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<document id="A073570EF97A62B8C3C03371016E083D" ID-CLB-Dataset="161923" ID-DOI="10.11646/phytotaxa.567.3.9" ID-GBIF-Dataset="4d517799-5bb6-4099-9a48-ee03357e2bb5" ID-ISSN="1179-3163" ID-Zenodo-Dep="7156782" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665138932319" checkinUser="plazi" docAuthor="Hoang, Thanh Son &amp; Lin, Che-Wei" docDate="2022" docId="0388B3556E32FF96FF35F9F6FC73F60C" docLanguage="en" docName="phytotaxa.567.3.9.pdf" docOrigin="Phytotaxa 567 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.567.3.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Begonia tuanii T. S. Hoang &amp; C. W. Lin. A, B. Habit 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="298" masterDocId="FFB1CB2D6E32FF93FFBDFF8EFF8AF728" masterDocTitle="Begonia tuanii (sect. Platycentrum, Begoniaceae), a new species from Lao Cai Province, North Vietnam" masterLastPageNumber="300" masterPageNumber="294" pageNumber="294" updateTime="1699390267279" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="6662534593C4CBF9839998C619642E70" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0BB43A7B741C45390D50C8089EA3A504">
<mods:title id="8189650B54EBCCB8763F6B1B83CCD289">Begonia tuanii (sect. Platycentrum, Begoniaceae), a new species from Lao Cai Province, North Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="74878FCA410BF314E236E4DD73198116" type="personal">
<mods:role id="FABA90A823C195C5B777C83E059FBCFB">
<mods:roleTerm id="60AE4DC9CABFD8F96FEFB86D1EDD3E6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3A6CE9C301D4E5ED769BF46DBB5FCC03">Hoang, Thanh Son</mods:namePart>
<mods:nameIdentifier id="F36410C31014AE31BA2E1D81E5523319" type="ORCID">0000-0003-0351-929X</mods:nameIdentifier>
<mods:affiliation id="B68E33CB468CD788E0657D252E484A3D">Institute of Applied Technology, Thu Dau Mot University, Binh Duong Province 820000, Vietnam &amp; hoangthanhson @ tdmu. edu. vn; https: // orcid. org / 0000 - 0003 - 0351 - 929 X</mods:affiliation>
<mods:nameIdentifier id="AE39484B80CFFF434FA62C980198F15F" type="email">hoangthanhson@tdmu.edu.vn</mods:nameIdentifier>
</mods:name>
<mods:name id="495053C86FF9B74396CD052D5F102A6C" type="personal">
<mods:role id="6A18A45348EBFA1715442DD811E7346F">
<mods:roleTerm id="209F17FFF6E056D327D36FBA44290A7A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9703E9C43A6C08C88C2389E62FCC6E5">Lin, Che-Wei</mods:namePart>
<mods:nameIdentifier id="B73C72697F09C0AB042C57A5A49032AA" type="ORCID">0000-0002-8564-5793</mods:nameIdentifier>
<mods:affiliation id="24803D790F51C6791BAD0D9C248337EE">Herbarium of Taiwan Forestry Research Institute, Taiwan Forestry Research Institute, No. 53, Nan-Hai Road, Taipei 100, Taiwan &amp; varalba @ gmail. com; https: // orcid. org / 0000 - 0002 - 8564 - 5793</mods:affiliation>
<mods:nameIdentifier id="88798304FCD17194B6CDFFB3781705FB" type="email">varalba@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="70C6ECE5FAEC002D87BDF88FCF857432">text</mods:typeOfResource>
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<mods:title id="E3F44167DC1642FFA99D7C39BC6E6298">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F7BF44796F2D94884F231DB681B48B76">
<mods:date id="825ED382293C96E0D782738CB5024011">2022</mods:date>
<mods:detail id="22951F48803186CC75B8C3FAB3F24AC8" type="pubDate">
<mods:number id="95B9F7DA535E857D7C72F0DCA77A3A40">2022-10-07</mods:number>
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<mods:number id="8EC7A6F6F3D416360D3BEE9E1B6EC7FE">567</mods:number>
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<mods:detail id="166EAE085CD14275644C87D5BD274CBA" type="issue">
<mods:number id="9CC2298FECA2B19070FCF79376F17E89">3</mods:number>
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<mods:start id="D79DFDAEFEE110841AED52A1817E665B">294</mods:start>
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<treatment id="0388B3556E32FF96FF35F9F6FC73F60C" ID-DOI="http://doi.org/10.5281/zenodo.7182708" ID-GBIF-Taxon="202581999" ID-Zenodo-Dep="7182708" LSID="urn:lsid:plazi:treatment:0388B3556E32FF96FF35F9F6FC73F60C" httpUri="http://treatment.plazi.org/id/0388B3556E32FF96FF35F9F6FC73F60C" lastPageId="5" lastPageNumber="298" pageId="0" pageNumber="294">
<subSubSection id="C33B51C86E32FF93FF35F9F6FCFAF1A5" box="[136,880,1656,1678]" pageId="0" pageNumber="294" type="nomenclature">
<paragraph id="8B9E02436E32FF93FF35F9F6FCFAF1A5" blockId="0.[136,1452,1656,1786]" box="[136,880,1656,1678]" pageId="0" pageNumber="294">
<taxonomicName id="4C2179C06E32FF93FF35F9F6FE71F1A6" ID-CoL="d2837769-b84b-44f0-8e87-090d911ef2f7" authority="T.S.Hoang &amp; C.W.Lin" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[136,507,1656,1678]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="0" pageNumber="294" phylum="Tracheophyta" rank="species" species="tuanii" status="sp. nov.">
<emphasis id="B955DE516E32FF93FF35F9F6FE96F1A5" bold="true" box="[136,284,1656,1677]" italics="true" pageId="0" pageNumber="294">Begonia tuanii</emphasis>
T.S.Hoang &amp; C.W.Lin
</taxonomicName>
,
<emphasis id="B955DE516E32FF93FDB9F9F6FDC5F1A5" bold="true" box="[516,591,1656,1677]" italics="true" pageId="0" pageNumber="294">
<taxonomicNameLabel id="A266632A6E32FF93FDB9F9F6FDC5F1A5" box="[516,591,1656,1677]" pageId="0" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
§
<taxonomicName id="4C2179C06E32FF93FDDAF9F6FD62F1A5" authorityName="A. de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" box="[615,744,1656,1677]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="0" pageNumber="294" phylum="Tracheophyta" rank="section" section="Platycentrum">
<emphasis id="B955DE516E32FF93FDDAF9F6FD62F1A5" box="[615,744,1656,1677]" italics="true" pageId="0" pageNumber="294">Platycentrum</emphasis>
</taxonomicName>
. (
<figureCitation id="131A1EC66E32FF93FD46F9F6FCB7F1A5" box="[763,829,1656,1678]" captionStart="FIGURE 2" captionStartId="2.[136,234,1959,1981]" captionTargetBox="[199,1389,264,1905]" captionTargetId="figure-47@2.[177,1410,245,1934]" captionTargetPageId="2" captionText="FIGURE 2. Begonia tuanii T.S.Hoang &amp; C.W.Lin.A. Habit; B, B’. Portion of leaf, upper and lower surfaces; C. Stipule; D–D’’. Bracts; D’’’. Bracteole; E, E’. Staminate flower, face and side views; F, F’, F’’. Stamens dorsal, ventral and side views; G, G’. Pistillate flower, face and side views; H, H’. Style and stigmatic band, dorsal and ventral views; I. Cross section of ovary." figureDoi="http://doi.org/10.5281/zenodo.7156786" httpUri="https://zenodo.org/record/7156786/files/figure.png" pageId="0" pageNumber="294">Figs. 2</figureCitation>
&amp;
<figureCitation id="131A1EC66E32FF93FCE1F9F6FCE2F1A5" box="[860,872,1656,1677]" captionStart="FIGURE 3" captionStartId="3.[136,234,1935,1957]" captionTargetBox="[151,1436,190,1911]" captionTargetId="figure-19@3.[151,1436,190,1911]" captionTargetPageId="3" captionText="FIGURE 3. Begonia tuanii T.S.Hoang &amp; C.W.Lin. A, B. Habit and habitat; C. Leaf, upper and lower views; D. Portion of leaf, upper surface, E. same, lower surface; F. Inflorescence; G. Stipule, also showing stem and petioles; H. Immature inflorescence, also showing bracts; I. Staminate flowers; J. Androecium; K. Pistillate flower; L. Cross section of ovary. All photos from by Hoang Thanh Son." figureDoi="http://doi.org/10.5281/zenodo.7156788" httpUri="https://zenodo.org/record/7156788/files/figure.png" pageId="0" pageNumber="294">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B51C86E32FF93FF35F912FDCBF1D1" pageId="0" pageNumber="294" type="materials_examined">
<paragraph id="8B9E02436E32FF93FF35F912FDCBF1D1" blockId="0.[136,1452,1656,1786]" pageId="0" pageNumber="294">
<emphasis id="B955DE516E32FF93FF35F912FF50F199" bold="true" box="[136,218,1692,1713]" pageId="0" pageNumber="294">Type:—</emphasis>
<materialsCitation id="3B49081E6E32FF93FF67F912FE1CF1D1" ID-GBIF-Occurrence="3922697302" collectingDate="2021-02-18" collectorName="Hoang Thanh Son &amp; Nguyen Van Tuan" country="Vietnam" elevation="1258" latitude="22.35639" location="Ta Phoi Commune" longLatPrecision="1" longitude="103.92227" municipality="City" pageId="0" pageNumber="294" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<collectingCountry id="F33642D36E32FF93FF67F912FEC7F199" box="[218,333,1692,1713]" name="Vietnam" pageId="0" pageNumber="294">VIETNAM</collectingCountry>
:
<collectingRegion id="49E5CCA16E32FF93FEEAF912FD8EF19A" box="[343,516,1692,1714]" country="Vietnam" name="Lao Cai" pageId="0" pageNumber="294">Lao Cai Province</collectingRegion>
,
<collectingRegion id="49E5CCA16E32FF93FDB2F912FDD7F19A" box="[527,605,1692,1714]" country="Vietnam" name="Lao Cai" pageId="0" pageNumber="294">Lao Cai</collectingRegion>
<collectingMunicipality id="6BFA98396E32FF93FDD8F912FD05F19A" box="[613,655,1692,1714]" pageId="0" pageNumber="294">City</collectingMunicipality>
,
<location id="8EFE54986E32FF93FD24F912FCD9F199" LSID="urn:lsid:plazi:treatment:0388B3556E32FF96FF35F9F6FC73F60C:8EFE54986E32FF93FD24F912FCD9F199" box="[665,851,1692,1714]" country="Vietnam" latitude="22.35639" longLatPrecision="1" longitude="103.92227" municipality="City" name="Ta Phoi Commune" pageId="0" pageNumber="294" stateProvince="Lao Cai">Ta Phoi Commune</location>
,
<geoCoordinate id="EE1564846E32FF93FCE0F912FC6AF199" box="[861,992,1692,1713]" degrees="103.922273" direction="east" orientation="longitude" pageId="0" pageNumber="294" precision="1" value="103.92227">103.922273E</geoCoordinate>
,
<geoCoordinate id="EE1564846E32FF93FC56F912FBEFF19A" box="[1003,1125,1692,1714]" degrees="22.356390" direction="north" orientation="latitude" pageId="0" pageNumber="294" precision="1" value="22.35639">22.356390N</geoCoordinate>
,
<quantity id="4CD9AFA66E32FF93FBCDF912FB4AF199" box="[1136,1216,1692,1714]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="0" pageNumber="294" unit="m" value="1258.0">
<elevation id="000CE5706E32FF93FBCDF912FB4AF199" box="[1136,1216,1692,1714]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="0" pageNumber="294" unit="m" value="1258.0">1,258 m</elevation>
</quantity>
elev., on the moist land and rocks in secondary broad-leaved evergreen forest,
<date id="FF9F24836E32FF93FD73F94EFCF6F1FD" box="[718,892,1728,1750]" pageId="0" pageNumber="294" value="2021-02-18">
<collectingDate id="EFDBDD6B6E32FF93FD73F94EFCF6F1FD" box="[718,892,1728,1750]" pageId="0" pageNumber="294" value="2021-02-18">18 February 2021</collectingDate>
</date>
, flowers and fruits,
<emphasis id="B955DE516E32FF93FB83F94EFF67F1D1" italics="true" pageId="0" pageNumber="294">
<collectorName id="26D467956E32FF93FB83F94EFB7AF1FD" box="[1086,1264,1728,1749]" pageId="0" pageNumber="294">Hoang Thanh Son</collectorName>
,
<collectorName id="26D467956E32FF93FB46F94EFA2DF1FD" box="[1275,1447,1728,1749]" pageId="0" pageNumber="294">Nguyen Van Tuan</collectorName>
, 8455
</emphasis>
(
<typeStatus id="549ABCE16E32FF93FF47F96AFEDAF1D2" box="[250,336,1764,1786]" pageId="0" pageNumber="294" type="holotype">holotype</typeStatus>
VAFS;
</materialsCitation>
<materialsCitation id="3B49081E6E32FF93FE21F96AFDCBF1D1" ID-GBIF-Occurrence="3922697301" box="[412,577,1764,1786]" collectingDate="2021-02-18" collectorName="Hoang Thanh Son &amp; Nguyen Van Tuan" country="Vietnam" elevation="1258" latitude="22.35639" location="Ta Phoi Commune" longLatPrecision="1" longitude="103.92227" municipality="City" pageId="0" pageNumber="294" specimenCount="1" stateProvince="Lao Cai" typeStatus="isotype">isotypes VNMN)</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B51C86E32FF92FF35F8A6FC4DF44C" lastPageId="1" lastPageNumber="295" pageId="0" pageNumber="294" type="description">
<paragraph id="8B9E02436E32FF92FF35F8A6FC4DF44C" blockId="0.[136,1452,1832,2075]" lastBlockId="1.[136,1452,158,869]" lastPageId="1" lastPageNumber="295" pageId="0" pageNumber="294">
Plant perennial, monoecious, terrestrial.
<emphasis id="B955DE516E32FF93FDF2F8A6FD06F06A" bold="true" box="[591,652,1832,1858]" pageId="0" pageNumber="294">Stem</emphasis>
erect, unbranched or little branched, yellowish-green to reddish-green,
<quantity id="4CD9AFA66E32FF93FF35F8C2FF70F04F" box="[136,250,1868,1895]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="0" pageNumber="294" unit="cm" value="20.0" valueMax="25.0" valueMin="15.0">15–25 cm</quantity>
tall,
<quantity id="4CD9AFA66E32FF93FE87F8C2FE11F04F" box="[314,411,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="0" pageNumber="294" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
across, covered with curved puberulous; internodes
<quantity id="4CD9AFA66E32FF93FC42F8C2FBDFF04F" box="[1023,1109,1868,1895]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="0" pageNumber="294" unit="cm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 cm</quantity>
long, nodes slightly swollen.
<emphasis id="B955DE516E32FF93FF35F8FEFF56F0A2" bold="true" box="[136,220,1904,1930]" pageId="0" pageNumber="294">Stipule</emphasis>
decidous, ovate to elliptic, 18–23 ×
<quantity id="4CD9AFA66E32FF93FDC9F8FEFD6AF0A3" box="[628,736,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="0" pageNumber="294" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, pale green to reddish-green, hyaline, keeled, abaxially sparsely curved puberulous, margin entire, apex cuspidate, cusp
<emphasis id="B955DE516E32FF93FD42F81BFC93F086" box="[767,793,1941,1966]" italics="true" pageId="0" pageNumber="294">ca</emphasis>
.
<quantity id="4CD9AFA66E32FF93FC95F81AFCE0F087" box="[808,874,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="294" unit="mm" value="1.0">1 mm</quantity>
long.
<emphasis id="B955DE516E32FF93FC0EF81AFB8EF086" bold="true" box="[947,1028,1940,1966]" pageId="0" pageNumber="294">Petiole</emphasis>
D-shaped, reddish-brown to pinkishgreen, grooved above,
<quantity id="4CD9AFA66E32FF93FE34F836FD8AF0FB" box="[393,512,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="10.0" metricValueMin="3.5" pageId="0" pageNumber="294" unit="cm" value="6.75" valueMax="10.0" valueMin="3.5">3.5–10 cm</quantity>
long,
<quantity id="4CD9AFA66E32FF93FDF5F836FD30F0FB" box="[584,698,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.0" metricValueMin="3.5" pageId="0" pageNumber="294" unit="mm" value="4.75" valueMax="6.0" valueMin="3.5">3.5–6 mm</quantity>
across, abaxially curved puberulous.
<emphasis id="B955DE516E32FF93FBDFF836FB3EF0FA" bold="true" box="[1122,1204,1976,2002]" pageId="0" pageNumber="294">Leaves</emphasis>
4–6, oblique, slightly pointing upward; lamina ovate, thinly succulent, basifixed, asymmetric, margin denticulate, crenulate to subentire, apex attenuate; lamina
<quantity id="4CD9AFA66E32FF93FE32F78EFD9EFF33" box="[399,532,2048,2075]" metricMagnitude="-1" metricUnit="m" metricValue="2.025" metricValueMax="2.7" metricValueMin="1.35" pageId="0" pageNumber="294" unit="cm" value="20.25" valueMax="27.0" valueMin="13.5">13.5–27 cm</quantity>
long (basal lobes included),
<quantity id="4CD9AFA66E32FF93FCE3F78EFC60FF33" box="[862,1002,2048,2075]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.42" metricValueMin="0.73" pageId="0" pageNumber="294" unit="cm" value="10.75" valueMax="14.2" valueMin="7.3">7.3–14.2 cm</quantity>
wide, broadside
<quantity id="4CD9AFA66E32FF93FB10F78EFAA1FF33" box="[1197,1323,2048,2075]" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="9.5" metricValueMin="4.8" pageId="0" pageNumber="294" unit="cm" value="7.15" valueMax="9.5" valueMin="4.8">4.8–9.5 cm</quantity>
wide, base unequal, basal lobes cordate on broadside,
<quantity id="4CD9AFA66E33FF92FDDCFF10FD42F791" box="[609,712,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="4.0" metricValueMin="2.2" pageId="1" pageNumber="295" unit="cm" value="3.1" valueMax="4.0" valueMin="2.2">2.2–4 cm</quantity>
long, adaxially olive green to dusky green with reddish veins, very sparsely scabridulous between veins; abaxially magenta to pale green tinged red, densely curved puberulous on veins; venation palmate-pinnate, midrib distinguishable,
<quantity id="4CD9AFA66E33FF92FD07FF68FCCFF629" box="[698,837,230,257]" metricMagnitude="-1" metricUnit="m" metricValue="1.51" metricValueMax="2.25" metricValueMin="0.77" pageId="1" pageNumber="295" unit="cm" value="15.1" valueMax="22.5" valueMin="7.7">7.7–22.5 cm</quantity>
long,
<emphasis id="B955DE516E33FF92FC36FF69FC2FF628" box="[907,933,231,256]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
. 3 major lateral veins on either side of midrib, other primary veins branching dichotomously, minor veins reticulate.
<emphasis id="B955DE516E33FF92FC21FE84FC55F60C" bold="true" box="[924,991,266,292]" pageId="1" pageNumber="295">Bract</emphasis>
decidous, pale yellowish-green to dusky red, hyaline, those at basal node of inflorescence ovate to elliptic, to 1.8 ×
<quantity id="4CD9AFA66E33FF92FC7DFEA0FB84F661" box="[960,1038,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="295" unit="cm" value="0.8">0.8 cm</quantity>
, apex obtuse, sometimes with a short apiculate, margin entire to sparsely denticulate or puberulous; bracts at summit of inflorescence similar but smaller.
<emphasis id="B955DE516E33FF92FF35FEF8FEACF6B8" bold="true" box="[136,294,374,400]" pageId="1" pageNumber="295">Inflorescence</emphasis>
axillary in upper leaf axils, bisexual, cymosely branching panicle
<quantity id="4CD9AFA66E33FF92FB98FEF8FB1DF6B9" box="[1061,1175,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="295" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
long, peduncle crimson to pinkish-green,
<emphasis id="B955DE516E33FF92FEEFFE15FEF9F69C" box="[338,371,411,436]" italics="true" pageId="1" pageNumber="295">ca.</emphasis>
<quantity id="4CD9AFA66E33FF92FEC0FE14FE41F69D" box="[381,459,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="295" unit="cm" value="5.5">5.5 cm</quantity>
long,
<emphasis id="B955DE516E33FF92FDAAFE15FDBBF69C" box="[535,561,411,436]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
. 2 orders of branching, erect or ascending, glabrous, protandrous.
<emphasis id="B955DE516E33FF92FA8FFE14FF54F6F0" bold="true" pageId="1" pageNumber="295">Staminate flower:</emphasis>
pedicel pink to yellowish-pink,
<quantity id="4CD9AFA66E33FF92FDFBFE30FD24F6F1" box="[582,686,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="5.0" metricValueMin="3.2" pageId="1" pageNumber="295" unit="cm" value="4.1" valueMax="5.0" valueMin="3.2">3.2–5 cm</quantity>
long, glabrous, tepals 4, white to pale pink, glabrous; outer 2 widely ovate, margin entire, 2.5–4 ×
<quantity id="4CD9AFA66E33FF92FE5DFE6CFDE9F6D5" box="[480,611,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="1" pageNumber="295" unit="cm" value="2.95" valueMax="3.5" valueMin="2.4">2.4–3.5 cm</quantity>
, inner 2 elliptic to obovate, 2–3.5 ×
<quantity id="4CD9AFA66E33FF92FBB3FE6CFBEDF6D5" box="[1038,1127,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="295" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
; androecium actinomorphic, stamens 90–120, filaments fused into a column to
<quantity id="4CD9AFA66E33FF92FD03FD88FD75F509" box="[702,767,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="295" unit="mm" value="5.0">5 mm</quantity>
long; anthers
<quantity id="4CD9AFA66E33FF92FC22FD88FB98F509" box="[927,1042,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="295" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
long, widely obovate, apex rounded or truncate, filaments
<quantity id="4CD9AFA66E33FF92FEC3FDA4FE56F56D" box="[382,476,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="295" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
long.
<emphasis id="B955DE516E33FF92FD99FDA4FD60F56C" bold="true" box="[548,746,554,580]" pageId="1" pageNumber="295">Pistillate flower:</emphasis>
pedicel pinkish-green to red,
<emphasis id="B955DE516E33FF92FB81FDA5FBDCF56C" box="[1084,1110,555,580]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
.
<quantity id="4CD9AFA66E33FF92FBD8FDA4FB38F56D" box="[1125,1202,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="295" unit="cm" value="4.5">4.5 cm</quantity>
long, sometimes with a pairs alternate ovate bracteoles
<emphasis id="B955DE516E33FF92FDB0FDC1FDADF540" box="[525,551,591,616]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
. 1 ×
<quantity id="4CD9AFA66E33FF92FDD6FDC0FD34F541" box="[619,702,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="295" unit="cm" value="0.7">0.7 cm</quantity>
; tepals 5, white to pinkish-white, glabrous, outer 4 tepals larger, suborbicular to widely obovate, 1.5–2.5 ×
<quantity id="4CD9AFA66E33FF92FDCBFDFCFD73F5A5" box="[630,761,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="1" pageNumber="295" unit="cm" value="2.1" valueMax="2.7" valueMin="1.5">1.5–2.7 cm</quantity>
, inner 1 tepal obovate, 1.5–2.2 ×
<quantity id="4CD9AFA66E33FF92FB3BFDFCFB7FF5A5" box="[1158,1269,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="1" pageNumber="295" unit="cm" value="1.2" valueMax="1.4" valueMin="1.0">1–1.4 cm</quantity>
; ovary green to reddish-green, body trigonous-ellipsoid,
<emphasis id="B955DE516E33FF92FDE1FD19FDFCF598" box="[604,630,663,688]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
.
<quantity id="4CD9AFA66E33FF92FD35FD18FD52F599" box="[648,728,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="295" unit="cm" value="1.5">1.5 cm</quantity>
long,
<quantity id="4CD9AFA66E33FF92FC9BFD18FCFCF599" box="[806,886,662,689]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="295" unit="cm" value="1.2">1.2 cm</quantity>
across, sparsely verrucose, 3-winged; wings red to green, subsequal, abaxial wing slightly wide than lateral ones, very narrowly crescent-shaped,
<emphasis id="B955DE516E33FF92FB62FD35FB73F5FC" box="[1247,1273,699,724]" italics="true" pageId="1" pageNumber="295">ca</emphasis>
. 20 ×
<quantity id="4CD9AFA66E33FF92FAFBFD34FA2DF5FD" box="[1350,1447,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="295" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
, margin entire, verrucose; ovary 3-locular, placenta bilamellated; styles 3, greenish-yellow, bifid, deeply divided to base, Y-shaped,
<emphasis id="B955DE516E33FF92FE83FC8DFEEAF434" box="[318,352,771,796]" italics="true" pageId="1" pageNumber="295">ca.</emphasis>
<quantity id="4CD9AFA66E33FF92FED5FC8DFE23F435" box="[360,425,771,797]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="295" unit="mm" value="7.0">7 mm</quantity>
long; stigmas in a spiral band and papillose all around.
<emphasis id="B955DE516E33FF92FB9FFC8CFA99F434" bold="true" box="[1058,1299,770,796]" pageId="1" pageNumber="295">Capsule (immature)</emphasis>
pendent on a stalk, tepals caducous, capsule ca.
<quantity id="4CD9AFA66E33FF92FDA9FCA8FDEFF469" box="[532,613,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="295" unit="mm" value="15.0">15 mm</quantity>
long,
<quantity id="4CD9AFA66E33FF92FD13FCA8FD75F469" box="[686,767,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="295" unit="mm" value="20.0">20 mm</quantity>
thick (wings included), slightly fleshy, wings unequal, very narrowly crescent-shaped, rounded to obtuse proximally, rounded distally.
</paragraph>
</subSubSection>
<caption id="DF5E52CB6E33FF92FF35F922FE92F1CF" ID-DOI="http://doi.org/10.5281/zenodo.7156784" ID-Zenodo-Dep="7156784" httpUri="https://zenodo.org/record/7156784/files/figure.png" pageId="1" pageNumber="295" startId="1.[136,234,1708,1730]" targetBox="[157,1430,896,1676]" targetPageId="1">
<paragraph id="8B9E02436E33FF92FF35F922FE92F1CF" blockId="1.[136,1451,1708,1767]" pageId="1" pageNumber="295">
<emphasis id="B955DE516E33FF92FF35F922FE8BF1E9" bold="true" box="[136,257,1708,1730]" pageId="1" pageNumber="295">FIGURE 1.</emphasis>
Distribution map of
<taxonomicName id="4C2179C06E33FF92FE7EF922FDDFF1E9" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[451,597,1708,1729]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E33FF92FE7EF922FDDFF1E9" bold="true" box="[451,597,1708,1729]" italics="true" pageId="1" pageNumber="295">Begonia tuanii</emphasis>
</taxonomicName>
(yellow star),
<taxonomicName id="4C2179C06E33FF92FD66F923FCA7F1EA" box="[731,813,1708,1730]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis id="B955DE516E33FF92FD66F923FCA7F1EA" bold="true" box="[731,813,1708,1730]" italics="true" pageId="1" pageNumber="295">B. fordii</emphasis>
</taxonomicName>
(pink circle),
<taxonomicName id="4C2179C06E33FF92FC12F923FBDAF1EA" authorityName="C. Y. Wu" authorityYear="1995" box="[943,1104,1708,1730]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="hymenocarpa">
<emphasis id="B955DE516E33FF92FC12F923FBDAF1EA" bold="true" box="[943,1104,1708,1730]" italics="true" pageId="1" pageNumber="295">B. hymenocarpa</emphasis>
</taxonomicName>
(green square) and
<taxonomicName id="4C2179C06E33FF92FAB5F923FA21F1EA" authorityName="C. M. Hu ex C. Y. Wu &amp; T. C. Ku. A, B. Habits" authorityYear="1995" box="[1288,1451,1708,1730]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="wenshanensis">
<emphasis id="B955DE516E33FF92FAB5F923FA21F1EA" bold="true" box="[1288,1451,1708,1730]" italics="true" pageId="1" pageNumber="295">B. wenshanensis</emphasis>
</taxonomicName>
(blue triangle).
</paragraph>
</caption>
<subSubSection id="C33B51C86E33FF92FF00F89BFAD0F0B3" pageId="1" pageNumber="295" type="distribution">
<paragraph id="8B9E02436E33FF92FF00F89BFAD0F0B3" blockId="1.[136,1452,1813,2092]" pageId="1" pageNumber="295">
<emphasis id="B955DE516E33FF92FF00F89BFD85F007" bold="true" box="[189,527,1813,1839]" pageId="1" pageNumber="295">Distribution and ecology:—</emphasis>
For now,
<taxonomicName id="4C2179C06E33FF92FD3CF898FCA0F007" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[641,810,1814,1839]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E33FF92FD3CF898FCA0F007" box="[641,810,1814,1839]" italics="true" pageId="1" pageNumber="295">Begonia tuanii</emphasis>
</taxonomicName>
is known only from Ta Phoi Commune,
<collectingRegion id="49E5CCA16E33FF92FAB2F898FAE6F007" box="[1295,1388,1813,1840]" country="Vietnam" name="Lao Cai" pageId="1" pageNumber="295">Lao Cai</collectingRegion>
City,
<collectingRegion id="49E5CCA16E33FF92FF35F8B4FEDAF07B" box="[136,336,1849,1876]" country="Vietnam" name="Lao Cai" pageId="1" pageNumber="295">Lao Cai Province</collectingRegion>
and its vicinity. The new species grows on humid lands in the secondary broad-leaved evergreen forest at elevations of
<emphasis id="B955DE516E33FF92FE36F8D0FE27F05F" box="[395,429,1886,1911]" italics="true" pageId="1" pageNumber="295">ca.</emphasis>
<quantity id="4CD9AFA66E33FF92FE0BF8D3FD9DF050" box="[438,535,1885,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.258" pageId="1" pageNumber="295" unit="m" value="1258.0">1,258 m</quantity>
, together with
<taxonomicName id="4C2179C06E33FF92FD7BF8D3FCB4F05F" box="[710,830,1885,1911]" class="Magnoliopsida" family="Urticaceae" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="family">Urticaceae</taxonomicName>
(
<taxonomicName id="4C2179C06E33FF92FCEDF8D3FC73F05F" box="[848,1017,1885,1911]" class="Magnoliopsida" family="Urticaceae" genus="Elatostema" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B955DE516E33FF92FCEDF8D3FC44F05F" box="[848,974,1885,1911]" italics="true" pageId="1" pageNumber="295">Elatostema</emphasis>
sp.
</taxonomicName>
),
<taxonomicName id="4C2179C06E33FF92FBAFF8D3FB24F05F" box="[1042,1198,1885,1911]" class="Magnoliopsida" family="Polygonaceae" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="family">Polygonaceae</taxonomicName>
(
<taxonomicName id="4C2179C06E33FF92FB7DF8D3FAE1F05F" box="[1216,1387,1885,1911]" class="Magnoliopsida" family="Polygonaceae" genus="Polygonum" kingdom="Plantae" order="Caryophyllales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B955DE516E33FF92FB7DF8D3FACAF05F" box="[1216,1344,1885,1911]" italics="true" pageId="1" pageNumber="295">Polygonum</emphasis>
sp.
</taxonomicName>
), and
<taxonomicName id="4C2179C06E33FF92FF35F80FFEBAF0B3" box="[136,304,1921,1947]" class="Magnoliopsida" family="Balsaminaceae" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="family">Balsaminaceae</taxonomicName>
(
<emphasis id="B955DE516E33FF92FEFDF80CFE27F0B3" box="[320,429,1922,1947]" italics="true" pageId="1" pageNumber="295">Impetiens</emphasis>
sp.),
<taxonomicName id="4C2179C06E33FF92FE56F80FFDF0F0B3" box="[491,634,1921,1947]" class="Magnoliopsida" family="Begoniaceae" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="family">Begoniaceae</taxonomicName>
(
<taxonomicName id="4C2179C06E33FF92FD36F80CFC23F0B3" authority="Irmsch." authorityName="Irmsch." box="[651,937,1921,1947]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="handelii">
<emphasis id="B955DE516E33FF92FD36F80CFCC1F0B3" box="[651,843,1921,1947]" italics="true" pageId="1" pageNumber="295">Begonia handelii</emphasis>
Irmsch.
</taxonomicName>
) and
<taxonomicName id="4C2179C06E33FF92FC5BF80FFB1EF0B3" box="[998,1172,1921,1947]" class="Lycopodiopsida" family="Selaginellaceae" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="family">Selaginellaceae</taxonomicName>
(
<taxonomicName id="4C2179C06E33FF92FB18F80FFAC6F0B3" box="[1189,1356,1921,1947]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B955DE516E33FF92FB18F80FFAA9F0B3" box="[1189,1315,1921,1947]" italics="true" pageId="1" pageNumber="295">Selaginella</emphasis>
sp.
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B51C86E33FF92FF00F82BFC4CF0CB" pageId="1" pageNumber="295" type="etymology">
<paragraph id="8B9E02436E33FF92FF00F82BFC4CF0CB" blockId="1.[136,1452,1813,2092]" pageId="1" pageNumber="295">
<emphasis id="B955DE516E33FF92FF00F82BFEEEF097" bold="true" box="[189,356,1957,1983]" pageId="1" pageNumber="295">Etymology:—</emphasis>
The specific epithet is named in honor of Mr. Nguyen Van Tuan of the Silviculture Research Institute (SRI), who first collected the
<typeStatus id="549ABCE16E33FF92FD85F844FDE2F0CC" box="[568,616,1994,2020]" pageId="1" pageNumber="295">type</typeStatus>
specimens of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C33B51C86E33FF91FF00F863FB05F7F4" lastPageId="2" lastPageNumber="296" pageId="1" pageNumber="295" type="conservation">
<paragraph id="8B9E02436E33FF91FF00F863FB05F7F4" blockId="1.[136,1452,1813,2092]" lastBlockId="2.[136,1451,158,221]" lastPageId="2" lastPageNumber="296" pageId="1" pageNumber="295">
<emphasis id="B955DE516E33FF92FF00F863FDCFFF2F" bold="true" box="[189,581,2029,2055]" pageId="1" pageNumber="295">Proposed conservation status:—</emphasis>
Currently,
<taxonomicName id="4C2179C06E33FF92FD7CF860FCE0FF2F" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[705,874,2030,2055]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="295" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E33FF92FD7CF860FCE0FF2F" box="[705,874,2030,2055]" italics="true" pageId="1" pageNumber="295">Begonia tuanii</emphasis>
</taxonomicName>
is only found in the Ta Phoi Commune which is likely to represent its full natural range. This new species has quite a narrow distribution that is not included in any protected forest, with only two known populations of fewer than 30 individuals. We consider an IUCN category of Endangered (EN D) to be appropriate (
<bibRefCitation id="EFB07FB26E30FF91FD86FF4CFB09F7F4" author="IUCN Standards &amp; Petitions Subcommittee" box="[571,1155,194,220]" pageId="2" pageNumber="296" refId="ref2837" refString="IUCN Standards and Petitions Subcommittee (2019) Guidelines for Using the IUCN Red List Categories and Criteria, Version 14. Prepared by the Standards and Petitions Subcommittee. Available from: https: // www. iucnredlist. org / resources / redlistguidelines (accessed 26 January 2022)" type="url" year="2019">IUCN Standards and Petitions Subcommittee, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5E52CB6E30FF91FF35F829FBF9FF2D" ID-DOI="http://doi.org/10.5281/zenodo.7156786" ID-Zenodo-Dep="7156786" httpUri="https://zenodo.org/record/7156786/files/figure.png" pageId="2" pageNumber="296" startId="2.[136,234,1959,1981]" targetBox="[199,1389,264,1905]" targetPageId="2">
<paragraph id="8B9E02436E30FF91FF35F829FBF9FF2D" blockId="2.[136,1452,1959,2053]" pageId="2" pageNumber="296">
<emphasis id="B955DE516E30FF91FF35F829FE88F094" bold="true" box="[136,258,1959,1981]" pageId="2" pageNumber="296">FIGURE 2.</emphasis>
<taxonomicName id="4C2179C06E30FF91FEB5F829FD5DF095" authority="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[264,727,1959,1981]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="2" pageNumber="296" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E30FF91FEB5F829FE16F094" bold="true" box="[264,412,1959,1980]" italics="true" pageId="2" pageNumber="296">Begonia tuanii</emphasis>
T.S.Hoang &amp; C.W.Lin. A. Habit
</taxonomicName>
; B, B’. Portion of leaf, upper and lower surfaces; C. Stipule; D–D’’. Bracts; D’’’. Bracteole; E, E’. Staminate flower, face and side views; F, F’, F’’. Stamens dorsal, ventral and side views; G, G’. Pistillate flower, face and side views; H, H’. Style and stigmatic band, dorsal and ventral views; I. Cross section of ovary.
</paragraph>
</caption>
<caption id="DF5E52CB6E31FF90FF35F801FAECF0C5" ID-DOI="http://doi.org/10.5281/zenodo.7156788" ID-Zenodo-Dep="7156788" httpUri="https://zenodo.org/record/7156788/files/figure.png" pageId="3" pageNumber="297" startId="3.[136,234,1935,1957]" targetBox="[151,1436,190,1911]" targetPageId="3">
<paragraph id="8B9E02436E31FF90FF35F801FAECF0C5" blockId="3.[136,1452,1935,2030]" pageId="3" pageNumber="297">
<emphasis id="B955DE516E31FF90FF35F801FE8EF08D" bold="true" box="[136,260,1935,1957]" pageId="3" pageNumber="297">FIGURE 3.</emphasis>
<taxonomicName id="4C2179C06E31FF90FEB6F81EFD71F08E" authority="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[267,763,1936,1958]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="297" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E31FF90FEB6F81EFE15F08D" bold="true" box="[267,415,1936,1957]" italics="true" pageId="3" pageNumber="297">Begonia tuanii</emphasis>
T.S.Hoang &amp; C.W.Lin. A, B. Habit
</taxonomicName>
and habitat; C. Leaf, upper and lower views; D. Portion of leaf, upper surface, E. same, lower surface; F. Inflorescence; G. Stipule, also showing stem and petioles; H. Immature inflorescence, also showing bracts; I. Staminate flowers; J. Androecium; K. Pistillate flower; L. Cross section of ovary. All photos from by Hoang Thanh Son.
</paragraph>
</caption>
<caption id="DF5E52CB6E36FF97FF35F97AFA8CF006" ID-DOI="http://doi.org/10.5281/zenodo.7156790" ID-Zenodo-Dep="7156790" httpUri="https://zenodo.org/record/7156790/files/figure.png" pageId="4" pageNumber="298" startId="4.[136,234,1780,1802]" targetBox="[158,1430,190,1756]" targetPageId="4">
<paragraph id="8B9E02436E36FF97FF35F97AFA8CF006" blockId="4.[136,1452,1780,1838]" pageId="4" pageNumber="298">
<emphasis id="B955DE516E36FF97FF35F97AFE8EF021" bold="true" box="[136,260,1780,1802]" pageId="4" pageNumber="298">FIGURE 4.</emphasis>
<taxonomicName id="4C2179C06E36FF97FEB1F97AFC20F022" authority="C. M. Hu ex C. Y. Wu &amp; T. C. Ku. A, B. Habits" authorityName="C. M. Hu ex C. Y. Wu &amp; T. C. Ku. A, B. Habits" authorityYear="1995" box="[268,938,1780,1802]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="wenshanensis">
<emphasis id="B955DE516E36FF97FEB1F97AFE65F022" bold="true" box="[268,495,1780,1802]" italics="true" pageId="4" pageNumber="298">Begonia wenshanensis</emphasis>
C.M.Hu
<emphasis id="B955DE516E36FF97FDF3F97BFDE9F022" box="[590,611,1781,1802]" italics="true" pageId="4" pageNumber="298">ex</emphasis>
C.Y.Wu &amp; T.C.Ku. A, B. Habits
</taxonomicName>
; C. Immature inflorescence, also showing bracts; D. Inflorescence; E. Staminate flowers, face and side views; F. Pistillate flower. All photos from
<emphasis id="B955DE516E36FF97FBB6F897FBF5F005" box="[1035,1151,1816,1838]" italics="true" pageId="4" pageNumber="298">Peng 20516</emphasis>
by C.-I Peng.
</paragraph>
</caption>
<subSubSection id="C33B51C86E36FF96FF00F8D3FC73F60C" lastPageId="5" lastPageNumber="299" pageId="4" pageNumber="298" type="discussion">
<paragraph id="8B9E02436E36FF96FF00F8D3FC73F60C" blockId="4.[136,1451,1885,2091]" lastBlockId="5.[136,1452,158,292]" lastPageId="5" lastPageNumber="299" pageId="4" pageNumber="298">
<emphasis id="B955DE516E36FF97FF00F8D3FEACF05F" bold="true" box="[189,294,1885,1911]" pageId="4" pageNumber="298">Notes:—</emphasis>
Like
<taxonomicName id="4C2179C06E36FF97FEE2F8D0FD6EF050" authority="Irmsch. (1939: 501)" authorityName="Irmsch." authorityPageNumber="501" authorityYear="1939" box="[351,740,1885,1912]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="fordii">
<emphasis id="B955DE516E36FF97FEE2F8D0FE75F05F" box="[351,511,1885,1911]" italics="true" pageId="4" pageNumber="298">Begonia fordii</emphasis>
Irmsch. (1939: 501)
</taxonomicName>
,
<taxonomicName id="4C2179C06E36FF97FD52F8D0FCC4F05F" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[751,846,1886,1911]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E36FF97FD52F8D0FCC4F05F" box="[751,846,1886,1911]" italics="true" pageId="4" pageNumber="298">B. tuanii</emphasis>
</taxonomicName>
has erect stem, ovate leaves and axillary inflorescence with 4-tepaled staminate flowers and 5-tepaled pistillate flower with 3-locular ovaries, however, both species are different in many respects (see
<tableCitation id="C6A337F86E36FF97FE52F82BFDC8F097" box="[495,578,1957,1983]" captionStart="TABLE 1" captionStartId="5.[136,224,340,364]" captionTargetPageId="5" captionText="TABLE 1. Comparison of Begonia tuanii T.S.Hoang &amp; C.W.Lin, B. fordii Irmsch., B. hymenocarpa C.Y.Wu and B. wenshanensis C.M.Hu ex C.Y.Wu &amp; T.C.Ku" httpUri="http://table.plazi.org/id/DF5E52CB6E37FF96FF35FEDAFDE8F6B8" pageId="4" pageNumber="298" tableUuid="DF5E52CB6E37FF96FF35FEDAFDE8F6B8">Table 1</tableCitation>
). Among the begonias of
<emphasis id="B955DE516E36FF97FCD2F828FC0AF097" box="[879,896,1958,1983]" italics="true" pageId="4" pageNumber="298">B</emphasis>
. sect.
<taxonomicName id="4C2179C06E36FF97FC76F82BFBE8F097" authorityName="A. de Candolle" authorityYear="1859" baseAuthorityName="Klotzsch" box="[971,1122,1957,1983]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="294" phylum="Tracheophyta" rank="section" section="Platycentrum">
<emphasis id="B955DE516E36FF97FC76F82BFBE8F097" box="[971,1122,1957,1983]" italics="true" pageId="4" pageNumber="298">Platycentrum</emphasis>
</taxonomicName>
in
<collectingCountry id="F33642D36E36FF97FB34F82BFB61F097" box="[1161,1259,1957,1983]" name="Vietnam" pageId="4" pageNumber="298">Vietnam</collectingCountry>
,
<taxonomicName id="4C2179C06E36FF97FB45F828FAD0F097" authorityName="T. S. Hoang &amp; C. W. Lin. A, B. Habit" authorityYear="2022" box="[1272,1370,1958,1983]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="tuanii">
<emphasis id="B955DE516E36FF97FB45F828FAD0F097" box="[1272,1370,1958,1983]" italics="true" pageId="4" pageNumber="298">B. tuanii</emphasis>
</taxonomicName>
is also somewhat similar with
<taxonomicName id="4C2179C06E36FF97FE2CF844FBBEF0CB" authority="C. M. Hu ex C. Y. Wu &amp; T. C. Ku (1995: 262)" authorityName="C. M. Hu ex C. Y. Wu &amp; T. C. Ku. A, B. Habits" authorityPageNumber="262" authorityYear="1995" box="[401,1076,1993,2020]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="wenshanensis">
<emphasis id="B955DE516E36FF97FE2CF844FDC1F0CB" box="[401,587,1993,2019]" italics="true" pageId="4" pageNumber="298">B. wenshanensis</emphasis>
C.M.Hu
<emphasis id="B955DE516E36FF97FD0AF844FD5AF0CB" box="[695,720,1994,2019]" italics="true" pageId="4" pageNumber="298">ex</emphasis>
C.Y.Wu &amp; T.C.Ku (1995: 262)
</taxonomicName>
(
<figureCitation id="131A1EC66E36FF97FBF9F847FB03F0CB" box="[1092,1161,1993,2019]" captionStart="FIGURE 4" captionStartId="4.[136,234,1780,1802]" captionTargetBox="[158,1430,190,1756]" captionTargetId="figure-17@4.[158,1430,190,1756]" captionTargetPageId="4" captionText="FIGURE 4. Begonia wenshanensis C.M.Hu ex C.Y.Wu &amp; T.C.Ku. A, B. Habits; C. Immature inflorescence, also showing bracts; D. Inflorescence; E. Staminate flowers, face and side views; F. Pistillate flower. All photos from Peng 20516 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.7156790" httpUri="https://zenodo.org/record/7156790/files/figure.png" pageId="4" pageNumber="298">Fig. 4</figureCitation>
), but the new species can be easily told apart from
<taxonomicName id="4C2179C06E36FF97FE12F860FDE1FF2F" authorityName="C. M. Hu ex C. Y. Wu &amp; T. C. Ku. A, B. Habits" authorityYear="1995" box="[431,619,2029,2055]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="298" phylum="Tracheophyta" rank="species" species="wenshanensis">
<emphasis id="B955DE516E36FF97FE12F860FDE1FF2F" box="[431,619,2029,2055]" italics="true" pageId="4" pageNumber="298">B. wenshanensis</emphasis>
</taxonomicName>
by its shorter height, pistillate flowers with 5 tepals (vs. 3) and abaxial wing very narrowly crescent-shaped (vs. narrowly triangular or falcate). In addition, the new species also resembles
<taxonomicName id="4C2179C06E37FF96FF35FF11FDA5F791" authority="C. Y. Wu (1995: 256)" authorityName="C. Y. Wu" authorityPageNumber="256" authorityYear="1995" box="[136,559,158,185]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="299" phylum="Tracheophyta" rank="species" species="hymenocarpa">
<emphasis id="B955DE516E37FF96FF35FF11FECAF790" box="[136,320,158,184]" italics="true" pageId="5" pageNumber="299">B. hymenocarpa</emphasis>
C.Y.
<bibRefCitation id="EFB07FB26E37FF96FECAFF10FDA5F791" author="Wu, C. Y. &amp; T. C. Ku" box="[375,559,158,185]" pageId="5" pageNumber="299" pagination="251 - 280" refId="ref3264" refString="Wu, C. Y. &amp; T. C. Ku (1995) New taxa of the Begonia L. (Begoniaceae) from China. Acta Phytotaxonomica Sinica 33 (3): 251 - 280." type="journal article" year="1995">Wu (1995: 256)</bibRefCitation>
</taxonomicName>
in vegetative characters, both have erect cane-like habit, ovate leaves, axillary inflorescence and 3-loculate ovary with the placentation axile and bilamellate placentae. However, it is clearly distinct from
<taxonomicName id="4C2179C06E37FF96FEA2FF69FE5CF628" authorityName="C. Y. Wu" authorityYear="1995" box="[287,470,230,256]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="299" phylum="Tracheophyta" rank="species" species="hymenocarpa">
<emphasis id="B955DE516E37FF96FEA2FF69FE5CF628" box="[287,470,230,256]" italics="true" pageId="5" pageNumber="299">B. hymenocarpa</emphasis>
</taxonomicName>
in its larger leaves and tepals, glabrous staminate flowers and slightly fleshy capsule. A comparison of the salient characters of the four species is presented in
<tableCitation id="C6A337F86E37FF96FC22FE84FC79F60C" box="[927,1011,266,292]" captionStart="TABLE 1" captionStartId="5.[136,224,340,364]" captionTargetPageId="5" captionText="TABLE 1. Comparison of Begonia tuanii T.S.Hoang &amp; C.W.Lin, B. fordii Irmsch., B. hymenocarpa C.Y.Wu and B. wenshanensis C.M.Hu ex C.Y.Wu &amp; T.C.Ku" httpUri="http://table.plazi.org/id/DF5E52CB6E37FF96FF35FEDAFDE8F6B8" pageId="5" pageNumber="299" tableUuid="DF5E52CB6E37FF96FF35FEDAFDE8F6B8">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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