254 lines
36 KiB
XML
254 lines
36 KiB
XML
<document id="FB4DE3423E903564D0E7C2013B2D97C1" ID-CLB-Dataset="53418" ID-DOI="10.11646/zootaxa.5238.1.1" ID-GBIF-Dataset="751096f2-4b5b-43c3-9748-4d07afe044c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7621793" ID-ZooBank="768E9932-2D18-4115-8359-3FF800328BCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1675871988450" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2023" docId="03C79010FFAAD742FF707FE420CFFADA" docLanguage="en" docName="zootaxa.5238.1.1.pdf" docOrigin="Zootaxa 5238 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5238.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chloeia piotrowskiae Salazar-Vallejo 2023, sp. n." docType="treatment" docVersion="8" lastPageNumber="94" masterDocId="FFFEE868FFF6D71FFFE77B0B2306FFEB" masterDocTitle="Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="93" updateTime="1699474919205" updateUser="ExternalLinkService">
|
||
<mods:mods id="C92B389A028949AB431252BA2B586FCF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4429CF2C54E1700B9A718A5013ECC49C">
|
||
<mods:title id="7A690716E1898157C605818762BF4FEE">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="43257370577D75E841208BFBA460D89F" type="personal">
|
||
<mods:role id="D34E9E7DD5E73D84CC86A14F094A7D47">
|
||
<mods:roleTerm id="E35E3410B6F2836CD31A5A6F914AB003">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4792C73EB0762A43ED70AFA404537F92">Salazar-Vallejo, Sergio I.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="022331B972B656B541F2C17ED433496D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="85283C67E5D21305D1687BB63ED9178C" type="host">
|
||
<mods:titleInfo id="1037DA1DE6B8F0F01A1BCE9B8D12BBEF">
|
||
<mods:title id="D3F9BE87ECA99291DDE190B7160E3335">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="CC3A84A250A7AE2841C2D7A3C3F2F6C1">
|
||
<mods:date id="0F3AE9660850F3F7F00DBC52325DAE43">2023</mods:date>
|
||
<mods:detail id="A733D866860C3894600B0C3D239D1FAC" type="pubDate">
|
||
<mods:number id="E96C29981893696DE8A75E36F5F853CA">2023-02-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A442F892941B1C4ECDB35ED1F64EA52D" type="volume">
|
||
<mods:number id="6BFF6ED547D1FE2809A9EDAEE9EAD610">5238</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C473B4094252007F3012D209ED605949" type="issue">
|
||
<mods:number id="E3977836B059E81F66E3625AD2ADB1A3">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4FB83565B0D3CB0C1F8877F57D0D2A11" unit="page">
|
||
<mods:start id="A9F3D73752061ED1EA716CEF55403097">1</mods:start>
|
||
<mods:end id="8C00F04C8B8A33A37A47AFB5D5214CE5">134</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="7E83069BF12A5BEFAE24AD1755CB02FC">
|
||
<mods:url id="B37FC9118B118B4E44866EC0BFBFF15D">http://dx.doi.org/10.11646/zootaxa.5238.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="5E913214B7B1EB04AABC515678BA218B">journal article</mods:classification>
|
||
<mods:identifier id="3B0AB3B73C5EBD6D3715A2B8EF26D0F5" type="CLB-Dataset">53418</mods:identifier>
|
||
<mods:identifier id="0E1D8BD0B190ED0FFE12CE156F68EF83" type="DOI">10.11646/zootaxa.5238.1.1</mods:identifier>
|
||
<mods:identifier id="B9A64737482FCF2D5D56F8DB8371305F" type="GBIF-Dataset">751096f2-4b5b-43c3-9748-4d07afe044c3</mods:identifier>
|
||
<mods:identifier id="C2554CCD7D76F510492498B2F944D0B9" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="022F022BB33D626D639B97FFD16A2794" type="Zenodo-Dep">7621793</mods:identifier>
|
||
<mods:identifier id="9EB4BD09CABF2289E70613ACC281A8E3" type="ZooBank">768E9932-2D18-4115-8359-3FF800328BCD</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03C79010FFAAD742FF707FE420CFFADA" ID-DOI="http://doi.org/10.5281/zenodo.7641429" ID-GBIF-Taxon="209295616" ID-Zenodo-Dep="7641429" LSID="urn:lsid:plazi:treatment:03C79010FFAAD742FF707FE420CFFADA" httpUri="http://treatment.plazi.org/id/03C79010FFAAD742FF707FE420CFFADA" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
|
||
<subSubSection id="C374728DFFAAD743FF707FE42071FAC3" pageId="92" pageNumber="93" type="nomenclature">
|
||
<paragraph id="8BD12106FFAAD743FF707FE422E6FAE2" blockId="92.[151,887,1263,1356]" box="[151,480,1263,1290]" pageId="92" pageNumber="93">
|
||
<heading id="D099966AFFAAD743FF707FE422E6FAE2" bold="true" box="[151,480,1263,1290]" fontSize="11" level="2" pageId="92" pageNumber="93" reason="4">
|
||
<emphasis id="B91AFD14FFAAD743FF707FE422E6FAE2" bold="true" box="[151,480,1263,1290]" pageId="92" pageNumber="93">
|
||
<taxonomicName id="4C6E5A85FFAAD743FF707FE42290FAE1" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[151,406,1263,1290]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="92" pageNumber="93" phylum="Annelida" rank="species" species="piotrowskiae" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFAAD743FF707FE42290FAE1" bold="true" box="[151,406,1263,1290]" italics="true" pageId="92" pageNumber="93">Chloeia piotrowskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFAAD743FE797FE422E6FAE2" box="[414,480,1263,1289]" pageId="92" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFAAD743FF707E1B2071FAC3" blockId="92.[151,887,1263,1356]" box="[151,887,1296,1320]" pageId="92" pageNumber="93">
|
||
<uri id="FFFF2D04FFAAD743FF707E1B2071FAC3" box="[151,887,1296,1320]" pageId="92" pageNumber="93">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFC81BD3FFAAD743FE687E1B2071FAC3" box="[399,887,1296,1320]" pageId="92" pageNumber="93">23EC1D3E-EF9A-459F-B3E7-05204F8DFF86</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFAAD743FF707E3923E1FAA7" box="[151,231,1330,1356]" pageId="92" pageNumber="93" type="description">
|
||
<paragraph id="8BD12106FFAAD743FF707E3923E1FAA7" blockId="92.[151,887,1263,1356]" box="[151,231,1330,1356]" pageId="92" pageNumber="93">
|
||
<figureCitation id="13553D83FFAAD743FF707E3923E1FAA7" box="[151,231,1330,1356]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="92" pageNumber="93">Fig. 43</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFAAD743FF707E7E2768FA10" pageId="92" pageNumber="93" type="materials_examined">
|
||
<paragraph id="8BD12106FFAAD743FF707E7E2768FA10" blockId="92.[151,1437,1396,2035]" pageId="92" pageNumber="93">
|
||
<emphasis id="B91AFD14FFAAD743FF707E7E223BFA65" bold="true" box="[151,317,1396,1423]" pageId="92" pageNumber="93">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B062B5BFFAAD743FEAD7E7F21F0FA64" ID-GBIF-Occurrence="4029882362" box="[330,758,1396,1423]" collectingDate="2011-06-02" collectionCode="CAS" collectorName="R. Mooi" country="Philippines" county="Ambil Island" elevation="84" latitude="13.91" location="Philippines" longLatPrecision="1" longitude="120.35" municipality="Lubang Islands" pageId="92" pageNumber="93" specimenCode="186957" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B91AFD14FFAAD743FEAD7E7F22D6FA65" bold="true" box="[330,464,1396,1422]" pageId="92" pageNumber="93">
|
||
<collectingCountry id="F3796196FFAAD743FEAD7E7F22D6FA65" box="[330,464,1396,1422]" name="Philippines" pageId="92" pageNumber="93">Philippines</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54D59FA4FFAAD743FE397E7E2140FA64" box="[478,582,1397,1423]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFAAD743FDB37E7E218AFA64" box="[596,652,1397,1423]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="92" pageNumber="93">CAS</collectionCode>
|
||
<specimenCode id="DBC8897DFFAAD743FD757E7F21ECFA64" box="[658,746,1396,1423]" pageId="92" pageNumber="93">186957</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="3B062B5BFFAAD743FD1A7E7E2768FA10" ID-GBIF-Occurrence="4029882509" collectingDate="2011-06-02" collectionCode="CAS" collectorName="R. Mooi" country="Philippines" county="Ambil Island" elevation="84" latitude="13.91" location="Hearst Philippine Biodiversity Expedition" longLatPrecision="1" longitude="120.35" municipality="Lubang Islands" pageId="92" pageNumber="93" specimenCode="186957" specimenCount="1" typeStatus="paratype">
|
||
and
|
||
<specimenCount id="9D68EA8FFFAAD743FCCC7E7E2053FA64" box="[811,853,1397,1423]" pageId="92" pageNumber="93">two</specimenCount>
|
||
<typeStatus id="54D59FA4FFAAD743FCBB7E7E20C1FA64" box="[860,967,1397,1423]" pageId="92" pageNumber="93" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFAAD743FC327E7E270BFA64" box="[981,1037,1397,1423]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="92" pageNumber="93">CAS</collectionCode>
|
||
<specimenCode id="DBC8897DFFAAD743FBF47E7F276DFA64" box="[1043,1131,1396,1423]" pageId="92" pageNumber="93">186957</specimenCode>
|
||
b),
|
||
<location id="8EB177DDFFAAD743FB6B7E7E2263FA58" LSID="urn:lsid:plazi:treatment:03C79010FFAAD742FF707FE420CFFADA:8EB177DDFFAAD743FB6B7E7E2263FA58" country="Philippines" county="Ambil Island" latitude="13.91" longLatPrecision="1" longitude="120.35" municipality="Lubang Islands" name="Hearst Philippine Biodiversity Expedition" pageId="92" pageNumber="93">Hearst Philippine Biodiversity Expedition</location>
|
||
2011, MV DA-BFAR,
|
||
<collectingMunicipality id="6BB5BB7CFFAAD743FD8B7E92201DFA58" box="[620,795,1433,1459]" pageId="92" pageNumber="93">Lubang Islands</collectingMunicipality>
|
||
, off
|
||
<collectingCounty id="62B0598AFFAAD743FCAB7E9220D8FA58" box="[844,990,1433,1459]" pageId="92" pageNumber="93">Ambil Island</collectingCounty>
|
||
(13.91° N, 120.35° E;
|
||
<geoCoordinate id="EE5A47C1FFAAD743FB057E93269EFA58" box="[1250,1432,1432,1459]" degrees="13" direction="north" minutes="54" orientation="latitude" pageId="92" pageNumber="93" precision="1" seconds="36.00" value="13.91">13°54’36.00” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFAAD743FF707EB6225EFA3C" box="[151,344,1469,1495]" degrees="120" direction="east" minutes="21" orientation="longitude" pageId="92" pageNumber="93" precision="1" seconds="00.00" value="120.35">120°21’00.00” E</geoCoordinate>
|
||
), mixed bottom, sand and rocks,
|
||
<quantity id="4C968CE3FFAAD743FD2A7EB62035FA3C" box="[717,819,1469,1495]" metricMagnitude="1" metricUnit="m" metricValue="8.4" metricValueMax="8.6" metricValueMin="8.2" pageId="92" pageNumber="93" unit="m" value="84.0" valueMax="86.0" valueMin="82.0">
|
||
<elevation id="0043C635FFAAD743FD2A7EB62035FA3C" box="[717,819,1469,1495]" metricMagnitude="1" metricUnit="m" metricValue="8.4" metricValueMax="8.6" metricValueMin="8.2" pageId="92" pageNumber="93" unit="m" value="84.0" valueMax="86.0" valueMin="82.0">82–86 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFD007C6FFAAD743FCD87EB620C5FA3C" box="[831,963,1469,1495]" pageId="92" pageNumber="93" value="2011-06-02">
|
||
<collectingDate id="EF94FE2EFFAAD743FCD87EB620C5FA3C" box="[831,963,1469,1495]" pageId="92" pageNumber="93" value="2011-06-02">2 Jun. 2011</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269B44D0FFAAD743FC287EB6272BFA3C" box="[975,1069,1469,1495]" pageId="92" pageNumber="93">R. Mooi</collectorName>
|
||
<emphasis id="B91AFD14FFAAD743FBD37EB6276AFA3C" box="[1076,1132,1469,1495]" italics="true" pageId="92" pageNumber="93">et al.</emphasis>
|
||
, coll. (paratypes complete, stiff, many chaetae broken, body 27.5–45.0 mm long,
|
||
<quantity id="4C968CE3FFAAD743FD117EEA2055FA10" box="[758,851,1505,1531]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="92" pageNumber="93" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
wide, 29–34 chaetigers).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFAAD743FF207D0F267BF98C" pageId="92" pageNumber="93" type="diagnosis">
|
||
<paragraph id="8BD12106FFAAD743FF207D0F267BF98C" blockId="92.[151,1437,1396,2035]" pageId="92" pageNumber="93">
|
||
<emphasis id="B91AFD14FFAAD743FF207D0F223EF9F5" bold="true" box="[199,312,1540,1566]" pageId="92" pageNumber="93">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFAAD743FEAE7D0E22A6F9F4" authorityName="Savigny" authorityYear="1818" box="[329,416,1541,1567]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="92" pageNumber="93" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91AFD14FFAAD743FEAE7D0E22A6F9F4" box="[329,416,1541,1567]" italics="true" pageId="92" pageNumber="93">Chloeia</emphasis>
|
||
</taxonomicName>
|
||
with bipinnate branchiae from chaetiger 4, decreasing in size posteriorly; middorsal band single, wide, continuous, running throughout body; caruncle pale; anterior notopodia with furcate harpoon chaetae and aciculars, median chaetigers with aciculars and harpoon chaetae without spurs; neurochaetae spurred and furcates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFAAD742FF207D7B220CFD1A" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="description">
|
||
<paragraph id="8BD12106FFAAD743FF207D7B2235F938" blockId="92.[151,1437,1396,2035]" pageId="92" pageNumber="93">
|
||
<emphasis id="B91AFD14FFAAD743FF207D7B2257F961" bold="true" box="[199,337,1648,1674]" pageId="92" pageNumber="93">Description</emphasis>
|
||
.
|
||
<typeStatus id="54D59FA4FFAAD743FEB87D7A22C1F960" box="[351,455,1649,1675]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
|
||
(CAS 186957) complete, stiff (fixed in 95% ethanol), slightly bent laterally, many chaetae broken, portions of venter along chaetigers 14–17 previously removed; body fusiform,
|
||
<quantity id="4C968CE3FFAAD743FB617D9F27D2F944" box="[1158,1236,1684,1711]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="92" pageNumber="93" unit="mm" value="35.0">35 mm</quantity>
|
||
long,
|
||
<quantity id="4C968CE3FFAAD743FAFE7D9E265FF944" box="[1305,1369,1685,1711]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="92" pageNumber="93" unit="mm" value="6.0">6 mm</quantity>
|
||
wide, 31 chaetigers.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFAAD743FF207DD6265AF888" blockId="92.[151,1437,1396,2035]" pageId="92" pageNumber="93">
|
||
<typeStatus id="54D59FA4FFAAD743FF207DD62229F91C" box="[199,303,1757,1783]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
|
||
colorful, chaetae colorless, almost transparent; middorsal band reddish present along body from chaetiger 4 (
|
||
<figureCitation id="13553D83FFAAD743FF157C0A225DF8F0" box="[242,347,1793,1819]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="92" pageNumber="93">Fig. 43A</figureCitation>
|
||
), continued to last chaetiger; middorsal band homogeneously wide throughout body (
|
||
<figureCitation id="13553D83FFAAD743FAC07C0A2689F8F0" box="[1319,1423,1793,1819]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="92" pageNumber="93">Fig. 43C</figureCitation>
|
||
); chaetigers 1–3 with similar bands bordering caruncle. Branchiae with orange stems and paler to white lateral branches. Dorsal cirri and ventral cirri of chaetiger 2 dark purple. Anterior parapodial surfaces with yellowish band.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFAAD743FF207C662115F818" blockId="92.[151,1437,1396,2035]" pageId="92" pageNumber="93">
|
||
Prostomium anteriorly entire, anterior prostomial area whitish (
|
||
<figureCitation id="13553D83FFAAD743FC627C6620ECF86C" box="[901,1002,1901,1927]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="92" pageNumber="93">Fig. 43B</figureCitation>
|
||
). Eyes blackish, anterior eyes 3× larger than posterior ones. Median antenna inserted at anterior caruncular margin, 1/3 as long as caruncle, 2× longer than lateral antennae. Lateral antennae bases separate, reddish medially, slightly longer than palps. Mouth vental on chaetiger 2. Pharynx not exposed.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFABD742FF207B9C23D1FF12" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">Caruncle pale, tapered, bent laterally, reaching chaetiger 4. Median ridge plicate, with about 30 vertical folds, most marked by a blackish spot, partially concealing lateral lobes. Lateral lobes narrow, with about 32 vertical folds.</paragraph>
|
||
<paragraph id="8BD12106FFABD742FF207A08269DFEAA" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">Bipinnate branchiae from chaetiger 4, roughly parallel along body, progressively larger to chaetigers 10–14, decreasing in size posteriorly. Median segments with 14–16 lateral branches, each with a few secondary filaments.</paragraph>
|
||
<paragraph id="8BD12106FFABD742FF207A402074FE3A" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">Parapodia biramous, notopodia with cirriform branchiae along chaetigers 1–3, shorter progressively. Dorsal cirri longer than bipinnate branchiae along median chaetigers, 2–3× longer in posterior chaetigers. Second ventral cirri with cirrophores 3–4× wider, and cirrostyles 3–4× longer than adjacent ones, directed dorsally. Other ventral cirri directed ventrolaterallly, as long as one subsequent segment.</paragraph>
|
||
<paragraph id="8BD12106FFABD742FF207AD0279CFD42" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
Chaetae generally broken in
|
||
<typeStatus id="54D59FA4FFABD742FDF07AD0217AFE1E" box="[535,636,475,501]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
|
||
, observed in largest
|
||
<typeStatus id="54D59FA4FFABD742FC897AD020C8FE1E" box="[878,974,475,501]" pageId="93" pageNumber="94" type="paratype">paratype</typeStatus>
|
||
for avoiding further damage. Complete chaetae without distal hoods, probably eroded. Anterior notochaetae a few aciculars and many harpoon-chaetae, a few without spurs, others with spurs 1/3–1/5 as long as denticulate (
|
||
<figureCitation id="13553D83FFABD742FC70792820FAFDD6" box="[919,1020,547,573]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="93" pageNumber="94">Fig. 43F</figureCitation>
|
||
). Median chaetigers with a few short aciculars and longest harpoon-shaped chaetae without spurs (
|
||
<figureCitation id="13553D83FFABD742FCA0794C20B6FD8A" box="[839,944,583,609]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="93" pageNumber="94">Fig. 43H</figureCitation>
|
||
). Neurochaetae mostly furcates and spurred chaetae, furcates with major tines thinner in anterior chaetigers (
|
||
<figureCitation id="13553D83FFABD742FC8E796020D7FD6E" box="[873,977,619,645]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="93" pageNumber="94">Fig. 43G</figureCitation>
|
||
), major tines about 4× longer than minor one; posterior chaetigers (
|
||
<figureCitation id="13553D83FFABD742FE5D79842112FD42" box="[442,532,655,681]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="93" pageNumber="94">Fig. 43I</figureCitation>
|
||
) with furcates, major tines 3—6×longer than minor ones.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFABD742FF2079B8220CFD1A" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
Posterior region tapered (
|
||
<figureCitation id="13553D83FFABD742FE0B79B8215EFD26" box="[492,600,691,717]" captionStart="FIGURE 43" captionStartId="92.[152,255,1024,1048]" captionTargetBox="[154,1435,184,1000]" captionTargetId="figure-28@92.[151,1436,181,1000]" captionTargetPageId="92" captionText="FIGURE 43. Chloeia piotrowski sp. n. A. Holotype (CAS 186957), anterior region, dorsal view. B. Same, anterior end, oblique left lateral view, after Methyl Green staining. C. Same, chaetigers 9-11, dorsal view. D. Same, posterior region, dorsal view. E. Largest paratype (CAS 186957b), anterior end, dorsal view, after Methyl green staining. F. Chaetiger 4, notochaetal tips. G. Same, neurochaetal tips. H. Chaetiger 10, notochaetal tips. I. Same, neurochaetal tips. Scale bars:A, 2 mm; B, 1 mm; C, 0.7 mm; D, 1.7 mm; E, 1.4 mm; F, 200 μm; G, I, 40 μm; H, 160 μm." figureDoi="http://doi.org/10.5281/zenodo.7621949" httpUri="https://zenodo.org/record/7621949/files/figure.png" pageId="93" pageNumber="94">Fig. 43D</figureCitation>
|
||
); pygidium with anus terminal; anal cirri unknown, lost in
|
||
<typeStatus id="54D59FA4FFABD742FAEF79B8266CFD26" box="[1288,1386,691,717]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54D59FA4FFABD742FF7079DC2200FD1A" box="[151,262,727,753]" pageId="93" pageNumber="94" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFABD742FF2079F127E0FCD2" pageId="93" pageNumber="94" type="description">
|
||
<paragraph id="8BD12106FFABD742FF2079F127E0FCD2" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
<emphasis id="B91AFD14FFABD742FF2079F122A4FCFF" bold="true" box="[199,418,762,788]" pageId="93" pageNumber="94">Live pigmentation</emphasis>
|
||
(after C. Piotrowski notes after collecting). Middorsal longitudinal band and branchiae red. Median caruncular rigde with black spots, Dorsal cirri, and ventral cirri of chaetiger 2 dark purple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFABD742FF207848217FFC22" pageId="93" pageNumber="94" type="etymology">
|
||
<paragraph id="8BD12106FFABD742FF207848217FFC22" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
<emphasis id="B91AFD14FFABD742FF207848224EFCB6" bold="true" box="[199,328,835,861]" pageId="93" pageNumber="94">Etymology</emphasis>
|
||
. This species is named after M. Sc. Christina Piotrowski, Collection Manager,
|
||
<collectingRegion id="49AAEFE4FFABD742FB5D7848262AFCB6" box="[1210,1324,835,861]" country="United States of America" name="California" pageId="93" pageNumber="94">California</collectingRegion>
|
||
Academy of Sciences, San Francisco, in recognition of her long-term efforts in collecting polychaetes from many tropical localities all over the world, and especially by her support to my research activities. The derived name is a noun in the genitive case (
|
||
<bibRefCitation id="EFFF5CF7FFABD742FE8778A422E3FC22" author="ICZN & International Commission of Zoological Nomenclature" box="[352,485,943,969]" pageId="93" pageNumber="94" refId="ref108505" refString="ICZN (International Commission of Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London. Available from: https: // code. iczn. org (accessed 22 June 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
, Art. 31.1.2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFABD742FF2078D92073FBDE" pageId="93" pageNumber="94" type="description">
|
||
<paragraph id="8BD12106FFABD742FF2078D92073FBDE" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
<emphasis id="B91AFD14FFABD742FF2078D9223EFC07" bold="true" box="[199,312,978,1004]" pageId="93" pageNumber="94">Variation</emphasis>
|
||
. Most morphological features are present in
|
||
<typeStatus id="54D59FA4FFABD742FCD678D820A6FC06" box="[817,928,979,1005]" pageId="93" pageNumber="94" type="paratype">paratypes</typeStatus>
|
||
. The middorsal reddish band has faded off in last few chaetigers. Anterior eyes are 4× as long as posterior ones in the smallest
|
||
<typeStatus id="54D59FA4FFABD742FBFA78FC2786FBFA" box="[1053,1152,1015,1041]" pageId="93" pageNumber="94" type="paratype">paratype</typeStatus>
|
||
, 3× as long in the longest one (
|
||
<figureCitation id="13553D83FFABD742FF287F102233FBDE" box="[207,309,1051,1077]" captionStart="FIGURE 41" captionStartId="88.[152,255,1522,1546]" captionTargetBox="[155,1433,348,1498]" captionTargetId="figure-114@88.[151,1436,345,1499]" captionTargetPageId="88" captionText="FIGURE 41. Chloeia murrayae sp. n., holotype (AM W28128). A. Anterior region, dorsal view. B. Anterior end, oblique dorsal view. C. Chaetiger 3, notochaetal tips. D. Same, neurochaetal tips. E. Chaetiger 15, notochaetal tips. F. Same, neurochaetal tips. G. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, 0.8 mm; C, E, 110 μm; D, F, 30 μm; G, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621943" httpUri="https://zenodo.org/record/7621943/files/figure.png" pageId="93" pageNumber="94">Fig. 41E</figureCitation>
|
||
). The median antenna is ¼–1/3 as long as caruncle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFABD742FF207F34265EFAE6" pageId="93" pageNumber="94" type="discussion">
|
||
<paragraph id="8BD12106FFABD742FF207F34265EFAE6" blockId="93.[151,1437,151,1329]" pageId="93" pageNumber="94">
|
||
<emphasis id="B91AFD14FFABD742FF207F342232FBB2" bold="true" box="[199,308,1087,1113]" pageId="93" pageNumber="94">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFABD742FEA27F34213EFBB2" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[325,568,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="93" pageNumber="94" phylum="Annelida" rank="species" species="piotrowskiae" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFABD742FEA27F34213EFBB2" box="[325,568,1087,1113]" italics="true" pageId="93" pageNumber="94">Chloeia piotrowskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFABD742FDA57F342187FBB2" box="[578,641,1087,1113]" pageId="93" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
|
||
is described with specimens from The
|
||
<collectingCountry id="F3796196FFABD742FBA87F3427D7FBB2" box="[1103,1233,1087,1113]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
|
||
; it belongs in the group venusta because it has a middorsal band, bipinnate branchiae from chaetiger 4, decreasing in size posteriorly. Because the middorsal band is regular, continuous throughout complete segments, it resembles
|
||
<taxonomicName id="4C6E5A85FFABD742FB3A7F8C2662FB4A" authorityName="McIntosh" authorityYear="1925" box="[1245,1380,1159,1185]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="93" pageNumber="94" phylum="Annelida" rank="species" species="gilchristi">
|
||
<emphasis id="B91AFD14FFABD742FB3A7F8C2662FB4A" box="[1245,1380,1159,1185]" italics="true" pageId="93" pageNumber="94">C. gilchristi</emphasis>
|
||
</taxonomicName>
|
||
Mc-Intosh, 1924 from
|
||
<collectingCountry id="F3796196FFABD742FE8C7FA022FBFB2E" box="[363,509,1195,1221]" name="South Africa" pageId="93" pageNumber="94">South Africa</collectingCountry>
|
||
, redescribed above,
|
||
<taxonomicName id="4C6E5A85FFABD742FD017FA02058FB2F" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[742,862,1195,1221]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="93" pageNumber="94" phylum="Annelida" rank="species" species="poupini" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFABD742FD017FA02058FB2F" box="[742,862,1195,1221]" italics="true" pageId="93" pageNumber="94">C. poupini</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFABD742FC827FA020A5FB2E" box="[869,931,1195,1221]" pageId="93" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
|
||
from the
|
||
<collectingCountry id="F3796196FFABD742FBF37FA027D1FB2E" box="[1044,1239,1195,1221]" name="French Polynesia" pageId="93" pageNumber="94">French Polynesia</collectingCountry>
|
||
described below, and
|
||
<taxonomicName id="4C6E5A85FFABD742FF2E7FC42138FB02" authority="de Quatrefages, 1866" authorityName="de Quatrefages" authorityYear="1866" box="[201,574,1231,1257]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="93" pageNumber="94" phylum="Annelida" rank="species" species="venusta">
|
||
<emphasis id="B91AFD14FFABD742FF2E7FC42244FB03" box="[201,322,1231,1257]" italics="true" pageId="93" pageNumber="94">C. venusta</emphasis>
|
||
<bibRefCitation id="EFFF5CF7FFABD742FEAD7FC42138FB02" author="De Quatrefages, A." box="[330,574,1231,1257]" pageId="93" pageNumber="94" pagination="578 - 600" refId="ref105083" refString="De Quatrefages, A. (1866 (1865 )) Histoire naturelle des Anneles marins et d'eau douce. Annelides et Gephyriens. Vol. 1. Librarie Encyclopedique de Roret, Paris, 588 pp. [publication date after Wright, E. P. (1866). Section 6 (Annelida, etc.). Zoological Record, 1866, 578 - 600] https: // doi. org / 10.5962 / bhl. title. 122818" type="book chapter" year="1866">de Quatrefages, 1866</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Mediterranean Sea, redescribed below. However,
|
||
<taxonomicName id="4C6E5A85FFABD742FB0D7FC4269BFB02" authorityName="Salazar-Vallejo" authorityYear="2023" box="[1258,1437,1231,1257]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="93" pageNumber="94" phylum="Annelida" rank="species" species="piotrowskiae">
|
||
<emphasis id="B91AFD14FFABD742FB0D7FC4269BFB02" box="[1258,1437,1231,1257]" italics="true" pageId="93" pageNumber="94">C. piotrowskiae</emphasis>
|
||
</taxonomicName>
|
||
separates from these species because it is the only species having harpoon notochaetae in anterior parapodia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFABD742FF207E1D20CFFADA" box="[199,969,1302,1329]" pageId="93" pageNumber="94" type="distribution">
|
||
<paragraph id="8BD12106FFABD742FF207E1D20CFFADA" blockId="93.[151,1437,151,1329]" box="[199,969,1302,1329]" pageId="93" pageNumber="94">
|
||
<emphasis id="B91AFD14FFABD742FF207E1D225FFADB" bold="true" box="[199,345,1302,1328]" pageId="93" pageNumber="94">Distribution</emphasis>
|
||
. The
|
||
<collectingCountry id="F3796196FFABD742FE7E7E1C211CFADA" box="[409,538,1303,1329]" name="Philippines" pageId="93" pageNumber="94">Philippines</collectingCountry>
|
||
, in sediments at
|
||
<quantity id="4C968CE3FFABD742FD337E1C2031FADA" box="[724,823,1303,1329]" metricMagnitude="1" metricUnit="m" metricValue="8.4" metricValueMax="8.6" metricValueMin="8.2" pageId="93" pageNumber="94" unit="m" value="84.0" valueMax="86.0" valueMin="82.0">82–86 m</quantity>
|
||
water depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |