161 lines
15 KiB
XML
161 lines
15 KiB
XML
<document id="09BE751390FF38A5E636E83F1271A955" ID-CLB-Dataset="28776" ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZooBank="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2018" docId="D7657B421ACAF4D00F17D9CE636622BE" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina fume Balke 1998" docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="19" updateTime="1701377518496" updateUser="ExternalLinkService">
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<mods:title id="2CF67CB0801F095771DA5C54F23D9B50">Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="5B809A8378308AE3170BEF453D0D40AC">Shaverdo, Helena</mods:namePart>
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<mods:roleTerm id="32EB47B91D985DEFFC2704252D172BBB">Author</mods:roleTerm>
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<mods:namePart id="1D17A2B46E02DC23A3D6755EE0FD229C">Sagata, Katayo</mods:namePart>
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<mods:namePart id="7750529AAC6B4F1CDE949AE2D695EAFE">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="3EC0B9F7C61557E81BAFAF4027EDE3D4">text</mods:typeOfResource>
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<mods:title id="82C20A1EB929FC872D933DB5A1886F62">ZooKeys</mods:title>
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<mods:part id="4989989B3F47052B7ADDC5AC0085686F">
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<mods:date id="D352B68148B62E74EC4A897EAB1D7802">2018</mods:date>
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<mods:detail id="49036845639BAEDEB4CD6C86D43C6B6B" type="volume">
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<mods:number id="2F49DB3ABB35342B15FAB30182B09481">803</mods:number>
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<mods:end id="83D95945D8786179B0AA35122A8CE4CB">70</mods:end>
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<mods:url id="D213FC84A0E50820712BA4CF1F7C8344">http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
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<mods:classification id="3450D63F93E312D3D917E4C51743E9D1">journal article</mods:classification>
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<mods:identifier id="5CBB485C0F0AB247DD4950ED618E9D3B" type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
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<mods:identifier id="F433C87FBA82AD4B066D2EC08B91DB38" type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
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<treatment id="D7657B421ACAF4D00F17D9CE636622BE" ID-GBIF-Taxon="154125595" LSID="urn:lsid:plazi:treatment:D7657B421ACAF4D00F17D9CE636622BE" httpUri="http://treatment.plazi.org/id/D7657B421ACAF4D00F17D9CE636622BE" lastPageNumber="19" pageId="12" pageNumber="19">
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<subSubSection id="88BA536284AFC8A5324E7F195E03FE66" pageId="12" pageNumber="19" type="nomenclature">
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<paragraph id="2A8F0655AE70F70F573BE3F269B19114" pageId="12" pageNumber="19">
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7.
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<taxonomicName id="C69FFF41D7452E0ADACBEB0E3CBD7E2A" ID-CoL="6HMQL" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume">Exocelina fume (Balke, 1998)</taxonomicName>
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Figs 3, 27
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</paragraph>
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</subSubSection>
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<subSubSection id="CE49058B6300EF25169C35127973CFDA" pageId="12" pageNumber="19" type="reference_group">
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<paragraph id="6BB2D4043DDAABEC97192AC27F2D35AD" pageId="12" pageNumber="19">
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<taxonomicName id="716A0389AA771DDCBD5F3C7326303941" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume" subGenus="Papuadytes">Copelatus (Papuadytes) fume</taxonomicName>
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Balke, 1998: 330;
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<bibRefCitation id="3ECB38546C905BA77F24BEC0A3DC91EA" author="Nilsson, AN" journalOrPublisher="Apollo Books, Stenstrup" pageId="39" pageNumber="46" title="Dytiscidae. World Catalogue of Insects Vol. 3." year="2001">Nilsson 2001</bibRefCitation>
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: 76 (catalogue).
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</paragraph>
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<paragraph id="B092C996EBAFAFCD4C46C01A6BD464FA" pageId="12" pageNumber="19">
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<taxonomicName id="0B06442E949A1258209624143D57739B" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume">Papuadytes fume</taxonomicName>
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(Balke, 1998):
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<bibRefCitation id="5DE1234B0588157F07FA1DF08531C301" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="39" pageNumber="46" pagination="55 - 74" title="World Catalogue of Dytiscidae-corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
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: 56 (comb. n.).
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</paragraph>
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<paragraph id="DFB8FF32C0378CCD1B9C0BBA421E7D60" pageId="12" pageNumber="19">
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<taxonomicName id="358B93B718D127D3B424A7077837C4C3" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume">Exocelina fume</taxonomicName>
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(Balke, 1998):
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<bibRefCitation id="D48D37C246FAEA3369F731CEFA93ACAC" pageId="12" pageNumber="19">Nilsson 2007</bibRefCitation>
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: 33 (comb. n.);
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<bibRefCitation id="AFCCD0A0ECDCC3576B8CB55F0A953BD8" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="40" pageNumber="47" url="http://waterbeetles.eu/documents/W_CAT_Dytiscidae_2017.pdf" year="2018">
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Nilsson and
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<normalizedToken id="42A6BE4B9338B94B27CCAAA7F7680308" originalValue="Hájek">Hajek</normalizedToken>
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2018
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</bibRefCitation>
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: 66 (catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection id="0905852728725C0440009832F8B44D48" pageId="12" pageNumber="19" type="type locality">
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<paragraph id="ABD7040C9579EB27DA61DD41D1D19EEA" pageId="12" pageNumber="19">Type locality.</paragraph>
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<paragraph id="DADDE24CFAD49BE800C07EB4D15920ED" pageId="12" pageNumber="19">
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Papua: Pegunungan Bintang Regency, Borme,
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<geoCoordinate id="E9E09CAF16E4B5F97B42A718E5809A6C" direction="south" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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,
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<geoCoordinate id="8E7FB1244525E016F5D8C5E58BBF057B" direction="east" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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, 1800 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="3C6355FF2BBC3462A2F3E91F8A4FE623" pageId="12" pageNumber="19" type="type material">
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<paragraph id="3B25CA52A4DC1B4097C89A93759B8503" pageId="12" pageNumber="19">Type material studied.</paragraph>
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<paragraph id="443DBC0D0E6393AE49707805C06C2CDA" pageId="12" pageNumber="19">
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Holotype: male "IRIAN JAYA Zentralmassive
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<geoCoordinate id="7639CC50AAE274D06652D704431F9651" direction="east" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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<geoCoordinate id="42D62772CE810F546FB9F7546EBF3699" direction="south" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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, "Borme, 1800m 16.8.1992 leg. Balke (12, 12A)",
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<normalizedToken id="7C417716B90D49176CBAB614288B7600" originalValue="“HOLOTYPUS”">"HOLOTYPUS"</normalizedToken>
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[red], "
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<taxonomicName id="D7559A3EFA2FF65B805222FB4D5A3A46" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume">Copelatus fume</taxonomicName>
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Balke des. 1997" [red] (NHMW). Paratypes: 9 males with the same label as the holotype and additionally with red labels "Paratypus
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<taxonomicName id="3387A5002DC74A9BB56EF009480CFFC5" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus fume" order="Coleoptera" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="fume">Copelatus fume</taxonomicName>
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Balke des. 1997", two of them with labels "M.Balke 3275" [green], and "M.Balke 3276" [green] and "M.Balke 6405" [green text] (NHMW). Note: There are two additional paratypes of
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<taxonomicName id="772939A78A0A31D65BE0E3D4939CDA87" lsidName="E. fume" pageId="12" pageNumber="19" rank="species" species="fume">E. fume</taxonomicName>
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, which do not belong to this species but to
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<taxonomicName id="A1BDBB120C860EA3D5D6BDB757EF794E" lsidName="E. ketembang" pageId="12" pageNumber="19" rank="species" species="ketembang">E. ketembang</taxonomicName>
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(Balke, 1998) and
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<taxonomicName id="A5684393F0C2AFD122E18417B00AA212" lsidName="E. erteldi" pageId="12" pageNumber="19" rank="species" species="erteldi">E. erteldi</taxonomicName>
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(Balke, 1998).
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</paragraph>
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</subSubSection>
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<subSubSection id="7E5E4389B128AA198350D39847C4E82C" pageId="12" pageNumber="19" type="diagnosis">
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<paragraph id="C235AD42942471A98264C949E71BDD18" pageId="12" pageNumber="19">Diagnosis.</paragraph>
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<paragraph id="23EB1B9BD8D7F9D4A1226182300A212F" pageId="12" pageNumber="19">
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For complete description, see
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<bibRefCitation id="9E19C9EB44FA86FF95D5A2C9450C2A3B" pageId="12" pageNumber="19">Balke (1998</bibRefCitation>
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: 330). Beetle medium-sized: TL-H 3.7-4.4 mm; oblong-oval; brown to dark brown, with reddish brown pronotal sides, head, and sometimes also sides of elytra, in some specimens, disc of pronotum and elytron almost piceous; submatt, with fine but rather dense punctation and strongly impressed microreticulation; pronotum without lateral bead; male antennae simple (Fig. 3); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, with anterior band of more than 60 and posterior row of 6 relatively long, thin setae (Fig. 27D); median lobe in lateral view slightly curved, with apex curved downwards, with visible angle on dorsal side, in ventral view, distally distinctly narrowed to truncate apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae long, only slightly sparser than subdistal ones (Fig. 27
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<normalizedToken id="24084250A9A2B5B4419572B9DD62872E" originalValue="A–C">A-C</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0872DBBF0287BF7A08B33B99EBA79070" pageId="12" pageNumber="19" type="affinities">
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<paragraph id="2A849B445B3ED4BD9F7B5640B47379B5" pageId="12" pageNumber="19">Affinities.</paragraph>
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<paragraph id="416D65CBA264A7EE6682493E63C696B5" pageId="12" pageNumber="19">
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In the area of its distribution,
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<taxonomicName id="9D84F35141AB66C3DC3671453023CEC9" lsidName="E. fume" pageId="12" pageNumber="19" rank="species" species="fume">E. fume</taxonomicName>
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co-occurs with
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<taxonomicName id="964417623F154A069FA6A553C756E46B" lsidName="E. takime" pageId="12" pageNumber="19" rank="species" species="takime">E. takime</taxonomicName>
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(Balke, 1998) and species of the
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<taxonomicName id="2178F01F5CD6ED1F7E08A3C14B5B108E" lsidName="ekari" pageId="12" pageNumber="19" rank="species" species="ekari">ekari</taxonomicName>
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-, aipo-, okbapensis-, aipomek-,
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<taxonomicName id="C59CE8B65AD40A04CAE3B5C036319105" lsidName="erteldi" pageId="12" pageNumber="19" rank="species" species="erteldi">erteldi</taxonomicName>
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-, and
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<taxonomicName id="044B941339D54ED35FF662A02AA6319E" lsidName="danae" pageId="12" pageNumber="19" rank="species" species="danae">danae</taxonomicName>
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-groups. From species of the
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<taxonomicName id="E6A3F8100D6D9EB9D9FC42B8F729CB40" lsidName="ekari" pageId="12" pageNumber="19" rank="species" species="ekari">ekari</taxonomicName>
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-group, the species differs in larger size, evidently stronger dorsal punctation and microreticulation, and the shape of the median lobe. In the latter two characters,
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<taxonomicName id="757A95154211710EAAEB14705E2F186B" lsidName="E. fume" pageId="12" pageNumber="19" rank="species" species="fume">E. fume</taxonomicName>
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differs also from the species of the remaining groups, as well as in absence of the pronotal bead and simple male antennae.
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</paragraph>
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<paragraph id="AC815334F5F3FC738C96A8C4DEFF578A" pageId="12" pageNumber="19">
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Within the
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<taxonomicName id="9D009D864AD18DC8796B4130E37C0568" lsidName="casuarina" pageId="12" pageNumber="19" rank="species" species="casuarina">casuarina</taxonomicName>
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-group, the species is more similar to
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<taxonomicName id="AF1FDB5636FEDB0D1B7953808EEF74D2" lsidName="E. casuarina" pageId="12" pageNumber="19" rank="species" species="casuarina">E. casuarina</taxonomicName>
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and
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<taxonomicName id="E8563BC72BF50B136FA5E9243FDC1CEC" lsidName="E. ibalimi" pageId="12" pageNumber="19" rank="species" species="ibalimi">E. ibalimi</taxonomicName>
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sp. n., especially the latter one, from which can be distinguished by paler coloration and the shape of the median lobe (see their
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<normalizedToken id="81E8B4923F212DB8E7F83914EA21A28C" originalValue="“Affinities”">"Affinities"</normalizedToken>
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and the
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<normalizedToken id="D764FD8C216D3F25A1D6AD37A95A949E" originalValue="“Key”">"Key"</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="EF449E50C78686947430E5ABDB59FE3E" pageId="12" pageNumber="19" type="distribution">
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<paragraph id="82A5F45B320093C3B5D2D409870BBC93" pageId="12" pageNumber="19">Distribution.</paragraph>
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<paragraph id="40709C031FBB80F9492C82EE7CFC4252" pageId="12" pageNumber="19">Papua: Pegunungan Bintang. The species is known only from the type locality (Fig. 50).</paragraph>
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</subSubSection>
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</treatment>
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</document> |