9F36B6FFC54B59B081A7DB17EDFE8A" blockId="63.[151,761,190,2069]" lastBlockId="64.[151,760,190,2069]" lastPageId="64" lastPageNumber="289" pageId="63" pageNumber="288">
<keyLead id="30D19583FFC54B59B081A7DB17EDFE8A" lastPageId="64" lastPageNumber="289" pageId="63" pageNumber="288">
12 Head disc with fine isodiametric microsculpture (
<figureCitation id="131B2A33FFC54B26B2DBA7DB1410F78A" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="63" pageNumber="288">Fig. K12A</figureCitation>
). Without C-sclerite of the internal sac (
<figureCitation id="131B2A33FFC54B59B2DBA7FB15ECFF4A" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" lastPageId="64" lastPageNumber="289" pageId="63" pageNumber="288">Fig. 16A</figureCitation>
). Found at alpine sites of Central and Southern Europe (
<figureCitation id="131B2A33FFBA4B59B136A0DB148CFF6A" box="[288,406,222,245]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="64" pageNumber="289">Fig. K12C</figureCitation>
). .................................................. ......................
<taxonomicName id="4C204D35FFBA4B59B175A0F817EDFE8A" ID-CoL="78Q9W" authority="Kiesenwetter, 1847" authorityName="Kiesenwetter" authorityYear="1847" box="[355,759,253,277]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B954EAA4FFBA4B59B175A0F81738FE8A" bold="true" box="[355,546,253,277]" italics="true" pageId="64" pageNumber="289">Quedius unicolor</emphasis>
<bibRefCitation id="EFB14B47FFBA4B59B23FA0FB17EDFE8A" author="KIESENWETTER H. &amp; MARKEL F." box="[553,759,254,277]" pageId="64" pageNumber="289" pagination="73 - 87" refId="ref85769" refString="KIESENWETTER H. ඏඈඇ &amp; MARKEL F. 1847: Eine entomologische Excursion im Riesengebirge im Juli 1846 (Fortsetzung). Entomologische Zeitung (Stettin) 8: 73 - 87." type="journal article" year="1847">Kiesenwetter, 1847</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A11B17EDFE4A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A11B17EDFE4A" pageId="64" pageNumber="289">
Head disc most often with microsculpture of fine transverse waves, rarely isodiametric (
<figureCitation id="131B2A33FFBA4B59B264A13B17F6FECA" box="[626,748,318,341]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="64" pageNumber="289">Fig. K12B</figureCitation>
). With clear C-sclerite of the internal sac (
<figureCitation id="131B2A33FFBA4B59B291A15B17F1FEEA" box="[647,747,350,373]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">Fig. 15C</figureCitation>
). Found disjunct in boreal Scandinavia and
<collectingCountry id="F3377626FFBA4B59B29AA17B17CCFE0A" box="[652,726,382,405]" name="Russia" pageId="64" pageNumber="289">Russia</collectingCountry>
, as well as alpine in
<collectingRegion id="49E4F854FFBA4B59B196A19B14AFFE2A" box="[384,437,414,437]" country="Russia" name="Altay" pageId="64" pageNumber="289">Altai</collectingRegion>
Mountains and Transbaikalia (
<figureCitation id="131B2A33FFBA4B59B0DAA1BB145EFE4A" box="[204,324,446,469]" captionStart="Fig" captionStartId="29.[151,182,2030,2049]" captionTargetBox="[151,1437,1368,2011]" captionTargetId="figure-672@29.[151,1437,1368,2011]" captionTargetPageId="29" captionText="Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange)." figureDoi="http://doi.org/10.5281/zenodo.7399730" httpUri="https://zenodo.org/record/7399730/files/figure.png" pageId="64" pageNumber="289">Fig. K12D</figureCitation>
). .....
<taxonomicName id="4C204D35FFBA4B59B190A1B817EDFE4A" ID-CoL="4R2Z8" authority="Korge, 1961" authorityName="Korge" authorityYear="1961" box="[390,759,445,469]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="subunicolor">
<emphasis id="B954EAA4FFBA4B59B190A1B81776FE4A" bold="true" box="[390,620,445,469]" italics="true" pageId="64" pageNumber="289">Quedius subunicolor</emphasis>
<bibRefCitation id="EFB14B47FFBA4B59B265A1BB17EDFE4A" author="KORGE H." box="[627,759,446,469]" pageId="64" pageNumber="289" pagination="81 - 83" refId="ref86145" refString="KORGE H. 1961: Quedius subunicolor n. sp. aus Nordeuropa (Col. Staphyl.). Mitteilungen der Deutschen Entomologischen Gesellschaft 20: 81 - 83." type="journal article" year="1961">Korge, 1961</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFBA4B59B081A1DB17E2FC8A" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A1DB17E2FDEA" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A1DB17E2FDEA" pageId="64" pageNumber="289">
13 Body brown. Elytra yellowish red. Apex of paramere pointing ventral, away from median lobe in form of a small blunt hook (
<figureCitation id="131B2A33FFBA4B59B1C7A21B1755FDAA" box="[465,591,542,565]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Figs K13B</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B249A21B178BFDAA" box="[607,657,542,565]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">15A</figureCitation>
). Central
<collectingCountry id="F3377626FFBA4B59B0D3A23B143EFDCA" box="[197,292,574,597]" name="Turkey" pageId="64" pageNumber="289">Anatolia</collectingCountry>
, Middle East, Transcauscasus, Central Asia (
<figureCitation id="131B2A33FFBA4B59B0DAA25B145EFDEA" box="[204,324,606,629]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Fig. K13A</figureCitation>
). .....
<taxonomicName id="4C204D35FFBA4B59B197A25817E2FDEA" ID-CoL="4R2ZB" authority="(Menetries, 1832)" baseAuthorityName="Menetries" baseAuthorityYear="1832" box="[385,760,605,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="B954EAA4FFBA4B59B197A2581728FDEA" bold="true" box="[385,562,605,629]" italics="true" pageId="64" pageNumber="289">Quedius vicinus</emphasis>
(
<bibRefCitation id="EFB14B47FFBA4B59B256A25B17EAFDEA" author="MENETRIES E." box="[576,752,606,629]" pageId="64" pageNumber="289" refId="ref87999" refString="MENETRIES E. 1832: Catalogue raisonne des objets de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontieres actuelles de la Perse entrepris par ordre de S. M. L'Empereur. L'Academie Imperiale des Sciences, St. Petersbourg, 271 + xxxii + iv pp." type="book" year="1832">Ménétriés, 1832</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A27B17E2FC8A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A27B17E2FC8A" pageId="64" pageNumber="289">
Body most often black, or at most brownish. Elytra brick or brownish red. Apex of paramere straight, not pointing ventrad, away from median lobe (
<figureCitation id="131B2A33FFBA4B59B2DEA2BB143DFD6A" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Figs K13CD</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B12DA2DB14B6FD6A" box="[315,428,734,757]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">15D, 15F</figureCitation>
). Distribution more northern (
<figureCitation id="131B2A33FFBA4B59B0DAA2FB1444FC8A" box="[204,350,766,789]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="64" pageNumber="289">Figs K14AB</figureCitation>
)...................................................... 14
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</paragraph>
</keyStep>
<keyStep id="30D42E13FFBA4B59B081A31B17E2FB8A" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A31B17EDFC0A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A31B17EDFC0A" pageId="64" pageNumber="289">
14 C-shaped sclerite of internal sac without spine-like basal extension (
<figureCitation id="131B2A33FFBA4B59B191A33B1712FCCA" box="[391,520,830,853]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Figs K13C</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B20DA33B1752FCCA" box="[539,584,830,853]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">15F</figureCitation>
). SE European distribution (
<figureCitation id="131B2A33FFBA4B59B158A35B14DFFCEA" box="[334,453,862,885]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="64" pageNumber="289">Fig. K14A</figureCitation>
). ........................................... ...........
<taxonomicName id="4C204D35FFBA4B59B10DA37817EDFC0A" ID-CoL="4R2YT" authority="Smetana, 1958" authorityName="Smetana A. Solodovnikov" authorityYear="1958" box="[283,759,893,917]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="meridiocarpathicus">
<emphasis id="B954EAA4FFBA4B59B10DA3781748FC0A" bold="true" box="[283,594,893,917]" italics="true" pageId="64" pageNumber="289">Quedius meridiocarpathicus</emphasis>
<bibRefCitation id="EFB14B47FFBA4B59B24FA37B17EDFC0A" author="SMETANA A." box="[601,759,894,917]" pageId="64" pageNumber="289" refId="ref91349" refString="SMETANA A. 1958: Fauna CSR. Svazek 12. Drabcikoviti - Staphylinidae I. Staphylinae (Coleoptera). [Fauna of the CSR. Volume 12. Rove beetles -- Staphylinidae I. Staphylinae (Coleoptera)]. Ceskoslovenska akademie ved, Praha, 435 pp (in Czech)." type="book" year="1958">Smetana, 1958</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A39B17E2FB8A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A39B17E2FB8A" pageId="64" pageNumber="289">
C-shaped sclerite of internal sac with spine-like basal extension (
<figureCitation id="131B2A33FFBA4B59B19EA3BB1711FC4A" box="[392,523,958,981]" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Figs K13D</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B209A3BB174AFC4A" box="[543,592,958,981]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">15D</figureCitation>
). N Palearctic distribution (
<figureCitation id="131B2A33FFBA4B59B158A3DB14D9FC6A" box="[334,451,990,1013]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A Q. unicolor Kiesenwetter, 1847; B Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C Q. hispanicus Bernhauer, 1898; D Q. pallipes Lucas, 1846; E Q. laticollis (Gravenhorst, 1802); F Q. strenuus Casey, 1915. I Parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="64" pageNumber="289">Fig. K14B</figureCitation>
). ........................................... ...............
<taxonomicName id="4C204D35FFBA4B59B123A3F817E2FB8A" ID-CoL="4R2YX" authority="(Gravenhorst, 1806)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[309,760,1021,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFBA4B59B123A3F81703FB8A" bold="true" box="[309,537,1021,1045]" italics="true" pageId="64" pageNumber="289">Quedius molochinus</emphasis>
(
<bibRefCitation id="EFB14B47FFBA4B59B231A3FB17EAFB8A" author="GRAVENHORST J. L. C." box="[551,752,1022,1045]" pageId="64" pageNumber="289" refId="ref83841" refString="GRAVENHORST J. L. C. 1806: Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae, 248 pp." type="book" year="1806">Gravenhorst, 1806</bibRefCitation>
)
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30D42E13FFBA4B59B081A41B17E2FAEA" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A41B17EDFB4A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A41B17EDFB4A" pageId="64" pageNumber="289">
15 Basal three antennomeres pale and antennae more elongate (antennomere 10 clearly longer than wide) (
<figureCitation id="131B2A33FFBA4B59B0DAA45B145DFBEA" box="[204,327,1118,1141]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">Fig. K15A</figureCitation>
). Elytra light to dark brown. Generally larger. Forebody length 5.495.93 (5.78) mm (
<figureCitation id="131B2A33FFBA4B59B2DBA47B15E8FB2A" captionStart="Fig" captionStartId="31.[151,182,2004,2023]" captionTargetBox="[207,1368,203,1953]" captionTargetId="figure-12@31.[191,1396,189,1962]" captionTargetPageId="31" captionText="Fig. 13. Aedeagus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. sundukovi Smetana, 2003; C Q. subunicolor Korge, 1961; D Q. molochinus (Gravenhorst, 1806); E Q. balticus Korge, 1960; F Q. meridiocarpathicus Smetana, 1958. I parameral view of whole aedeagus; II apices of paramere in antiparameral view with peg setae; III lateral view of whole aedeagus; IV apices of median lobe in parameral view." figureDoi="http://doi.org/10.5281/zenodo.7399732" httpUri="https://zenodo.org/record/7399732/files/figure.png" pageId="64" pageNumber="289">Fig. 13F</figureCitation>
). Boreal North America (Fig. 28). ...................... .....................
<taxonomicName id="4C204D35FFBA4B59B14CA4B817EDFB4A" ID-CoL="4R2YL" authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" box="[346,759,1213,1237]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="labradorensis">
<emphasis id="B954EAA4FFBA4B59B14CA4B81749FB4A" bold="true" box="[346,595,1213,1237]" italics="true" pageId="64" pageNumber="289">Quedius labradorensis</emphasis>
Smetana, 1965
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A4DB17E2FAEA" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A4DB17E2FAEA" pageId="64" pageNumber="289">
Basal three antennomeres darkened and antennae more robust (antennomere 10 more or less transverse) (
<figureCitation id="131B2A33FFBA4B59B147A51B14D4FAAA" box="[337,462,1310,1333]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="64" pageNumber="289">Fig. K15B</figureCitation>
). Elytra black or reddish. Generally smaller. Forebody length 4.975.86 (5.39) mm. ........................................................... 16
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</keyStep>
<keyStep id="30D42E13FFBA4B59B081A57B17E2F90A" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A57B17EDF9CA" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A57B17EDF9CA" pageId="64" pageNumber="289">
16 Paramere with basal attenuation and broadening towards apex forming a spoon like shape, peg setae situated in a semi-circle along the edge of the paramere (
<figureCitation id="131B2A33FFBA4B59B172A5DB14F8FA6A" box="[356,482,1502,1525]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="64" pageNumber="289">Figs K16A</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B1E6A5DB14E5FA6A" box="[496,511,1502,1525]" captionStart="Fig" captionStartId="18.[151,182,2027,2046]" captionTargetBox="[189,1393,195,2007]" captionTargetId="figure-13@18.[174,1413,185,2007]" captionTargetPageId="18" captionText="Fig. 8. Habitus of Quedius s. str. A Q. vicinus (Ménétriés, 1832); B Q. meridiocarpathicus Smetana, 1958; C Q. molochinus (Gravenhorst, 1806) (red elytra variant); D Q. molochinus (black elytra variant); E Q. balticus Korge, 1960; F Q. sundukovi Smetana, 2003." figureDoi="http://doi.org/10.5281/zenodo.7399722" httpUri="https://zenodo.org/record/7399722/files/figure.png" pageId="64" pageNumber="289">8</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B219A5DB1724FA6A" box="[527,574,1502,1525]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="64" pageNumber="289">17B</figureCitation>
). Coastal to mid elevation from
<collectingRegion id="49E4F854FFBA4B59B17FA5FB14A3F98A" box="[361,441,1534,1557]" country="United States of America" name="Oregon" pageId="64" pageNumber="289">Oregon</collectingRegion>
to Southern
<collectingRegion id="49E4F854FFBA4B59B254A5FB1796F98A" box="[578,652,1534,1557]" country="United States of America" name="Alaska" pageId="64" pageNumber="289">Alaska</collectingRegion>
including Vancouver and Haida Gwaii. ..................................... ......................................
<taxonomicName id="4C204D35FFBA4B59B1D9A63817EDF9CA" ID-CoL="4R2YH" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[463,759,1597,1621]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="horni">
<emphasis id="B954EAA4FFBA4B59B1D9A6381775F9CA" bold="true" box="[463,623,1597,1621]" italics="true" pageId="64" pageNumber="289">Quedius horni</emphasis>
<bibRefCitation id="EFB14B47FFBA4B59B263A63B17EDF9CA" author="HATCH M. H." box="[629,759,1598,1621]" pageId="64" pageNumber="289" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A65B17E2F90A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A65B17E2F90A" pageId="64" pageNumber="289">
Paramere only with at most slight attenuation forming a lanceolate shape (
<figureCitation id="131B2A33FFBA4B59B183A67B1735F90A" box="[405,559,1662,1685]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="64" pageNumber="289">Figs K16BD</figureCitation>
). ...................... 17
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFBA4B59B081A69B17E2F80A" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A69B17EDF88A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A69B17EDF88A" pageId="64" pageNumber="289">
17 Distributed across Boreal North America. South of the Northern Rocky Mountains (
<figureCitation id="131B2A33FFBA4B59B23BA6BB17B2F94A" box="[557,680,1726,1749]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="64" pageNumber="289">Fig. K17A</figureCitation>
). Teeth of median lobe clearly folded inwards (
<figureCitation id="131B2A33FFBA4B59B26FA6DB17EEF96A" box="[633,756,1758,1781]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="64" pageNumber="289">Figs K16B</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B0D3A6FB15E9F88A" box="[197,243,1790,1813]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="64" pageNumber="289">17E</figureCitation>
). .........
<taxonomicName id="4C204D35FFBA4B59B146A6F817EDF88A" ID-CoL="4R2YW" authority="Smetana, 1965" authorityName="Smetana" authorityYear="1965" box="[336,759,1789,1813]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="molochinoides">
<emphasis id="B954EAA4FFBA4B59B146A6F81749F88A" bold="true" box="[336,595,1789,1813]" italics="true" pageId="64" pageNumber="289">Quedius molochinoides</emphasis>
Smetana, 1965
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBA4B59B081A71B17E2F80A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A71B17E2F80A" pageId="64" pageNumber="289">
Central and Southern Rocky Mountains, Cascades, Blue Mountains and Sierra
<collectingRegion id="49E4F854FFBA4B59B21EA73B1743F8CA" box="[520,601,1854,1877]" country="United States of America" name="Nevada" pageId="64" pageNumber="289">Nevada</collectingRegion>
(
<figureCitation id="131B2A33FFBA4B59B279A73B17F1F8CA" box="[623,747,1854,1877]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="64" pageNumber="289">Fig. K17B</figureCitation>
). Teeth of median lobe either protruding or not clearly formed (
<figureCitation id="131B2A33FFBA4B59B136A77B14A0F80A" box="[288,442,1918,1941]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="64" pageNumber="289">Figs K16CD</figureCitation>
). ....................................... 19
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFBA4B58B081A79B17E2FEAA" lastPageId="65" lastPageNumber="290" pageId="64" pageNumber="289">
<paragraph id="8B9F36B6FFBA4B59B081A79B17EDF78A" blockId="64.[151,760,190,2069]" pageId="64" pageNumber="289">
<keyLead id="30D19583FFBA4B59B081A79B17EDF78A" pageId="64" pageNumber="289">
18. Found in the Cascades, Blue Mountains and Sierra
<collectingRegion id="49E4F854FFBA4B59B0D3A7BB140CF84A" box="[197,278,1982,2005]" country="United States of America" name="Nevada" pageId="64" pageNumber="289">Nevada</collectingRegion>
(
<figureCitation id="131B2A33FFBA4B59B13AA7BB14B6F84A" box="[300,428,1982,2005]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="64" pageNumber="289">Fig. K18A</figureCitation>
). Paramere and median lobe clearly broader (
<figureCitation id="131B2A33FFBA4B59B165A7DB14F6F86A" box="[371,492,2014,2037]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="64" pageNumber="289">Figs K16C</figureCitation>
,
<figureCitation id="131B2A33FFBA4B59B1E0A7DB1732F86A" box="[502,552,2014,2037]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="64" pageNumber="289">17D</figureCitation>
). ............................ .......................................
<taxonomicName id="4C204D35FFBA4B59B1C3A7F817EDF78A" ID-CoL="4R2YM" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[469,759,2045,2069]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="64" pageNumber="289" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B954EAA4FFBA4B59B1C3A7F81775F78A" bold="true" box="[469,623,2045,2069]" italics="true" pageId="64" pageNumber="289">Quedius lanei</emphasis>
<bibRefCitation id="EFB14B47FFBA4B59B263A7FB17EDF78A" author="HATCH M. H." box="[629,759,2046,2069]" pageId="64" pageNumber="289" pagination="1 - 384" refId="ref84249" refString="HATCH M. H. 1957: The beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology 16 (2): 1 - 384." type="journal article" year="1957">Hatch, 1957</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBB4B58B081A0BB17E2FEAA" blockId="65.[151,760,190,757]" pageId="65" pageNumber="290">
<keyLead id="30D19583FFBB4B58B081A0BB17E2FEAA" pageId="65" pageNumber="290">
Found in Central and Southern Rocky Mountains (
<figureCitation id="131B2A33FFBB4B58B0DAA0DB145BFF6A" box="[204,321,222,245]" captionStart="Fig" captionStartId="57.[151,182,2034,2053]" captionTargetBox="[192,1396,190,2012]" captionTargetId="figure-13@57.[192,1397,189,2013]" captionTargetPageId="57" captionText="Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399744" httpUri="https://zenodo.org/record/7399744/files/figure.png" pageId="65" pageNumber="290">Fig. K18B</figureCitation>
). Paramere and median lobe clearly more slender (
<figureCitation id="131B2A33FFBB4B58B136A0FB1483FE8A" box="[288,409,254,277]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="65" pageNumber="290">Figs K16D</figureCitation>
,
<figureCitation id="131B2A33FFBB4B58B1B2A0FB14CEFE8A" box="[420,468,254,277]" captionStart="Fig" captionStartId="52.[151,182,1666,1685]" captionTargetBox="[170,1418,189,1655]" captionTargetId="figure-13@52.[170,1418,189,1659]" captionTargetPageId="52" captionText="Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath." figureDoi="http://doi.org/10.5281/zenodo.7399742" httpUri="https://zenodo.org/record/7399742/files/figure.png" pageId="65" pageNumber="290">17C</figureCitation>
). ........................................ ...........................................
<emphasis id="B954EAA4FFBB4B58B1FAA11817E2FEAA" bold="true" box="[492,760,285,309]" pageId="65" pageNumber="290">
<taxonomicName id="4C204D35FFBB4B58B1FAA1181786FEAA" authority="Hansen &amp; Brunke &amp; Simonsen &amp; Solodovnikov, 2022" authorityName="Hansen &amp; Brunke &amp; Simonsen &amp; Solodovnikov" authorityYear="2022" box="[492,668,285,309]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="290" phylum="Arthropoda" rank="species" species="altanai" status="sp. nov.">
<emphasis id="B954EAA4FFBB4B58B1FAA1181786FEAA" bold="true" box="[492,668,285,309]" italics="true" pageId="65" pageNumber="290">Quedius altanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757DFFFBB4B58B2B4A11B17E2FEAA" box="[674,760,286,309]" pageId="65" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFBB4B58B081A13B17E2FE6A" pageId="65" pageNumber="290">
<paragraph id="8B9F36B6FFBB4B58B081A13B17E2FE2A" blockId="65.[151,760,190,757]" pageId="65" pageNumber="290">
<keyLead id="30D19583FFBB4B58B081A13B17E2FE2A" pageId="65" pageNumber="290">
19 Paramere rather slender and clear C-sclerite of the internal sac (
<figureCitation id="131B2A33FFBB4B58B15AA15B14DFFEEA" box="[332,453,350,373]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="65" pageNumber="290">Figs K19A</figureCitation>
,
<figureCitation id="131B2A33FFBB4B58B1D9A15B171BFEEA" box="[463,513,350,373]" captionStart="Fig" captionStartId="39.[152,183,2033,2052]" captionTargetBox="[284,1300,1425,2027]" captionTargetId="figure-692@39.[276,1311,1425,2030]" captionTargetPageId="39" captionText="Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens." figureDoi="http://doi.org/10.5281/zenodo.7399736" httpUri="https://zenodo.org/record/7399736/files/figure.png" pageId="65" pageNumber="290">15D</figureCitation>
). Introduced in parts of Eastern North America, seems to be spreading. ........ ...............
<taxonomicName id="4C204D35FFBB4B58B123A19817E2FE2A" ID-CoL="4R2YX" authority="(Gravenhorst, 1806)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[309,760,413,437]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="290" phylum="Arthropoda" rank="species" species="molochinus">
<emphasis id="B954EAA4FFBB4B58B123A1981703FE2A" bold="true" box="[309,537,413,437]" italics="true" pageId="65" pageNumber="290">Quedius molochinus</emphasis>
(
<bibRefCitation id="EFB14B47FFBB4B58B231A19B17EAFE2A" author="GRAVENHORST J. L. C." box="[551,752,414,437]" pageId="65" pageNumber="290" refId="ref83841" refString="GRAVENHORST J. L. C. 1806: Monographia Coleopterorum Micropterorum. H. Dieterich, Gottingae, 248 pp." type="book" year="1806">Gravenhorst, 1806</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBB4B58B081A1BB17E2FE6A" blockId="65.[151,760,190,757]" pageId="65" pageNumber="290">
<keyLead id="30D19583FFBB4B58B081A1BB17E2FE6A" pageId="65" pageNumber="290">
Paramere rather broad and No C-sclerite of internal sac (
<figureCitation id="131B2A33FFBB4B58B0E3A1DB1494FE6A" box="[245,398,478,501]" captionStart="Fig" captionStartId="58.[151,182,2028,2047]" captionTargetBox="[194,1393,189,2000]" captionTargetId="figure-13@58.[194,1393,189,2003]" captionTargetPageId="58" captionText="Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399746" httpUri="https://zenodo.org/record/7399746/files/figure.png" pageId="65" pageNumber="290">Figs K19BC</figureCitation>
).
<emphasis id="B954EAA4FFBB4B58B1B6A1D81771FE6A" bold="true" box="[416,619,477,501]" pageId="65" pageNumber="290">
<taxonomicName id="4C204D35FFBB4B58B1B6A1D81707FE6A" ID-CoL="4R2YN" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[416,541,477,501]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="290" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFBB4B58B1B6A1D81707FE6A" bold="true" box="[416,541,477,501]" italics="true" pageId="65" pageNumber="290">Q. laticollis</emphasis>
</taxonomicName>
-group
</emphasis>
.............. 20
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D42E13FFBB4B58B081A1FB17EDFD6A" pageId="65" pageNumber="290">
<paragraph id="8B9F36B6FFBB4B58B081A1FB17E2FDEA" blockId="65.[151,760,190,757]" pageId="65" pageNumber="290">
<keyLead id="30D19583FFBB4B58B081A1FB17E2FDEA" pageId="65" pageNumber="290">
20 Peg setae of paramere forming a solid triangular field, usually with two wide and short extensions posteriorly (
<figureCitation id="131B2A33FFBB4B58B150A23B14A5FDCA" box="[326,447,574,597]" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="65" pageNumber="290">Figs K20A</figureCitation>
,
<figureCitation id="131B2A33FFBB4B58B1DFA23B14E2FDCA" box="[457,504,574,597]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="65" pageNumber="290">16E</figureCitation>
). E North America (
<figureCitation id="131B2A33FFBB4B58B2DBA23B1413FDEA" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="65" pageNumber="290">Fig. K20B</figureCitation>
). ........
<taxonomicName id="4C204D35FFBB4B58B14FA25817E2FDEA" ID-CoL="4R2YN" authority="(Gravenhorst, 1802)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[345,760,605,629]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="290" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B954EAA4FFBB4B58B14FA2581702FDEA" bold="true" box="[345,536,605,629]" italics="true" pageId="65" pageNumber="290">Quedius laticollis</emphasis>
(
<bibRefCitation id="EFB14B47FFBB4B58B231A25B17EAFDEA" author="GRAVENHORST J. L. C." box="[551,752,606,629]" pageId="65" pageNumber="290" refId="ref83803" refString="GRAVENHORST J. L. C. 1802: Coleoptera Microptera Brunsvicensia nec non exoticorum quotquot exstant in collectionibus entomologorum Brunsvicensium in genera, familiae et species distribuit. C. Reichard, Brunsuigae, 206 pp." type="book" year="1802">Gravenhorst, 1802</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9F36B6FFBB4B58B081A27B17EDFD6A" blockId="65.[151,760,190,757]" pageId="65" pageNumber="290">
<keyLead id="30D19583FFBB4B58B081A27B17EDFD6A" pageId="65" pageNumber="290">
Peg setae of paramere arranged into two short longitudinal rows, which more or less diverge posteriorly (
<figureCitation id="131B2A33FFBB4B58B150A2BB14A4FD4A" box="[326,446,702,725]" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="65" pageNumber="290">Figs K20C</figureCitation>
,
<figureCitation id="131B2A33FFBB4B58B1DEA2BB14EFFD4A" box="[456,501,702,725]" captionStart="Fig" captionStartId="47.[858,889,1894,1913]" captionTargetBox="[168,1437,204,2073]" captionTargetId="figure-12@47.[151,1436,189,2079]" captionTargetPageId="47" captionText="Fig. 16. Aedeagus of Quedius s. str. A Q. labradorensis Smetana, 1965; B Q. horni Hatch, 1957; C Q. altanai Hansen &amp; Brunke, sp. nov.; D Q. lanei Hatch, 1957; E Q. molochinoides Smetana, 1965. I parameral view of whole aedeagus, II apices of paramere in antiparameral view with peg setae, III lateral view of whole aedeagus, IV apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399740" httpUri="https://zenodo.org/record/7399740/files/figure.png" pageId="65" pageNumber="290">16F</figureCitation>
). Great American Plains (
<figureCitation id="131B2A33FFBB4B58B0DAA2DB145EFD6A" box="[204,324,734,757]" captionStart="Fig" captionStartId="59.[151,182,2028,2047]" captionTargetBox="[190,1392,189,2003]" captionTargetId="figure-13@59.[190,1393,189,2003]" captionTargetPageId="59" captionText="Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399748" httpUri="https://zenodo.org/record/7399748/files/figure.png" pageId="65" pageNumber="290">Fig. K20D</figureCitation>
). ...........
<taxonomicName id="4C204D35FFBB4B58B1BBA2D817EDFD6A" ID-CoL="4R2Z5" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[429,759,733,757]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="290" phylum="Arthropoda" rank="species" species="strenuus">
<emphasis id="B954EAA4FFBB4B58B1BBA2D81775FD6A" bold="true" box="[429,623,733,757]" italics="true" pageId="65" pageNumber="290">Quedius strenuus</emphasis>
<bibRefCitation id="EFB14B47FFBB4B58B260A2DB17EDFD6A" author="CASEY T. L." box="[630,759,734,757]" pageId="65" pageNumber="290" pagination="395 - 450" refId="ref81365" refString="CASEY T. L. 1915: Studies in some staphylinid genera of North America. Memoirs on the Coleoptera 6: 395 - 450." type="journal article" year="1915">Casey, 1915</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>