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<mods:title>Phytoseiid mites (Acari: Phytoseiidae) from Egypt, with new records, descriptions of new species, and a key to species</mods:title>
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<mods:namePart>Abo-Shnaf, Reham I. A.</mods:namePart>
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<taxonomicName authority="Wainstein &amp; Arutunjan" authorityName="Wainstein &amp; Arutunjan" box="[151,672,1741,1767]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="segnis">
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<emphasis bold="true" box="[151,367,1741,1767]" italics="true" pageId="27" pageNumber="28">Neoseiulus segnis</emphasis>
Wainstein &amp; Arutunjan
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<emphasis box="[151,335,1812,1834]" italics="true" pageId="27" pageNumber="28">Amblyseius segnis</emphasis>
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Wainstein &amp; Arutunjan, 
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: 
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; 
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<bibRefCitation author="Wainstein" box="[718,899,1812,1834]" pageId="27" pageNumber="28" refString="Wainstein, B. A., (1977 b) Family Phytoseiidae Ber 1 ese, 1916. In: Ghi 1 arov, B. M. (Ed.), Key to Soil - Inhabiting Mites (Mesostigmata). Ed. &quot; Nauka &quot;, Leningrad, pp. 227 - 244." type="journal article" year="1977" yearSuffix="b">
Wainstein, 
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b
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<treatmentCitation author="Beglyarov" box="[959,1168,1812,1834]" page="40" pageId="27" pageNumber="28" year="1981">
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Beglyarov, 
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: 
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. 
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<emphasis box="[151,266,1842,1864]" italics="true" pageId="27" pageNumber="28">Amblyseius</emphasis>
(
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) 
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. 
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Arutunjan, 
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: 
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<emphasis box="[151,331,1873,1895]" italics="true" pageId="27" pageNumber="28">Neoseiulus segnis</emphasis>
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. 
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Moraes 
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., 
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: 
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; 
<treatmentCitation author="Chant" box="[724,1025,1873,1895]" page="35" pageId="27" pageNumber="28" year="2003">
<bibRefCitation author="Chant" box="[724,987,1873,1895]" pageId="27" pageNumber="28" refString="Chant, D. A. &amp; McMurtry, J. A. (2003) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology, 29, 3 - 46. http: // dx. doi. org / 10.1080 / 01647950308684319" type="journal article" year="2003">
Chant &amp; McMurtry, 
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: 
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. 
<taxonomicName authority="Basha, Yousef, Ibrahim &amp; Mostafa" authorityName="Basha, Yousef, Ibrahim &amp; Mostafa" box="[151,737,1904,1926]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis box="[151,375,1904,1926]" italics="true" pageId="27" pageNumber="28">Neoseiulus seminudus</emphasis>
Basha, Yousef, Ibrahim &amp; Mostafa
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, in 
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<bibRefCitation author="Basha" box="[773,959,1904,1926]" pageId="27" pageNumber="28" refString="Basha, A. E., Yousef, A. A., Ibrahim, M. H. &amp; Mostafa, E. M. (2001) Five new phytoseiids from Egypt (Acari: Gamasida: Phytoseiidae). Al - Azhar Journal of Agricultural Research, 33, 371 - 386." type="journal article" year="2001">
Basha 
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., 
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: 
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(
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<emphasis bold="true" box="[151,238,1970,1995]" pageId="27" pageNumber="28">Female</emphasis>
(one specimen).
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Dorsal shield smooth; 
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long and 
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wide, with 
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pairs of setae. Setae 
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j 
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, 
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j 
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j 
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j 
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Z 
<number box="[514,525,152,176]" pageId="28" pageNumber="29" value="1.0">1</number>
</emphasis>
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, 
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Z 
<number box="[592,605,152,176]" pageId="28" pageNumber="29" value="4.0">4</number>
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S 
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r 
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, 
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R 
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<number box="[1158,1189,152,176]" pageId="28" pageNumber="29" value="20.0">20</number>
. Dorsal setae smooth, except 
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Z 
<number box="[247,260,189,212]" pageId="28" pageNumber="29" value="5.0">5</number>
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, slightly serrate. Peritreme extending forward to level of 
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j 
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Venter. Sternal shield smooth with few lateral striae, with three pairs of setae and two pairs of lyrifissures. Distances between 
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st 
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–st 
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, 
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st 
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–st 
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st 
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–st 
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st 
<number box="[779,793,260,284]" pageId="28" pageNumber="29" value="4.0">4</number>
–st 
<number box="[826,840,260,284]" pageId="28" pageNumber="29" value="4.0">4</number>
</emphasis>
<number box="[848,879,260,285]" pageId="28" pageNumber="29" value="61.0">61</number>
. Genital shield smooth; distance between 
<emphasis box="[1356,1436,261,284]" italics="true" pageId="28" pageNumber="29">
st 
<number box="[1375,1389,261,284]" pageId="28" pageNumber="29" value="5.0">5</number>
–st 
<number box="[1422,1436,261,284]" pageId="28" pageNumber="29" value="5.0">5</number>
</emphasis>
<number box="[151,182,295,320]" pageId="28" pageNumber="29" value="55.0">55</number>
. Ventrianal shield subpentagonal, with sparse transverse striae anteriorly to pre-anal pores and reticulate posteriorly; 
<number box="[286,328,332,357]" pageId="28" pageNumber="29" value="105.0">105</number>
long, 
<number box="[398,426,332,356]" pageId="28" pageNumber="29" value="91.0">91</number>
wide at 
<emphasis box="[519,566,332,356]" italics="true" pageId="28" pageNumber="29">
ZV 
<number box="[554,566,332,356]" pageId="28" pageNumber="29" value="2.0">2</number>
</emphasis>
level and 
<number box="[680,708,332,356]" pageId="28" pageNumber="29" value="82.0">82</number>
wide at level of anus; with three pairs of pre-anal setae and a pair of pre-anal pores. Seta 
<emphasis box="[410,454,368,391]" italics="true" pageId="28" pageNumber="29">
JV 
<number box="[441,454,368,391]" pageId="28" pageNumber="29" value="5.0">5</number>
</emphasis>
<number box="[460,491,368,392]" pageId="28" pageNumber="29" value="39.0">39</number>
. Ventral setae smooth. Two pairs of metapodal plates.
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" box="[199,983,404,429]" pageId="28" pageNumber="29">
Spermatheca. Calyx of spermatheca saccular, 
<number box="[713,741,404,428]" pageId="28" pageNumber="29" value="21.0">21</number>
long; atrium distinct.
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" pageId="28" pageNumber="29">
Gnathosoma. Corniculi parallel to each other; basal width of corniculus 
<number box="[1010,1026,439,464]" pageId="28" pageNumber="29" value="5.0">5</number>
, distance between bases of corniculi 
<number box="[151,167,476,500]" pageId="28" pageNumber="29" value="8.0">8</number>
. Movable cheliceral digit 
<number box="[460,488,476,500]" pageId="28" pageNumber="29" value="31.0">31</number>
long, with one tooth; fixed digit 
<number box="[858,886,476,500]" pageId="28" pageNumber="29" value="30.0">30</number>
long, with four teeth.
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" box="[199,1297,511,536]" pageId="28" pageNumber="29">
Legs. Macroseta sharp-tipped: 
<emphasis box="[546,598,512,536]" italics="true" pageId="28" pageNumber="29">
St 
<date box="[573,598,513,536]" pageId="28" pageNumber="29" value="1954-04">IV</date>
</emphasis>
<number box="[607,638,511,536]" pageId="28" pageNumber="29" value="54.0">54</number>
; chaetotaxy of genu 
<date box="[870,911,512,536]" pageId="28" pageNumber="29">
II 
<number box="[895,911,512,536]" pageId="28" pageNumber="29" value="2.2">2</number>
</date>
, 2/ 
<number box="[947,987,512,536]" pageId="28" pageNumber="29" value="0.2">0, 2</number>
/ 
<number box="[997,1038,512,536]" pageId="28" pageNumber="29" value="0.1">0, 1</number>
; genu 
<date box="[1113,1162,512,536]" pageId="28" pageNumber="29">
III 
<number box="[1148,1162,512,536]" pageId="28" pageNumber="29" value="1.2">1</number>
</date>
, 2/ 
<number box="[1203,1240,512,536]" pageId="28" pageNumber="29" value="1.2">1, 2</number>
/ 
<number box="[1250,1291,512,536]" pageId="28" pageNumber="29" value="0.1">0, 1</number>
.
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[199,456,548,573]" pageId="28" pageNumber="29">Specimens examined.</emphasis>
One female from cucumber leaves, at Senuris, Fayoum governorate, 
<date box="[1252,1436,548,573]" pageId="28" pageNumber="29" value="2006-09">
September 
<number box="[1380,1436,548,573]" pageId="28" pageNumber="29" value="2006.0">2006</number>
</date>
(coll. R.I.A. Abo-Shnaf).
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" box="[199,1023,620,645]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[199,550,620,645]" pageId="28" pageNumber="29">
Previous records from 
<collectingCountry box="[472,546,620,645]" name="Egypt" pageId="28" pageNumber="29">Egypt</collectingCountry>
.
</emphasis>
Gharbia governorate (
<bibRefCitation author="Basha" box="[803,1011,620,645]" pageId="28" pageNumber="29" refString="Basha, A. E., Yousef, A. A., Ibrahim, M. H. &amp; Mostafa, E. M. (2001) Five new phytoseiids from Egypt (Acari: Gamasida: Phytoseiidae). Al - Azhar Journal of Agricultural Research, 33, 371 - 386." type="journal article" year="2001">
Basha 
<emphasis box="[880,928,620,645]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
., 
<number box="[950,1011,620,644]" pageId="28" pageNumber="29" value="2001.0">2001</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="discussion">
<paragraph blockId="28.[151,1437,151,1077]" pageId="28" pageNumber="29">
<emphasis bold="true" box="[199,315,655,680]" pageId="28" pageNumber="29">Remarks.</emphasis>
<taxonomicName box="[334,537,655,680]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="segnis">
<emphasis box="[334,537,655,680]" italics="true" pageId="28" pageNumber="29">Neoseiulus segnis</emphasis>
</taxonomicName>
was originally described from the 
<typeStatus box="[942,1040,655,680]" pageId="28" pageNumber="29">holotype</typeStatus>
female and four 
<typeStatus box="[1243,1339,656,680]" pageId="28" pageNumber="29">paratype</typeStatus>
females collected in 
<collectingCountry box="[287,389,692,717]" name="Armenia" pageId="28" pageNumber="29">Armenia</collectingCountry>
. The original description was brief, but with illustrations and setal measurements. 
<taxonomicName class="Arachnida" family="Phytoseiidae" genus="Amblyseius" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis italics="true" pageId="28" pageNumber="29">Amblyseius seminudus</emphasis>
</taxonomicName>
was originally described from the 
<typeStatus box="[658,756,727,752]" pageId="28" pageNumber="29">holotype</typeStatus>
female and five 
<typeStatus box="[944,1041,728,752]" pageId="28" pageNumber="29">paratype</typeStatus>
females collected in Zifta, Gharbia governorate, 
<collectingCountry box="[299,370,765,789]" name="Egypt" pageId="28" pageNumber="29">Egypt</collectingCountry>
. The original description was detailed, with illustrations and setal measurements.
</paragraph>
<paragraph blockId="28.[151,1437,151,1077]" pageId="28" pageNumber="29">
Measurements of the specimen collected are similar to those of the original description of 
<taxonomicName box="[1201,1349,799,824]" class="Arachnida" family="Phytoseiidae" genus="Amblyseius" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis box="[1201,1218,800,823]" italics="true" pageId="28" pageNumber="29">A</emphasis>
. 
<emphasis box="[1231,1349,799,824]" italics="true" pageId="28" pageNumber="29">seminudus</emphasis>
</taxonomicName>
, except for 
<emphasis box="[190,218,836,860]" italics="true" pageId="28" pageNumber="29">
S 
<number box="[204,218,836,860]" pageId="28" pageNumber="29" value="4.0">4</number>
</emphasis>
; although it is described as 
<number box="[530,558,836,860]" pageId="28" pageNumber="29" value="13.0">13</number>
long in the text of the original description, the illustration shows it to be about as long as 
<emphasis box="[289,317,872,896]" italics="true" pageId="28" pageNumber="29">
S 
<number box="[304,317,872,896]" pageId="28" pageNumber="29" value="2.0">2</number>
</emphasis>
(
<number box="[339,368,872,896]" pageId="28" pageNumber="29" value="22.0">22</number>
according to the original description). Additionally, the specimens collected fit well the description of 
<taxonomicName box="[313,415,909,932]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="segnis">
<emphasis box="[313,332,909,932]" italics="true" pageId="28" pageNumber="29">N</emphasis>
. 
<emphasis box="[345,415,909,932]" italics="true" pageId="28" pageNumber="29">segnis</emphasis>
</taxonomicName>
. The measurements of the specimen collected and of those of the original description of 
<taxonomicName class="Arachnida" family="Phytoseiidae" genus="Amblyseius" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis box="[1412,1429,908,931]" italics="true" pageId="28" pageNumber="29">A</emphasis>
. 
<emphasis box="[151,269,943,968]" italics="true" pageId="28" pageNumber="29">seminudus</emphasis>
</taxonomicName>
are similar to those of the original description of 
<taxonomicName box="[830,933,944,968]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="segnis">
<emphasis box="[830,849,945,968]" italics="true" pageId="28" pageNumber="29">N</emphasis>
. 
<emphasis box="[863,933,945,968]" italics="true" pageId="28" pageNumber="29">segnis</emphasis>
</taxonomicName>
, except for 
<emphasis box="[1068,1098,944,968]" italics="true" pageId="28" pageNumber="29">
Z 
<number box="[1085,1098,944,968]" pageId="28" pageNumber="29" value="4.0">4</number>
</emphasis>
, 
<emphasis box="[1113,1142,944,967]" italics="true" pageId="28" pageNumber="29">
Z 
<number box="[1129,1142,944,967]" pageId="28" pageNumber="29" value="5.0">5</number>
</emphasis>
, 
<emphasis box="[1157,1181,944,968]" italics="true" pageId="28" pageNumber="29">
s 
<number box="[1167,1181,944,968]" pageId="28" pageNumber="29" value="4.0">4</number>
</emphasis>
, 
<emphasis box="[1196,1224,944,968]" italics="true" pageId="28" pageNumber="29">
S 
<number box="[1211,1224,944,968]" pageId="28" pageNumber="29" value="2.0">2</number>
</emphasis>
, 
<emphasis box="[1239,1267,944,968]" italics="true" pageId="28" pageNumber="29">
S 
<number box="[1253,1267,944,968]" pageId="28" pageNumber="29" value="4.0">4</number>
</emphasis>
and macroseta of basitarsus IV (respectively 
<number box="[490,564,980,1005]" pageId="28" pageNumber="29" value="42.6">42, 60</number>
, 
<number box="[575,648,980,1005]" pageId="28" pageNumber="29" value="33.35">33, 35</number>
, 
<number box="[661,689,980,1004]" pageId="28" pageNumber="29" value="30.0">30</number>
and 
<quantity box="[745,803,980,1005]" metricMagnitude="0" metricUnit="m" metricValue="1.905" pageId="28" pageNumber="29" unit="in" value="75.0">75 in</quantity>
the 
<typeStatus box="[853,901,981,1005]" pageId="28" pageNumber="29">type</typeStatus>
specimens). Those differences are attributed to the larger size of the 
<typeStatus box="[391,439,1016,1040]" pageId="28" pageNumber="29">type</typeStatus>
specimen of 
<taxonomicName box="[592,694,1016,1040]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="segnis">
<emphasis box="[592,609,1016,1039]" italics="true" pageId="28" pageNumber="29">A</emphasis>
. 
<emphasis box="[624,694,1017,1040]" italics="true" pageId="28" pageNumber="29">segnis</emphasis>
</taxonomicName>
(according to the original description, dorsal shield 
<number box="[1288,1330,1016,1040]" pageId="28" pageNumber="29" value="340.0">340</number>
long and 
<number box="[151,193,1052,1076]" pageId="28" pageNumber="29" value="190.0">190</number>
wide), and thus 
<taxonomicName box="[379,527,1052,1077]" class="Arachnida" family="Phytoseiidae" genus="Amblyseius" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis box="[379,396,1052,1075]" italics="true" pageId="28" pageNumber="29">A</emphasis>
. 
<emphasis box="[409,527,1052,1077]" italics="true" pageId="28" pageNumber="29">seminudus</emphasis>
</taxonomicName>
is considered a junior synonym of 
<taxonomicName box="[922,1025,1053,1076]" class="Arachnida" family="Phytoseiidae" genus="Neoseiulus" kingdom="Animalia" order="Mesostigmata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="segnis">
<emphasis box="[922,941,1053,1076]" italics="true" pageId="28" pageNumber="29">N</emphasis>
. 
<emphasis box="[955,1025,1053,1076]" italics="true" pageId="28" pageNumber="29">segnis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>