204 lines
21 KiB
XML
204 lines
21 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.4.6.1" ID-GBIF-Dataset="3c53535f-d5c5-44af-8421-a9c7750b8cf5" ID-ISSN="2624-2834" ID-Zenodo-Dep="5780148" checkinTime="1639468614706" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Ansorge, Jörg & Khramov, Alexander V." docDate="2021" docId="5849C365FFCCFFA95A53F96CAC6FA6DF" docLanguage="en" docName="palaeoentomology.4.6.1.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.3:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="3" docTitle="Epigambria longipennis Handlirsch 1939" docType="treatment" docVersion="5" lastPageNumber="518" masterDocId="A470BB1DFFCDFFAB5979FFA7A90EA051" masterDocTitle="Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera)" masterLastPageNumber="531" masterPageNumber="516" pageNumber="517" updateTime="1673523423440" updateUser="felipe">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ansorge, Jörg</mods:namePart>
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<mods:affiliation>Institute of Geography and Geology, University of Greifswald, Friedrich-Ludwig-Jahn-Strasse 17 a, D- 17489 Greifswald, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Khramov, Alexander V.</mods:namePart>
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<mods:affiliation>Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya Str. 123, 117997, Moscow, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>516</mods:start>
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<mods:end>531</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.4.6.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3c53535f-d5c5-44af-8421-a9c7750b8cf5</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5780148</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5779456" ID-GBIF-Taxon="191567584" ID-Zenodo-Dep="5779456" LSID="urn:lsid:plazi:treatment:5849C365FFCCFFA95A53F96CAC6FA6DF" httpUri="http://treatment.plazi.org/id/5849C365FFCCFFA95A53F96CAC6FA6DF" lastPageId="2" lastPageNumber="518" pageId="1" pageNumber="517">
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<subSubSection box="[810,1300,1739,1766]" pageId="1" pageNumber="517" type="nomenclature">
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<paragraph blockId="1.[810,1300,1739,1802]" box="[810,1300,1739,1766]" pageId="1" pageNumber="517">
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<heading bold="true" box="[810,1300,1739,1766]" fontSize="10" level="2" pageId="1" pageNumber="517" reason="3">
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<taxonomicName ID-CoL="6G5GV" authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[810,1300,1739,1766]" class="Insecta" family="Epigambriidae" genus="Epigambria" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="longipennis">
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<emphasis bold="true" box="[810,1300,1739,1766]" pageId="1" pageNumber="517">
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<emphasis bold="true" box="[810,1089,1739,1766]" italics="true" pageId="1" pageNumber="517">Epigambrialongipennis</emphasis>
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<bibRefCitation author="Handlirsch, A." bookContentInfo="16 pls." box="[1096,1300,1739,1765]" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="1" pageNumber="517" pagination="1 - 240" part="49" refId="ref9191" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil. Annalen des Naturhistorischen Museums in Wien, 49, 1 - 240, 16 pls." title="Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil" type="book chapter" year="1939">Handlirsch, 1939</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[810,894,1775,1802]" pageId="1" pageNumber="517" type="description">
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<paragraph blockId="1.[810,1300,1739,1802]" box="[810,894,1775,1802]" pageId="1" pageNumber="517">
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(
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<figureCitation box="[818,882,1775,1802]" captionStart="FIGURE 1" captionStartId="2.[136,235,1224,1248]" captionTargetBox="[312,1276,189,1190]" captionTargetId="figure-285@2.[312,1276,189,1201]" captionTargetPageId="2" captionText="FIGURE 1. Epigambria longipennis Handlirsch, 1939, holotype FGWG 123/51. A, Part. B, counterpart. C, Hind wing venation. Scale bars = 1 mm (A, B to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5780150" httpUri="https://zenodo.org/record/5780150/files/figure.png" pageId="1" pageNumber="517">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="517" type="reference_group">
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<paragraph blockId="1.[810,1452,1851,1981]" pageId="1" pageNumber="517">
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<treatmentCitationGroup pageId="1" pageNumber="517">
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<treatmentCitation author="Handlirsch, A." box="[810,1276,1851,1873]" page="76" pageId="1" pageNumber="517" year="1939">
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<taxonomicName ID-CoL="6G5GV" authority="Handlirsch, 1939: 76" authorityName="Handlirsch" authorityPageNumber="76" authorityYear="1939" box="[810,1276,1851,1873]" class="Insecta" family="Epigambriidae" genus="Epigambria" kingdom="Animalia" order="Neuroptera" pageId="1" pageNumber="517" phylum="Arthropoda" rank="species" species="longipennis">
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<emphasis box="[810,1048,1851,1872]" italics="true" pageId="1" pageNumber="517">Epigambrialongipennis</emphasis>
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<bibRefCitation author="Handlirsch, A." bookContentInfo="16 pls." box="[1060,1276,1851,1873]" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="1" pageNumber="517" pagination="1 - 240" part="49" refId="ref9191" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil. Annalen des Naturhistorischen Museums in Wien, 49, 1 - 240, 16 pls." title="Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil" type="book chapter" year="1939">Handlirsch, 1939: 76</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, Pl. 7, Fig. 119;
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<treatmentCitation author="Martynova, O. M." box="[863,1085,1887,1909]" page="167" pageId="1" pageNumber="517" year="1949">
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<bibRefCitation author="Martynova, O. M." box="[863,1085,1887,1909]" journalOrPublisher="Trudy Paleontologicheskogo Instituta" pageId="1" pageNumber="517" pagination="150 - 170" part="20" refId="ref10394" refString="Martynova, O. M. (1949) Mesozoic lacewings (Neuroptera) and their bearing on concepts of phylogeny and systematics of the order. Trudy Paleontologicheskogo Instituta, 20, 150 - 170. [In Russian]" title="Mesozoic lacewings (Neuroptera) and their bearing on concepts of phylogeny and systematics of the order" type="journal article" year="1949">Martynova, 1949: 167</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Lambkin, K. J." box="[1097,1298,1887,1909]" page="449" pageId="1" pageNumber="517" year="1988">
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<bibRefCitation author="Lambkin, K. J." box="[1097,1298,1887,1909]" journalOrPublisher="Memoirs of the Queensland Museum" pageId="1" pageNumber="517" pagination="445 - 458" part="25" refId="ref9621" refString="Lambkin, K. J. (1988) A re-examination of Lithosmylidia Riek from the Triassic of Queensland with notes on Mesozoic ' osmylidlike' fossil Neuroptera (Insecta: Neuroptera). Memoirs of the Queensland Museum, 25, 445 - 458." title="A re-examination of Lithosmylidia Riek from the Triassic of Queensland with notes on Mesozoic ' osmylidlike' fossil Neuroptera (Insecta: Neuroptera)" type="journal article" year="1988">Lambkin, 1988: 449</bibRefCitation>
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</treatmentCitation>
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, 453;
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<treatmentCitation author="Herrig, E. & Nestler, H." page="33" pageId="1" pageNumber="517" year="1989">
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<bibRefCitation author="Herrig, E. & Nestler, H." bookContentInfo="80 pp." journalOrPublisher="Ernst-Moritz-Arndt-Universitat, Sektion Geologische Wissenschaften, Greifswald" pageId="1" pageNumber="517" refId="ref9239" refString="Herrig, E. & Nestler, H. (1989) Katalog der palaozoologischen Typen. Ernst-Moritz-Arndt-Universitat, Sektion Geologische Wissenschaften, Greifswald, 80 pp." title="Katalog der palaozoologischen Typen" type="book" year="1989">Herrig & Nestler,1989:33</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Carpenter, F. M." box="[1031,1230,1923,1945]" page="355" pageId="1" pageNumber="517" year="1992">
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<bibRefCitation author="Carpenter, F. M." box="[1031,1230,1923,1945]" editor="Kaesler, R. L." journalOrPublisher="vol." pageId="1" pageNumber="517" pagination="279 - 655" part="4" refId="ref8788" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vols. 3 and 4. Geological Society of America, Boulder, CO & University of Kansas, Lawrence, KS, xxi + 1 - 277 [vol. 3], ii + 279 - 655 [vol. 4]." title="Superclass Hexapoda" type="journal article" volumeTitle="Treatise on Invertebrate Paleontology. Part R. Arthropoda" year="1992">Carpenter,1992:355</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Ross, A. J. & Jarzembowski, E. A." page="418" pageId="1" pageNumber="517" year="1993">
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<bibRefCitation author="Ross, A. J. & Jarzembowski, E. A." editor="Benton, M. J." journalOrPublisher="Chapman and Hall, London" pageId="1" pageNumber="517" pagination="363 - 426" refId="ref11185" refString="Ross, A. J. & Jarzembowski, E. A. (1993) Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (Ed.), The fossil record 2. Chapman and Hall, London, pp. 363 - 426." title="Arthropoda (Hexapoda; Insecta)" type="book chapter" volumeTitle="The fossil record 2" year="1993">Ross& Jarzembowski, 1993: 418</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Makarkin, V. N. & Archibald, S. B." box="[972,1303,1959,1981]" page="176" pageId="1" pageNumber="517" year="2003">
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<bibRefCitation author="Makarkin, V. N. & Archibald, S. B." box="[972,1303,1959,1981]" journalOrPublisher="Annals of the Entomological Society of America" pageId="1" pageNumber="517" pagination="171 - 180" part="96" refId="ref10018" refString="Makarkin, V. N. & Archibald, S. B. (2003) Family affinity of the genus Palaeopsychops Andersen with description of a new species from the Early Eocene of British Columbia (Neuroptera: Polystoechotidae). Annals of the Entomological Society of America, 96, 171 - 180." title="Family affinity of the genus Palaeopsychops Andersen with description of a new species from the Early Eocene of British Columbia (Neuroptera: Polystoechotidae)" type="journal article" year="2003">Makarkin & Archibald, 2003: 176</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="518" pageId="1" pageNumber="517" type="materials_examined">
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<paragraph blockId="1.[810,1451,2027,2089]" lastBlockId="2.[136,778,1350,2093]" lastPageId="2" lastPageNumber="518" pageId="1" pageNumber="517">
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<emphasis bold="true" box="[810,921,2027,2053]" pageId="1" pageNumber="517">Material.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3418314302" country="Germany" county="University of Greifswald" lastPageId="2" lastPageNumber="518" location="Institute of Geography" municipality="Geology" pageId="1" pageNumber="517" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[937,1041,2027,2053]" pageId="1" pageNumber="517">Holotype</typeStatus>
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FGWG 123/51, deposited in the collections of the
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<location LSID="urn:lsid:plazi:treatment:5849C365FFCCFFA95A53F96CAC6FA6DF:D53F24A8FFCCFFAA5A86F7A8AC08A878" box="[1023,1286,2063,2089]" country="Germany" county="University of Greifswald" municipality="Geology" name="Institute of Geography" pageId="1" pageNumber="517">Institute of Geography</location>
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and
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<collectingMunicipality box="[1350,1447,2063,2089]" pageId="1" pageNumber="517">Geology</collectingMunicipality>
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,
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<collectingCounty box="[136,425,1350,1376]" pageId="2" pageNumber="518">University of Greifswald</collectingCounty>
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,
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<collectingCountry box="[439,545,1350,1376]" name="Germany" pageId="2" pageNumber="518">Germany</collectingCountry>
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. Apoorly preserved crumpled complete hind wing.
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<collectingCountry box="[528,637,1388,1414]" name="Germany" pageId="2" pageNumber="518">Germany</collectingCountry>
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: Dobbertin. Early Jurassic (early Toarcian, “Grüne Serie”,
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<taxonomicName baseAuthorityName="Sowerby" baseAuthorityYear="1820" class="Cephalopoda" family="Hildoceratidae" genus="Harpoceras" kingdom="Animalia" order="Ammonoidea" pageId="2" pageNumber="518" phylum="Mollusca" rank="species" species="falciferum">
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<emphasis italics="true" pageId="2" pageNumber="518">Harpoceras falciferum</emphasis>
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</taxonomicName>
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ammonite zone).
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</materialsCitation>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5780150" ID-Zenodo-Dep="5780150" httpUri="https://zenodo.org/record/5780150/files/figure.png" pageId="2" pageNumber="518" startId="2.[136,235,1224,1248]" targetBox="[312,1276,189,1190]" targetPageId="2">
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<paragraph blockId="2.[136,1452,1224,1285]" pageId="2" pageNumber="518">
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<emphasis bold="true" box="[136,260,1224,1248]" pageId="2" pageNumber="518">FIGURE 1.</emphasis>
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<taxonomicName authority="Handlirsch, 1939" authorityName="Handlirsch" authorityYear="1939" box="[267,697,1224,1249]" class="Insecta" family="Epigambriidae" genus="Epigambria" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="518" phylum="Arthropoda" rank="species" species="longipennis">
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<emphasis box="[267,513,1224,1248]" italics="true" pageId="2" pageNumber="518">Epigambrialongipennis</emphasis>
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<bibRefCitation author="Handlirsch, A." bookContentInfo="16 pls." box="[520,697,1224,1249]" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="2" pageNumber="518" pagination="1 - 240" part="49" refId="ref9191" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil. Annalen des Naturhistorischen Museums in Wien, 49, 1 - 240, 16 pls." title="Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil" type="book chapter" year="1939">Handlirsch, 1939</bibRefCitation>
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</taxonomicName>
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, holotype FGWG 123/51.
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<emphasis bold="true" box="[968,986,1224,1248]" pageId="2" pageNumber="518">A</emphasis>
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, Part.
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<emphasis bold="true" box="[1053,1070,1225,1249]" pageId="2" pageNumber="518">B</emphasis>
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, counterpart.
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<emphasis bold="true" box="[1212,1230,1224,1248]" pageId="2" pageNumber="518">C</emphasis>
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, Hindwing venation. Scale bars = 1 mm (
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<emphasis bold="true" box="[340,358,1260,1284]" pageId="2" pageNumber="518">A</emphasis>
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,
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<emphasis bold="true" box="[371,388,1261,1285]" pageId="2" pageNumber="518">B</emphasis>
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to same scale).
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</paragraph>
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</caption>
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<paragraph blockId="2.[136,778,1350,2093]" lastBlockId="2.[810,1452,1350,1678]" pageId="2" pageNumber="518">
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<emphasis bold="true" box="[189,372,1501,1527]" pageId="2" pageNumber="518">Re-description.</emphasis>
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Hindwing 6.9mmlong,1.8 mmwide aspreserved (estimated completewidth ≥
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<quantity box="[615,685,1539,1565]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="518" unit="mm" value="2.0">2 mm</quantity>
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). Costal space relatively broad, slightly broadened proximally, moderately broadened distally. Subcostal veinlets widely spaced; basally simple, perpendicular to Sc; more proximally bent toward wing apex, simple or shallowly forked; distally once forked. Subcostal space moderately broad; crossveins not detected. Sc stout, approaching
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<collectionCode box="[136,173,1803,1829]" pageId="2" pageNumber="518">RA</collectionCode>
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distally, with or without fusion unknown.
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<collectionCode box="[669,706,1803,1829]" pageId="2" pageNumber="518">RA</collectionCode>
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stout, relatively long, entering margin somewhat proximad wing apex; with four long veinlets, once to twice forked.
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<collectionCode box="[136,173,1916,1942]" pageId="2" pageNumber="518">RA</collectionCode>
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space narrow, with four preserved crossveins (nearly perpendicular to
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<collectionCode box="[325,368,1953,1979]" pageId="2" pageNumber="518">RA</collectionCode>
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).
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<collectionCode box="[386,419,1954,1980]" pageId="2" pageNumber="518">RP</collectionCode>
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originating relatively near wing base, with seven pectinate branches.
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<collectionCode box="[545,586,1991,2017]" pageId="2" pageNumber="518">RP</collectionCode>
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1,
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<collectionCode box="[606,644,1991,2017]" pageId="2" pageNumber="518">RP</collectionCode>
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3,
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<collectionCode box="[668,705,1991,2017]" pageId="2" pageNumber="518">RP</collectionCode>
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4,
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<collectionCode box="[729,764,1991,2017]" pageId="2" pageNumber="518">RP</collectionCode>
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6 deeply dichotomously forked;
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<collectionCode box="[486,521,2029,2055]" pageId="2" pageNumber="518">RP</collectionCode>
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2 similarly (but not so deep) branched;
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<collectionCode box="[321,357,2066,2093]" pageId="2" pageNumber="518">RP</collectionCode>
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5 once forked. Proximal part of
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<collectionCode box="[711,778,2067,2093]" pageId="2" pageNumber="518">Mnot</collectionCode>
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preserved.
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<collectionCode box="[936,984,1350,1376]" pageId="2" pageNumber="518">MA</collectionCode>
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,
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<collectionCode box="[998,1037,1351,1377]" country="0" name="Mohonk Preserve, Inc." pageId="2" pageNumber="518">MP</collectionCode>
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deeply dichotomously forked.
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<collectionCode box="[1400,1451,1350,1376]" pageId="2" pageNumber="518">CuA</collectionCode>
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pectinately branched; its anterior trace, three branches once forked.
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<collectionCode box="[966,1013,1426,1452]" pageId="2" pageNumber="518">CuP</collectionCode>
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not clearly discernible (possibly few branched). Anal veins very poorly preserved. Crossveins in radial space very poorly preserved, probably forming one gradate series (three crossveins discernible).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="518" type="discussion">
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<paragraph blockId="2.[810,1452,1350,1678]" pageId="2" pageNumber="518">
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<emphasis bold="true" box="[863,979,1576,1602]" pageId="2" pageNumber="518">Remarks.</emphasis>
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The drawing of Handlirsch (
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<bibRefCitation author="Handlirsch, A." bookContentInfo="16 pls." journalOrPublisher="Annalen des Naturhistorischen Museums in Wien" pageId="2" pageNumber="518" pagination="1 - 240" part="49" refId="ref9191" refString="Handlirsch, A. (1939) Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil. Annalen des Naturhistorischen Museums in Wien, 49, 1 - 240, 16 pls." title="Neue Untersuchungen uber die fossilen Insekten mit Erganzungen und Nachtragen sowie Ausblicken auf phylogenetische, palaeogeographische und allgemein biologische Probleme. II. Theil" type="book chapter" year="1939">Handlirsch, 1939</bibRefCitation>
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: Pl. 7, Fig. 119) is largely incorrect, especially in the area of the cubital space; we revise it here (
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<figureCitation box="[1295,1360,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[136,235,1224,1248]" captionTargetBox="[312,1276,189,1190]" captionTargetId="figure-285@2.[312,1276,189,1201]" captionTargetPageId="2" captionText="FIGURE 1. Epigambria longipennis Handlirsch, 1939, holotype FGWG 123/51. A, Part. B, counterpart. C, Hind wing venation. Scale bars = 1 mm (A, B to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5780150" httpUri="https://zenodo.org/record/5780150/files/figure.png" pageId="2" pageNumber="518">Fig. 1</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |