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<mods:title id="EE4C1DB171CAEB41D49605ED244E24F1">Taxonomic revision of Marma Simon, 1902 (Araneae: Salticidae: Euophryini)</mods:title>
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<taxonomicName id="4CE9F584FF9BFF85538BFBDB0DCDFBD0" ID-CoL="3Y7VQ" authority="Simon, 1902" authorityName="Simon" authorityYear="1902" box="[151,471,1086,1113]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
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<emphasis id="B99D5215FF9BFF85538BFBDB0D2CFBD1" bold="true" box="[151,310,1086,1113]" italics="true" pageId="5" pageNumber="292">Marma baeri</emphasis>
<bibRefCitation id="EF78F3F6FF9BFF855221FBDA0DCDFBD0" author="Simon, E." box="[317,471,1086,1112]" pageId="5" pageNumber="292" pagination="24 - 56" refId="ref29638" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406. https: // doi. org / 10.5962 / bhl. part. 19237" type="journal article" year="1902">Simon, 1902</bibRefCitation>
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<figureCitation id="13D29282FF9BFF85538BFB870D36FBF5" box="[151,300,1123,1149]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStart-9="FIGURE 11" captionStartId-0="6.[151,250,1476,1500]" captionStartId-1="7.[151,250,1348,1372]" captionStartId-2="8.[151,250,1616,1640]" captionStartId-3="9.[151,250,1181,1205]" captionStartId-4="10.[151,250,1503,1527]" captionStartId-5="11.[151,250,1853,1877]" captionStartId-6="12.[151,250,1856,1880]" captionStartId-7="13.[151,250,1261,1285]" captionStartId-8="14.[151,250,1371,1395]" captionStartId-9="15.[151,250,1485,1509]" captionTargetBox-0="[151,1435,237,1452]" captionTargetBox-1="[151,1435,181,1324]" captionTargetBox-2="[151,1435,181,1592]" captionTargetBox-3="[156,1431,185,1129]" captionTargetBox-4="[151,1435,181,1480]" captionTargetBox-5="[158,1432,205,1821]" captionTargetBox-6="[151,1435,181,1832]" captionTargetBox-7="[151,1435,181,1237]" captionTargetBox-8="[151,1435,181,1347]" captionTargetBox-9="[151,1435,181,1462]" captionTargetId-0="figure-48@6.[151,1435,237,1452]" captionTargetId-1="figure-17@7.[151,1435,181,1324]" captionTargetId-2="figure-24@8.[151,1435,181,1592]" captionTargetId-3="figure-17@9.[151,1435,181,1157]" captionTargetId-4="figure-24@10.[151,1435,181,1480]" captionTargetId-5="figure-17@11.[151,1435,181,1830]" captionTargetId-6="figure-24@12.[151,1435,181,1832]" captionTargetId-7="figure-17@13.[151,1435,181,1237]" captionTargetId-8="figure-24@14.[151,1435,181,1347]" captionTargetId-9="figure-17@15.[151,1435,181,1462]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURE 2. Marma baeri Simon, 1902. Live specimens from Montañita/Santa Elena/Ecuador. AB male (A dorsal, B lateral); CD female (C dorsal, D lateral)." captionText-1="FIGURE 3. Marma baeri Simon, 1902. Live specimens from Montañita/Santa Elena/Ecuador.AB male (A frontal, B detail of face); CD female (C frontal, D detail of face). Black arrow in A shows black tarsus in male front leg." captionText-2="FIGURE 4. Marma baeri Simon, 1902 (Collection of E. Simon deposited in the MNHN). AC male holotype (A dorsal; B ventral; C left male palp, retroventral). DF female allotype (D dorsal; E ventral; F epigyne, ventral). G label data." captionText-3="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." captionText-4="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." captionText-5="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." captionText-6="FIGURE 8. Female of Marma baeri Simon, 1902.AF cephalothorax structures (A detail of a white scale and a bristle; B bristle base; C scale base; D carapace, dorsal; E detail of brown and white scales; F detail of scales and bristles around the ALE). Ab- breviations: ALE—anterior lateral eye; AME—anterior median eye; PLE—posterior lateral eye; PME—posterior median eye. Black arrows show black bristles; white arrows show white scales; brown arrow in E shows brown scale." captionText-7="FIGURE 9. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador.AF left palp (A bulb, ventral; B tip of embolus and PED, ventral; C dissected embolic disc and embolus, dorsal; D detail of embolus shaft, dorsal; E cymbial conductor, ventral; F proximal region of cymbium, ventral).Abbreviations: cc—cymbial conductor; e—embolus shaft; ed—em- bolic disc; PED—process on embolic disc. Black arrow in C shows groove; white arrows in D show spikes of embolus shaft and white arrow in F shows retroventral projection of cymbium." captionText-8="FIGURE 10. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador. AD Epigyne/vulva (A epigynal plate, ventral; B right half, dorsal; C detail of right primary spermatheca and fertilization duct, dorsal; D detail of initial portion of right copulatory duct, dorsal). Abbreviations: BG—Bennetts gland; co—copulatory openings; fd—fertilization duct; PS—primary spermatheca; SS—secondary spermatheca. White arrows in D show glands of the copulatory duct near the copulatory opening." captionText-9="FIGURE 11. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador.AF spinnerets, ventro-distal (A all spinnerets, B right anterior lateral spinneret, C left anterior lateral spinneret, D right posterior lateral spinneret, E left posterior lateral spinneret, F posterior median spinnerets). G setae around anal tubercle. H epiandric spigots. I detail of right group of epiandric spigots.Abbreviations: ac—aciniform spigot; ALS—anterior lateral spinneret; eps—epiandric spigots; MAP—ma- jor ampullate spigot; mAP—minor ampullate spigot; pi—piriform spigot; PLS—posterior lateral spinneret; PMS—posterior median spinneret; ta—tartipore. Yellow arrows in A and G show white scales." figureDoi-0="http://doi.org/10.5281/zenodo.4456837" figureDoi-1="http://doi.org/10.5281/zenodo.4456841" figureDoi-2="http://doi.org/10.5281/zenodo.4456843" figureDoi-3="http://doi.org/10.5281/zenodo.4456845" figureDoi-4="http://doi.org/10.5281/zenodo.4456849" figureDoi-5="http://doi.org/10.5281/zenodo.4456851" figureDoi-6="http://doi.org/10.5281/zenodo.4456855" figureDoi-7="http://doi.org/10.5281/zenodo.4456859" figureDoi-8="http://doi.org/10.5281/zenodo.4456863" figureDoi-9="http://doi.org/10.5281/zenodo.4456865" httpUri-0="https://zenodo.org/record/4456837/files/figure.png" httpUri-1="https://zenodo.org/record/4456841/files/figure.png" httpUri-2="https://zenodo.org/record/4456843/files/figure.png" httpUri-3="https://zenodo.org/record/4456845/files/figure.png" httpUri-4="https://zenodo.org/record/4456849/files/figure.png" httpUri-5="https://zenodo.org/record/4456851/files/figure.png" httpUri-6="https://zenodo.org/record/4456855/files/figure.png" httpUri-7="https://zenodo.org/record/4456859/files/figure.png" httpUri-8="https://zenodo.org/record/4456863/files/figure.png" httpUri-9="https://zenodo.org/record/4456865/files/figure.png" pageId="5" pageNumber="292">Figures 211</figureCitation>
,
<figureCitation id="13D29282FF9BFF85522BFB860D70FBF5" box="[311,362,1122,1149]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="5" pageNumber="292">57A</figureCitation>
,
<figureCitation id="13D29282FF9BFF855269FB860DBCFBF5" box="[373,422,1122,1149]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="5" pageNumber="292">58A</figureCitation>
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<emphasis id="B99D5215FF9BFF85538BFB4C0D07FB37" box="[151,285,1191,1215]" italics="true" pageId="5" pageNumber="292">Marma baeri</emphasis>
<bibRefCitation id="EF78F3F6FF9BFF85523FFB430DCDFB37" author="Simon, E." box="[291,471,1191,1215]" pageId="5" pageNumber="292" pagination="24 - 56" refId="ref29638" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406. https: // doi. org / 10.5962 / bhl. part. 19237" type="journal article" year="1902">Simon, 1902: 376</bibRefCitation>
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(
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♁:
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<collectingCountry id="F3FECE97FF9BFF855176FB430EF4FB37" box="[618,750,1191,1215]" name="Ecuador" pageId="5" pageNumber="292">ECUADOR</collectingCountry>
:
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Túmbez [
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,
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], leg. G.A. Baer, deposited in MNHN, collection of E. Simon;
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, same data as
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, both examined by photographs);
<treatmentCitation id="0A48A816FF9BFF8557FCFB220983FB56" author="Galiano, M. E." box="[1248,1433,1222,1246]" page="36" pageId="5" pageNumber="292" year="1962">
<bibRefCitation id="EF78F3F6FF9BFF8557FCFB220983FB56" author="Galiano, M. E." box="[1248,1433,1222,1246]" pageId="5" pageNumber="292" pagination="31 - 44" refId="ref28892" refString="Galiano, M. E. (1962) Los generos Amphidraus Simon y Marma Simon (Araneae, Salticidae). Acta Zoologica Lilloana, 18, 31 - 44." type="journal article" year="1962">Galiano, 1962: 36</bibRefCitation>
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, pl. I, figs 15;
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<bibRefCitation id="EF78F3F6FF9BFF855240FB000E3BFB74" author="Galiano, M. E." box="[348,545,1252,1276]" pageId="5" pageNumber="292" pagination="273 - 470" refId="ref28924" refString="Galiano, M. E. (1963) Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C), 23, 273 - 470." type="journal article" year="1963">Galiano, 1963: 395</bibRefCitation>
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, pl. XXVIII, figs 710.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B568E07FF9BFF85538BFACF0E0DFAE1" blockId="5.[151,1437,1323,2033]" pageId="5" pageNumber="292">
<emphasis id="B99D5215FF9BFF85538BFACF0CCCFACD" bold="true" box="[151,214,1323,1349]" pageId="5" pageNumber="292">Note.</emphasis>
Both sexes are redescribed based on newly collected specimens. Photographs of the
<typeStatus id="545230A5FF9BFF8557B2FACF08C4FACD" box="[1198,1246,1323,1349]" pageId="5" pageNumber="292">type</typeStatus>
specimens were provided by MNHN (
<figureCitation id="13D29282FF9BFF855295FAAB0DCAFAE1" box="[393,464,1359,1385]" captionStart="FIGURE 4" captionStartId="8.[151,250,1616,1640]" captionTargetBox="[151,1435,181,1592]" captionTargetId="figure-24@8.[151,1435,181,1592]" captionTargetPageId="8" captionText="FIGURE 4. Marma baeri Simon, 1902 (Collection of E. Simon deposited in the MNHN). AC male holotype (A dorsal; B ventral; C left male palp, retroventral). DF female allotype (D dorsal; E ventral; F epigyne, ventral). G label data." figureDoi="http://doi.org/10.5281/zenodo.4456843" httpUri="https://zenodo.org/record/4456843/files/figure.png" pageId="5" pageNumber="292">Figs 4</figureCitation>
AG).
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFF9BFF8553DBFA960D9BF8B5" pageId="5" pageNumber="292" type="diagnosis">
<paragraph id="8B568E07FF9BFF8553DBFA960D9BF8B5" blockId="5.[151,1437,1323,2033]" pageId="5" pageNumber="292">
<emphasis id="B99D5215FF9BFF8553DBFA960D58FA04" bold="true" box="[199,322,1394,1420]" pageId="5" pageNumber="292">Diagnosis.</emphasis>
The males of
<taxonomicName id="4CE9F584FF9BFF8552C2FA970E27FA05" authorityName="Simon" authorityYear="1902" box="[478,573,1395,1421]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF8552C2FA970E27FA05" box="[478,573,1395,1421]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="4CE9F584FF9BFF85506BFA970FC9FA05" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[887,979,1395,1421]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FF9BFF85506BFA970FC9FA05" box="[887,979,1395,1421]" italics="true" pageId="5" pageNumber="292">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FF9BFF8550C6FA970828FA05" bold="true" box="[986,1074,1395,1421]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A2AEEF6EFF9BFF8550C6FA970828FA05" box="[986,1074,1395,1421]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a short process on the embolic disc (PED; 5C, 7A, 13C, 15A), but differ by having a thicker embolic disc (
<figureCitation id="13D29282FF9BFF85575FFA730882FA39" box="[1091,1176,1430,1457]" captionStart="FIGURE 57" captionStartId="62.[151,250,1843,1867]" captionTargetBox="[159,1422,193,1813]" captionTargetId="figure-24@62.[151,1435,181,1819]" captionTargetPageId="62" captionText="FIGURE 57. Left male palp, ventral [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. pipa sp. nov.; D M. wesolowskae sp. nov.; E M. nigritarsis (Simon, 1900); F M. sinuosa sp. nov., G M. argentina (Mello-Leit„o, 1941), H M. rosea (Mello- Leit„o, 1941), I M. spelunca sp. nov.]. Colors represent homologous structures." figureDoi="http://doi.org/10.5281/zenodo.4456979" httpUri="https://zenodo.org/record/4456979/files/figure.png" pageId="5" pageNumber="292">Figs 57</figureCitation>
AB). Besides, the embolus shaft (e) in
<taxonomicName id="4CE9F584FF9BFF85524BFA5F0DACFA5D" authorityName="Simon" authorityYear="1902" box="[343,438,1467,1493]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF85524BFA5F0DACFA5D" box="[343,438,1467,1493]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
emerges from a less proximal portion of embolic disc and is consequently shorter than the embolus of
<taxonomicName id="4CE9F584FF9BFF855207FA3B0D63FA71" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[283,377,1503,1529]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FF9BFF855207FA3B0D63FA71" box="[283,377,1503,1529]" italics="true" pageId="5" pageNumber="292">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FF9BFF85529CFA3B0DC3FA71" bold="true" box="[384,473,1503,1529]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A2AEEF6EFF9BFF85529CFA3B0DC3FA71" box="[384,473,1503,1529]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(best visualized cleared in clove oil;
<figureCitation id="13D29282FF9BFF855067FA3B0FCBFA71" box="[891,977,1503,1529]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="5" pageNumber="292">Figs 7F</figureCitation>
,
<figureCitation id="13D29282FF9BFF8550C0FA3B0FE4FA71" box="[988,1022,1503,1529]" captionStart="FIGURE 9" captionStartId="13.[151,250,1261,1285]" captionTargetBox="[151,1435,181,1237]" captionTargetId="figure-17@13.[151,1435,181,1237]" captionTargetPageId="13" captionText="FIGURE 9. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador.AF left palp (A bulb, ventral; B tip of embolus and PED, ventral; C dissected embolic disc and embolus, dorsal; D detail of embolus shaft, dorsal; E cymbial conductor, ventral; F proximal region of cymbium, ventral).Abbreviations: cc—cymbial conductor; e—embolus shaft; ed—em- bolic disc; PED—process on embolic disc. Black arrow in C shows groove; white arrows in D show spikes of embolus shaft and white arrow in F shows retroventral projection of cymbium." figureDoi="http://doi.org/10.5281/zenodo.4456859" httpUri="https://zenodo.org/record/4456859/files/figure.png" pageId="5" pageNumber="292">9C</figureCitation>
,
<figureCitation id="13D29282FF9BFF855716FA3A0820FA71" box="[1034,1082,1502,1529]" captionStart="FIGURE 15" captionStartId="19.[151,250,1015,1039]" captionTargetBox="[281,1294,194,968]" captionTargetId="figure-17@19.[279,1308,181,991]" captionTargetPageId="19" captionText="FIGURE 15. Marma linae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ven- tral); FG epigyne/vulva (F ventral; G cleared, ventral). Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456879" httpUri="https://zenodo.org/record/4456879/files/figure.png" pageId="5" pageNumber="292">15E</figureCitation>
,
<figureCitation id="13D29282FF9BFF85575AFA3B0863FA72" box="[1094,1145,1503,1530]" captionStart="FIGURE 16" captionStartId="19.[151,250,1904,1928]" captionTargetBox="[280,1306,1172,1881]" captionTargetId="figure-80@19.[280,1306,1172,1881]" captionTargetPageId="19" captionText="FIGURE 16. Male of Marma linae sp. nov. AD left palp (A dissected embolic disc, dorsal; B detail of embolus shaft, dorsal; C detail of apical region of cymbium, ventral; D detail of proximal region of cymbium, ventral). Abbreviations: cc—cymbial conductor; e—embolus shaft; PED—process on embolic disc. Black arrow in A shows groove; white arrows in B show spikes of embolus shaft; white arrow in D shows retroventral projection of cymbium." figureDoi="http://doi.org/10.5281/zenodo.4456883" httpUri="https://zenodo.org/record/4456883/files/figure.png" pageId="5" pageNumber="292">16A</figureCitation>
). The females of
<taxonomicName id="4CE9F584FF9BFF855620FA3B0986FA71" authorityName="Simon" authorityYear="1902" box="[1340,1436,1503,1529]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF855620FA3B0986FA71" box="[1340,1436,1503,1529]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4CE9F584FF9BFF85529CF9E70DC5F995" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[384,479,1539,1565]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FF9BFF85529CF9E70DC5F995" box="[384,479,1539,1565]" italics="true" pageId="5" pageNumber="292">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FF9BFF8552F4F9E70E59F995" bold="true" box="[488,579,1539,1565]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A2AEEF6EFF9BFF8552F4F9E70E59F995" box="[488,579,1539,1565]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having the copulatory openings separated by approximately three times the girth of the copulatory duct and by its placement anterior to the primary spermathecae (
<figureCitation id="13D29282FF9BFF855783F9C308E1F9C9" box="[1183,1275,1575,1601]" captionStart="FIGURE 6" captionStartId="10.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-24@10.[151,1435,181,1480]" captionTargetPageId="10" captionText="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456849" httpUri="https://zenodo.org/record/4456849/files/figure.png" pageId="5" pageNumber="292">Figs 6C</figureCitation>
,
<figureCitation id="13D29282FF9BFF855614F9C3090FF9C9" box="[1288,1301,1575,1601]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="5" pageNumber="292">7</figureCitation>
GH, 58A), whereas they are very close (separated by less than 1/2 their diameter) and placed at the same level of the primary spermathecae in
<taxonomicName id="4CE9F584FF9BFF85524FF98B0DAAF901" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[339,432,1647,1673]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FF9BFF85524FF98B0DAAF901" box="[339,432,1647,1673]" italics="true" pageId="5" pageNumber="292">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FF9BFF8552ABF98B0E0BF901" bold="true" box="[439,529,1647,1673]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A2AEEF6EFF9BFF8552ABF98B0E0BF901" box="[439,529,1647,1673]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13D29282FF9BFF85513CF98B0E93F901" box="[544,649,1647,1673]" captionStart="FIGURE 14" captionStartId="18.[151,250,1372,1396]" captionTargetBox="[157,1407,279,1323]" captionTargetId="figure-65@18.[151,1435,273,1349]" captionTargetPageId="18" captionText="FIGURE 14. Female paratype of Marma linae sp. nov. (MPEG 34346). AB habitus (A dorsal, B ventral); CD cleared epi- gyne/vulva (C ventral, D dorsal).Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fer- tilization duct; PS—primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456877" httpUri="https://zenodo.org/record/4456877/files/figure.png" pageId="5" pageNumber="292">Figs 14C</figureCitation>
,
<figureCitation id="13D29282FF9BFF855189F98A0EB5F901" box="[661,687,1646,1673]" captionStart="FIGURE 15" captionStartId="19.[151,250,1015,1039]" captionTargetBox="[281,1294,194,968]" captionTargetId="figure-17@19.[279,1308,181,991]" captionTargetPageId="19" captionText="FIGURE 15. Marma linae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ven- tral); FG epigyne/vulva (F ventral; G cleared, ventral). Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456879" httpUri="https://zenodo.org/record/4456879/files/figure.png" pageId="5" pageNumber="292">15</figureCitation>
FG, 58B). Also, females of
<taxonomicName id="4CE9F584FF9BFF8550E8F98B084EF901" authorityName="Simon" authorityYear="1902" box="[1012,1108,1647,1673]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF8550E8F98B084EF901" box="[1012,1108,1647,1673]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
have the distal portion of the primary spermathecae projecting laterally (
<figureCitation id="13D29282FF9BFF855167F9770ECDF925" box="[635,727,1683,1709]" captionStart="FIGURE 6" captionStartId="10.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-24@10.[151,1435,181,1480]" captionTargetPageId="10" captionText="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456849" httpUri="https://zenodo.org/record/4456849/files/figure.png" pageId="5" pageNumber="292">Figs 6C</figureCitation>
,
<figureCitation id="13D29282FF9BFF8551FFF9770F1DF925" box="[739,775,1683,1709]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="5" pageNumber="292">7H</figureCitation>
,
<figureCitation id="13D29282FF9BFF855008F9760F5DF925" box="[788,839,1682,1709]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="5" pageNumber="292">58A</figureCitation>
), whereas they project anteriorly in
<taxonomicName id="4CE9F584FF9BFF8557C1F9770921F925" authority="Salgado &amp; Ruiz, 2020" authorityName="Salgado &amp; Ruiz" authorityYear="2020" box="[1245,1339,1683,1709]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="linae" status="sp. nov.">
<emphasis id="B99D5215FF9BFF8557C1F9770921F925" box="[1245,1339,1683,1709]" italics="true" pageId="5" pageNumber="292">M. linae</emphasis>
</taxonomicName>
<emphasis id="B99D5215FF9BFF85565EF9770986F925" bold="true" box="[1346,1436,1683,1709]" pageId="5" pageNumber="292">
<taxonomicNameLabel id="A2AEEF6EFF9BFF85565EF9770986F925" box="[1346,1436,1683,1709]" pageId="5" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13D29282FF9BFF855383F9530D12F959" box="[159,264,1719,1745]" captionStart="FIGURE 14" captionStartId="18.[151,250,1372,1396]" captionTargetBox="[157,1407,279,1323]" captionTargetId="figure-65@18.[151,1435,273,1349]" captionTargetPageId="18" captionText="FIGURE 14. Female paratype of Marma linae sp. nov. (MPEG 34346). AB habitus (A dorsal, B ventral); CD cleared epi- gyne/vulva (C ventral, D dorsal).Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fer- tilization duct; PS—primary spermatheca; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456877" httpUri="https://zenodo.org/record/4456877/files/figure.png" pageId="5" pageNumber="292">Figs 14C</figureCitation>
,
<figureCitation id="13D29282FF9BFF855208F9520D5DF959" box="[276,327,1718,1745]" captionStart="FIGURE 15" captionStartId="19.[151,250,1015,1039]" captionTargetBox="[281,1294,194,968]" captionTargetId="figure-17@19.[279,1308,181,991]" captionTargetPageId="19" captionText="FIGURE 15. Marma linae sp. nov. AE left male palp (A ventral; B retrolateral; C proventral; D dorsal; E cleared bulb, ven- tral); FG epigyne/vulva (F ventral; G cleared, ventral). Abbreviations: BG—Bennetts gland; cc—cymbial conductor; cdcopulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PED—process on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456879" httpUri="https://zenodo.org/record/4456879/files/figure.png" pageId="5" pageNumber="292">15G</figureCitation>
,
<figureCitation id="13D29282FF9BFF85524EF9520D9FF959" box="[338,389,1718,1745]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="5" pageNumber="292">58B</figureCitation>
). Females of
<taxonomicName id="4CE9F584FF9BFF855106F9530E60F959" authorityName="Simon" authorityYear="1902" box="[538,634,1719,1745]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF855106F9530E60F959" box="[538,634,1719,1745]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
are also very similar to those of
<taxonomicName id="4CE9F584FF9BFF8550F4F9530845F959" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[1000,1119,1719,1745]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="femella">
<emphasis id="B99D5215FF9BFF8550F4F9530845F959" box="[1000,1119,1719,1745]" italics="true" pageId="5" pageNumber="292">M. femella</emphasis>
</taxonomicName>
, both with copulatory openings far from each other and the distal portion of primary spermathecae projecting laterally (
<figureCitation id="13D29282FF9BFF85578DF93F0907F97D" box="[1169,1309,1754,1781]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="5" pageNumber="292">Figs 58A, D</figureCitation>
). However, in
<taxonomicName id="4CE9F584FF9BFF8553A8F91B0D0EF891" authorityName="Simon" authorityYear="1902" box="[180,276,1791,1817]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF9BFF8553A8F91B0D0EF891" box="[180,276,1791,1817]" italics="true" pageId="5" pageNumber="292">M. baeri</emphasis>
</taxonomicName>
the initial portion of spermathecae is dilated, whereas the spermathecae of
<taxonomicName id="4CE9F584FF9BFF855743F91B08CCF891" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[1119,1238,1791,1817]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="292" phylum="Arthropoda" rank="species" species="femella">
<emphasis id="B99D5215FF9BFF855743F91B08CCF891" box="[1119,1238,1791,1817]" italics="true" pageId="5" pageNumber="292">M. femella</emphasis>
</taxonomicName>
have an elliptical shape (
<figureCitation id="13D29282FF9BFF8553FAF8C70D6FF8B5" box="[230,373,1826,1853]" captionStart="FIGURE 58" captionStartId="63.[151,250,1595,1619]" captionTargetBox="[156,1429,271,1549]" captionTargetId="figure-41@63.[151,1435,237,1571]" captionTargetPageId="63" captionText="FIGURE 58. Epigyne/vulva, AC, EK ventral, D dorsal [A Marma baeri Simon, 1902; B M. linae sp. nov.; C M. abaira sp. nov.; D M. femella (Caporiacco, 1955); E M. pipa sp. nov.; F M. wesolowskae sp. nov.; G M. nigritarsis (Simon, 1900); H M. sinuosa sp. nov.; I M. argentina (Mello-Leitão, 1941); J M. rosea (Mello-Leitão, 1941); K M. spelunca sp. nov.]. Colors represent homologous structures; arrow in D shows hypothetical position of secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456981" httpUri="https://zenodo.org/record/4456981/files/figure.png" pageId="5" pageNumber="292">Figs 58A, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFF9BFF8753DBF8A20CE4F848" lastPageId="7" lastPageNumber="294" pageId="5" pageNumber="292" type="description">
<paragraph id="8B568E07FF9BFF8553DBF8A20872F821" blockId="5.[151,1437,1323,2033]" pageId="5" pageNumber="292">
<emphasis id="B99D5215FF9BFF8553DBF8A20E14F8E8" bold="true" box="[199,526,1862,1889]" pageId="5" pageNumber="292">
Description.
<emphasis id="B99D5215FF9BFF85527CF8A20D81F8E9" bold="true" box="[352,411,1862,1889]" italics="true" pageId="5" pageNumber="292">Male</emphasis>
(QCAZ).
</emphasis>
Total length: 3.12. Carapace 1.64 long, 1.27 wide, 0.97 high. Ocular quadrangle 0.90 long. Anterior eye row 1.11 wide, posterior 0.96 wide. Legs 1432. Length of legs: I 4.30 (1.29 + 1.75 + 1.26); II 2.95 (0.99 + 1.04 + 0.92); III 3.55 (1.20 + 1.16 + 1.19); IV 3.66 (1.10 + 1.22 + 1.34).
</paragraph>
<paragraph id="8B568E07FF9BFF8653DBF8570E3BFF5D" blockId="5.[151,1437,1323,2033]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="293" pageId="5" pageNumber="292">Leg macrosetae: femur I d1-1-1, p1di, r0; II d1-1-1, p1di, r1di; III d1-1-1, p1di, r0; IV d1-1-1, p0 (or p1di), r1di; patella III 0, IIIIV p0, r1; tibia I p0-1-0, r0, v1r-2-2; II p0-1-0, r0, v1r-2-1p; III p0-1-1-0, r0-1-1-0, v1p-0- 0-2 (v2di); IV p0-1-1-0, r1-1-1-0, v2di; metatarsus I p1di, r0, v2-2; II p1-1, r1di, v2-2; III d1p-0-0, p1-0-2, r1-0-2, v2-0-2; IV p1-1-2, r1-1-2, v1p-0-2.</paragraph>
<caption id="DF96DE8FFF98FF86538BFA200DF0F988" ID-DOI="http://doi.org/10.5281/zenodo.4456837" ID-Zenodo-Dep="4456837" httpUri="https://zenodo.org/record/4456837/files/figure.png" pageId="6" pageNumber="293" startId="6.[151,250,1476,1500]" targetBox="[151,1435,237,1452]" targetPageId="6">
<paragraph id="8B568E07FF98FF86538BFA200DF0F988" blockId="6.[151,1436,1476,1536]" pageId="6" pageNumber="293">
<emphasis id="B99D5215FF98FF86538BFA200D09FA54" bold="true" box="[151,275,1476,1500]" pageId="6" pageNumber="293">FIGURE 2.</emphasis>
<taxonomicName id="4CE9F584FF98FF865205FA210E31FA54" authority="Simon, 1902" authorityName="Simon" authorityYear="1902" box="[281,555,1476,1500]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="293" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF98FF865205FA210D85FA54" box="[281,415,1476,1500]" italics="true" pageId="6" pageNumber="293">Marma baeri</emphasis>
<bibRefCitation id="EF78F3F6FF98FF8652BAFA200E31FA54" author="Simon, E." box="[422,555,1476,1500]" pageId="6" pageNumber="293" pagination="24 - 56" refId="ref29638" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406. https: // doi. org / 10.5962 / bhl. part. 19237" type="journal article" year="1902">Simon, 1902</bibRefCitation>
</taxonomicName>
. Live specimens from Montañita/Santa Elena/Ecuador. AB male (A dorsal, B lateral); CD female (C dorsal, D lateral).
</paragraph>
</caption>
<paragraph id="8B568E07FF98FF8653DBF9CA0F41F974" blockId="6.[151,1437,1582,2005]" pageId="6" pageNumber="293">
Color in alcohol (
<figureCitation id="13D29282FF98FF865291F9CA0DC9F9C1" box="[397,467,1582,1609]" captionStart="FIGURE 5" captionStartId="9.[151,250,1181,1205]" captionTargetBox="[156,1431,185,1129]" captionTargetId="figure-17@9.[151,1435,181,1157]" captionTargetPageId="9" captionText="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456845" httpUri="https://zenodo.org/record/4456845/files/figure.png" pageId="6" pageNumber="293">Figs 5</figureCitation>
AB): carapace with triangle of scales pointing backwards; regions beside the triangle with less concentrated scales; abdomen pale ventrally, with three dark brown longitudinal stripes; legs: femur I with dark distal ring and proximal region with dark prolateral spot, II with dark distal ring and proximal region with prolateral and retrolateral dark spot, IIIIV with dark distal ring and proximal region with dark proventral spot; tibia I with dark proximal ring and distal region with dark retrolateral spot, IIIV with proximal and distal dark ring; tarsus I almost black, II with dark tip, IIIIV with dark proximal ring.
</paragraph>
<paragraph id="8B568E07FF98FF8653DBF8E20F82F8E1" blockId="6.[151,1437,1582,2005]" pageId="6" pageNumber="293">
Palp: RTA finger-shaped (
<figureCitation id="13D29282FF98FF8652FBF8E20E5EF8A9" box="[487,580,1798,1825]" captionStart="FIGURE 5" captionStartId="9.[151,250,1181,1205]" captionTargetBox="[156,1431,185,1129]" captionTargetId="figure-17@9.[151,1435,181,1157]" captionTargetPageId="9" captionText="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456845" httpUri="https://zenodo.org/record/4456845/files/figure.png" pageId="6" pageNumber="293">Figs 5D</figureCitation>
,
<figureCitation id="13D29282FF98FF865153F8E20E69F8A9" box="[591,627,1798,1825]" captionStart="FIGURE 6" captionStartId="10.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-24@10.[151,1435,181,1480]" captionTargetPageId="10" captionText="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456849" httpUri="https://zenodo.org/record/4456849/files/figure.png" pageId="6" pageNumber="293">6C</figureCitation>
); embolic disc with both edges curved (
<figureCitation id="13D29282FF98FF865737F8E2089CF8A9" box="[1067,1158,1798,1825]" captionStart="FIGURE 5" captionStartId="9.[151,250,1181,1205]" captionTargetBox="[156,1431,185,1129]" captionTargetId="figure-17@9.[151,1435,181,1157]" captionTargetPageId="9" captionText="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456845" httpUri="https://zenodo.org/record/4456845/files/figure.png" pageId="6" pageNumber="293">Figs 5C</figureCitation>
,
<figureCitation id="13D29282FF98FF86578DF8E208ACF8A8" box="[1169,1206,1798,1824]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="6" pageNumber="293">7A</figureCitation>
); PED with approximately same length of exposed portion of embolic disc and emerging from the middle-distal part of embolic disc (
<figureCitation id="13D29282FF98FF865383F8AA0CE1F8E1" box="[159,251,1870,1897]" captionStart="FIGURE 5" captionStartId="9.[151,250,1181,1205]" captionTargetBox="[156,1431,185,1129]" captionTargetId="figure-17@9.[151,1435,181,1157]" captionTargetPageId="9" captionText="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456845" httpUri="https://zenodo.org/record/4456845/files/figure.png" pageId="6" pageNumber="293">Figs 5C</figureCitation>
,
<figureCitation id="13D29282FF98FF86521AF8AA0D36F8E0" box="[262,300,1870,1896]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="6" pageNumber="293">7A</figureCitation>
); tip of embolus aligned with tip of PED (
<figureCitation id="13D29282FF98FF86501EF8AA0F44F8E1" box="[770,862,1870,1897]" captionStart="FIGURE 5" captionStartId="9.[151,250,1181,1205]" captionTargetBox="[156,1431,185,1129]" captionTargetId="figure-17@9.[151,1435,181,1157]" captionTargetPageId="9" captionText="FIGURE 5. Male of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD left male palp (C ventral; D retrolateral)." figureDoi="http://doi.org/10.5281/zenodo.4456845" httpUri="https://zenodo.org/record/4456845/files/figure.png" pageId="6" pageNumber="293">Figs 5C</figureCitation>
,
<figureCitation id="13D29282FF98FF865076F8AB0F91F8E1" box="[874,907,1871,1897]" captionStart="FIGURE 7" captionStartId="11.[151,250,1853,1877]" captionTargetBox="[158,1432,205,1821]" captionTargetId="figure-17@11.[151,1435,181,1830]" captionTargetPageId="11" captionText="FIGURE 7. Marma baeri Simon, 1902. A, CH specimens from Montañita/Santa Elena/Ecuador. B holotype illustrated by Galiano (1963), reflected.AF left male palp (AB ventral; C retrolateral; D proventral; E dorsal; F cleared bulb, ventral). GH Epigyne/vulva (G ventral; H cleared, ventral). Abbreviations: BG—Bennett´s gland; cc—cymbial conductor; cd—copulatory duct; co—copulatory opening; dh—distal hematodocha; e—embolus shaft; ed—embolic disc; fd—fertilization duct; PEDprocess on embolic disc; PS—primary spermatheca; RTA—retrolateral tibial apophysis; SS—secondary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4456851" httpUri="https://zenodo.org/record/4456851/files/figure.png" pageId="6" pageNumber="293">7F</figureCitation>
).
</paragraph>
<paragraph id="8B568E07FF98FF8653DBF8950FCEF85C" blockId="6.[151,1437,1582,2005]" pageId="6" pageNumber="293">
<emphasis id="B99D5215FF98FF8653DBF8950D8BF804" bold="true" box="[199,401,1905,1932]" pageId="6" pageNumber="293">
<emphasis id="B99D5215FF98FF8653DBF8950D04F804" bold="true" box="[199,286,1905,1932]" italics="true" pageId="6" pageNumber="293">Female</emphasis>
(QCAZ).
</emphasis>
Total length: 3.51. Carapace 1.70 long, 1.20 wide, 0.81 high. Ocular quadrangle 0.73 long. Anterior eye row 1.13 wide, posterior 0.94 wide. Legs 4312. Length of legs: I 2.71 (0.86 + 1.03 + 0.82); II 2.57 (0.86 + 0.92 + 0.79); III 3.39 (1.13 + 1.16 + 1.10); IV 3.65 (1.11 + 1.23 + 1.31).
</paragraph>
<caption id="DF96DE8FFF99FF87538BFAA008BBFA08" ID-DOI="http://doi.org/10.5281/zenodo.4456841" ID-Zenodo-Dep="4456841" httpUri="https://zenodo.org/record/4456841/files/figure.png" pageId="7" pageNumber="294" startId="7.[151,250,1348,1372]" targetBox="[151,1435,181,1324]" targetPageId="7">
<paragraph id="8B568E07FF99FF87538BFAA008BBFA08" blockId="7.[151,1436,1348,1408]" pageId="7" pageNumber="294">
<emphasis id="B99D5215FF99FF87538BFAA00D08FAD4" bold="true" box="[151,274,1348,1372]" pageId="7" pageNumber="294">FIGURE 3.</emphasis>
<taxonomicName id="4CE9F584FF99FF875204FAA10E32FAD4" authority="Simon, 1902" authorityName="Simon" authorityYear="1902" box="[280,552,1348,1372]" class="Arachnida" family="Salticidae" genus="Marma" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="294" phylum="Arthropoda" rank="species" species="baeri">
<emphasis id="B99D5215FF99FF875204FAA10D84FAD4" box="[280,414,1348,1372]" italics="true" pageId="7" pageNumber="294">Marma baeri</emphasis>
<bibRefCitation id="EF78F3F6FF99FF8752B8FAA00E32FAD4" author="Simon, E." box="[420,552,1348,1372]" pageId="7" pageNumber="294" pagination="24 - 56" refId="ref29638" refString="Simon, E. (1902) Description d'arachnides nouveaux de la famille des Salticidae (Attidae) (suite). Annales de la Societe Entomologique de Belgique, 46, 24 - 56 + 363 - 406. https: // doi. org / 10.5962 / bhl. part. 19237" type="journal article" year="1902">Simon, 1902</bibRefCitation>
</taxonomicName>
. Live specimens from Montañita/Santa Elena/Ecuador. AB male (A frontal, B detail of face); CD female (C frontal, D detail of face). Black arrow in A shows black tarsus in male front leg.
</paragraph>
</caption>
<paragraph id="8B568E07FF99FF8753DBFA4B08D5F9BC" blockId="7.[151,1437,1454,2021]" pageId="7" pageNumber="294">Leg macrosetae: Femur I d1-1-1, p1di, r0; II d1-1-1, p1di, r1di; III d1-1-1, p1di, r0; IV d1-1-1, p1di (or p0), r0 (or r1di). Patella III 0, IIIIV p0, r1. Tibia III p0-1-0, r0, v1r-2-2; III p0-1-1-0, r0-1-1-0, v1p-0-0-1; IV d0-1p-0-0, p0-1-1-1 (p1-0-1-0), r1-1-1-0, v1p-0-0-2 (or 2di). Metatarsus I p1di, r0, v2-2; II p1-1, r1di, v2-2; III d1p-0-0, p1-0- 2, r1-0-2, v2-0-2; IV d1p-0-0 (or d0), p1-0-2 (or p1-1-2), r1-0-2 (or r1-1-2), v2-0-2 (or v1p-0-2).</paragraph>
<paragraph id="8B568E07FF99FF8753DBF9DA0FD7F960" blockId="7.[151,1437,1454,2021]" pageId="7" pageNumber="294">
Color in alcohol (
<figureCitation id="13D29282FF99FF875291F9DA0DCEF9D1" box="[397,468,1598,1625]" captionStart="FIGURE 6" captionStartId="10.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-24@10.[151,1435,181,1480]" captionTargetPageId="10" captionText="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456849" httpUri="https://zenodo.org/record/4456849/files/figure.png" pageId="7" pageNumber="294">Figs 6</figureCitation>
AB): carapace as in male, abdomen ventrally with three dark longitudinal stripes and with dark brown edge with pale circles; legs: femur I with dark distal ring and proximal region with dark retroventral spot, IIIII with dark distal ring and proximal region with incomplete dark ring (dorsal portion not totally colored), IV with dark distal ring and proximal region with dark proventral spot; tibia IIV with proximal and distal dark ring; tarsus III without dark marks, IIIIV with dark proximal ring.
</paragraph>
<paragraph id="8B568E07FF99FF8753DBF9160F27F8F0" blockId="7.[151,1437,1454,2021]" pageId="7" pageNumber="294">
Epigyne (
<figureCitation id="13D29282FF99FF87522FF9160D62F884" box="[307,376,1778,1804]" captionStart="FIGURE 6" captionStartId="10.[151,250,1503,1527]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-24@10.[151,1435,181,1480]" captionTargetPageId="10" captionText="FIGURE 6. Female of Marma baeri Simon, 1902, specimen from Montañita/Santa Elena/Ecuador (QCAZ). AB habitus (A dorsal, B ventral); CD cleared epigyne/vulva (C ventral, cleared; D details of secondary spermathecae, dorsal). Abbreviations: BG—Bennetts gland; cd—copulatory duct; co—copulatory opening; fd—fertilization duct; e—embolus shaft (broken inside the duct); PS—Primary spermatheca; SS—secondary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4456849" httpUri="https://zenodo.org/record/4456849/files/figure.png" pageId="7" pageNumber="294">Figs 6</figureCitation>
CD, 7GH, 10AD): copulatory openings separated by approximately three times the girth of copulatory ducts, placed anteriorly to primary spermathecae; copulatory ducts short; proximal copulatory duct with approximately same length as distal section; primary spermathecae with initial portion dilated, tapering towards fertilization ducts; primary spermathecae laterally projected.
</paragraph>
<paragraph id="8B568E07FF99FF8753DBF8660CE4F848" blockId="7.[151,1437,1454,2021]" pageId="7" pageNumber="294">
<emphasis id="B99D5215FF99FF8753DBF8660DE0F814" bold="true" box="[199,506,1922,1948]" pageId="7" pageNumber="294">Other material examined.</emphasis>
<materialsCitation id="3B81845AFF99FF87511DF8660CE0F848" ID-GBIF-Occurrence="3020125362" collectionCode="QCAZ" country="Ecuador" latitude="-1.8171666" location="Santa Elena" longLatPrecision="1" longitude="-80.75808" municipality="Montanita" pageId="7" pageNumber="294" specimenCount="1" specimenCount-female="1">
<emphasis id="B99D5215FF99FF87511DF8660E82F814" bold="true" box="[513,664,1922,1948]" pageId="7" pageNumber="294">
<collectingCountry id="F3FECE97FF99FF87511DF8660E89F814" box="[513,659,1922,1948]" name="Ecuador" pageId="7" pageNumber="294">ECUADOR</collectingCountry>
:
</emphasis>
<emphasis id="B99D5215FF99FF875182F8670F3FF814" box="[670,805,1922,1949]" italics="true" pageId="7" pageNumber="294">
<location id="8E36D8DCFF99FF875182F8670F3FF814" LSID="urn:lsid:plazi:treatment:03403F11FF9BFF87538BFBDB08C6F86D:8E36D8DCFF99FF875182F8670F3FF814" box="[670,805,1922,1949]" country="Ecuador" latitude="-1.8171666" longLatPrecision="1" longitude="-80.75808" municipality="Montanita" name="Santa Elena" pageId="7" pageNumber="294">Santa Elena</location>
</emphasis>
:
<collectingMunicipality id="6B32147DFF99FF875028F8660FB0F814" box="[820,938,1922,1948]" pageId="7" pageNumber="294">Montañita</collectingMunicipality>
,
<geoCoordinate id="EEDDE8C0FF99FF8750A8F866084AF814" box="[948,1104,1922,1948]" degrees="01" direction="south" minutes="49" orientation="latitude" pageId="7" pageNumber="294" precision="1" seconds="01.8" value="-1.8171666">01°4901.8”S</geoCoordinate>
,
<geoCoordinate id="EEDDE8C0FF99FF875747F866091AF815" box="[1115,1280,1922,1949]" degrees="80" direction="west" minutes="45" orientation="longitude" pageId="7" pageNumber="294" precision="1" seconds="29.1" value="-80.75808">80°4529.1”W</geoCoordinate>
, 2019, 1♁
<specimenCount id="9DEF458EFF99FF875665F8660986F814" box="[1401,1436,1922,1948]" pageId="7" pageNumber="294" type="female">1♀</specimenCount>
(
<collectionCode id="EDF816C2FF99FF875383F8420CE8F848" box="[159,242,1958,1984]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="7" pageNumber="294" type="Museum">QCAZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3F3DD8CFF99FF8753DBF82E08C6F86D" box="[199,1244,1994,2021]" pageId="7" pageNumber="294" type="distribution">
<paragraph id="8B568E07FF99FF8753DBF82E08C6F86D" blockId="7.[151,1437,1454,2021]" box="[199,1244,1994,2021]" pageId="7" pageNumber="294">
<emphasis id="B99D5215FF99FF8753DBF82E0D7BF86C" bold="true" box="[199,353,1994,2020]" pageId="7" pageNumber="294">Distribution.</emphasis>
Known from the provinces of Imbabura and
<collectingCountry id="F3FECE97FF99FF875047F82E0FFBF86C" box="[859,993,1994,2020]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="7" pageNumber="294">Santa Elena</collectingCountry>
(
<collectingCountry id="F3FECE97FF99FF8750F3F82E084AF86D" box="[1007,1104,1994,2021]" name="Ecuador" pageId="7" pageNumber="294">Ecuador</collectingCountry>
) (
<figureCitation id="13D29282FF99FF87577BF82E08D5F86D" box="[1127,1231,1994,2021]" captionStart="FIGURE 59" captionStartId="64.[151,250,1837,1861]" captionTargetBox="[162,1435,181,1814]" captionTargetId="figure-24@64.[151,1435,181,1814]" captionTargetPageId="64" captionText="FIGURE 59. Maps of South America showing records of Marma species (A species previously described; B new species). Record from the state of Amazonas (Brazil) in Fig. 59A is taken from Zhang &amp; Maddison (2015)." figureDoi="http://doi.org/10.5281/zenodo.4456983" httpUri="https://zenodo.org/record/4456983/files/figure.png" pageId="7" pageNumber="294">Fig. 59A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>