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<document ID-DOI="10.11646/palaeoentomology.4.6.1" ID-GBIF-Dataset="3c53535f-d5c5-44af-8421-a9c7750b8cf5" ID-ISSN="2624-2834" ID-Zenodo-Dep="5780148" checkinTime="1639468614706" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Ansorge, Jörg &amp; Khramov, Alexander V." docDate="2021" docId="5849C365FFC5FFA25A53FABBABCAA59E" docLanguage="en" docName="palaeoentomology.4.6.1.pdf" docOrigin="Palaeoentomology 4 (6)" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.3:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="3" docTitle="Epigambriinae Handlirsch 1939" docType="treatment" docVersion="6" lastPageNumber="525" masterDocId="A470BB1DFFCDFFAB5979FFA7A90EA051" masterDocTitle="Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera)" masterLastPageNumber="531" masterPageNumber="516" pageNumber="524" updateTime="1673523423440" updateUser="felipe">
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<mods:titleInfo>
<mods:title>Revision of Epigambriinae Handlirsch, stat. nov., a subfamily of Early Jurassic Ithonidae s. l. (Neuroptera)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Makarkin, Vladimir N.</mods:namePart>
<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ansorge, Jörg</mods:namePart>
<mods:affiliation>Institute of Geography and Geology, University of Greifswald, Friedrich-Ludwig-Jahn-Strasse 17 a, D- 17489 Greifswald, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khramov, Alexander V.</mods:namePart>
<mods:affiliation>Borissiak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya Str. 123, 117997, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Palaeoentomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>516</mods:start>
<mods:end>531</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/palaeoentomology.4.6.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">3c53535f-d5c5-44af-8421-a9c7750b8cf5</mods:identifier>
<mods:identifier type="ISSN">2624-2834</mods:identifier>
<mods:identifier type="Zenodo-Dep">5780148</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5779470" ID-GBIF-Taxon="191567592" ID-Zenodo-Dep="5779470" LSID="urn:lsid:plazi:treatment:5849C365FFC5FFA25A53FABBABCAA59E" httpUri="http://treatment.plazi.org/id/5849C365FFC5FFA25A53FABBABCAA59E" lastPageId="9" lastPageNumber="525" pageId="8" pageNumber="524">
<subSubSection box="[810,1189,1308,1334]" pageId="8" pageNumber="524" type="nomenclature">
<paragraph blockId="8.[810,1189,1308,1370]" box="[810,1189,1308,1334]" pageId="8" pageNumber="524">
<heading bold="true" box="[810,1189,1308,1334]" fontSize="10" level="2" pageId="8" pageNumber="524" reason="3">
<emphasis bold="true" box="[810,1189,1308,1334]" pageId="8" pageNumber="524">
<taxonomicName authorityName="Handlirsch" authorityYear="1939" box="[810,989,1308,1334]" family="Ithonidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="517" rank="subFamily" subFamily="Epigambriinae">Epigambriinae</taxonomicName>
gen. et sp. indet.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[810,893,1344,1370]" pageId="8" pageNumber="524" type="description">
<paragraph blockId="8.[810,1189,1308,1370]" box="[810,893,1344,1370]" pageId="8" pageNumber="524">
(
<figureCitation box="[818,884,1344,1370]" captionStart="FIGURE 8" captionStartId="9.[136,235,1084,1108]" captionTargetBox="[312,1276,189,1051]" captionTargetId="figure-384@9.[312,1276,189,1061]" captionTargetPageId="9" captionText="FIGURE 8. Epigambriinae gen. et sp. indet., PIN 5552/48. A, Photograph. B, Hind wing venation. Scale bar = 1 mm (both to same scale)." figureDoi="http://doi.org/10.5281/zenodo.5780164" httpUri="https://zenodo.org/record/5780164/files/figure.png" pageId="8" pageNumber="524">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="524" type="materials_examined">
<paragraph blockId="8.[810,1452,1416,2092]" pageId="8" pageNumber="524">
<emphasis bold="true" box="[810,921,1416,1442]" pageId="8" pageNumber="524">Material.</emphasis>
PIN 5552/48, deposited in PIN. An incomplete crumpled hind wing.
</paragraph>
<paragraph blockId="8.[810,1452,1416,2092]" pageId="8" pageNumber="524">
<emphasis bold="true" box="[863,1120,1488,1515]" pageId="8" pageNumber="524">Locality and horizon.</emphasis>
<collectingCountry box="[1128,1237,1489,1515]" name="Germany" pageId="8" pageNumber="524">Germany</collectingCountry>
: Schandelah. Early Jurassic (early Toarcian,
<emphasis box="[1117,1202,1525,1551]" italics="true" pageId="8" pageNumber="524">boreale</emphasis>
nodule,
<taxonomicName baseAuthorityName="Sowerby" baseAuthorityYear="1820" class="Cephalopoda" family="Hildoceratidae" genus="Harpoceras" kingdom="Animalia" order="Ammonoidea" pageId="8" pageNumber="524" phylum="Mollusca" rank="species" species="falciferum">
<emphasis italics="true" pageId="8" pageNumber="524">Harpoceras falciferum</emphasis>
</taxonomicName>
ammonite zone).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="525" pageId="8" pageNumber="524" type="description">
<paragraph blockId="8.[810,1452,1416,2092]" lastBlockId="9.[136,778,1203,1487]" lastPageId="9" lastPageNumber="525" pageId="8" pageNumber="524">
<emphasis bold="true" box="[863,1009,1597,1623]" pageId="8" pageNumber="524">Description.</emphasis>
Hind wing
<quantity box="[1151,1299,1597,1624]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="8" pageNumber="524" unit="mm" value="5.8">5.8 mmlong</quantity>
as preserved (estimated completelength
<emphasis box="[1115,1148,1634,1659]" italics="true" pageId="8" pageNumber="524">ca.</emphasis>
<quantity box="[1153,1291,1633,1660]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="8" pageNumber="524" unit="mm" value="6.35" valueMax="6.5" valueMin="6.2">6.26.5 mm</quantity>
),
<quantity box="[1308,1451,1633,1660]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="524" unit="mm" value="2.3">2.3 mmwide</quantity>
aspreserved(estimated completelength
<emphasis box="[1249,1275,1670,1695]" italics="true" pageId="8" pageNumber="524">ca</emphasis>
. 2.5).Preserved portion of costal space relatively broad; broadened distally. Subcostal veinlets mainly oblique; simple or once shallowly forked medially; once to twice forked distally. Subcostal space rather narrow as preserved. Sc, RA stout. RA space narrow, with three preserved crossveins nearly perpendicular to RA (distal slightly oblique). RP with seven branches. RP1RP3 deeply dichotomously forked; other branches shallowly forked or their terminations not preserved. MA, MP dichotomously forked. CuA pectinate with at least five branches. CuP not completely preserved, probably few branched. Presumed A1, A2 each shallowly forked once (alternatively, A1 dichotomously forked, A2 not preserved). One distal crossvein between presumed A1, A2.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5780164" ID-Zenodo-Dep="5780164" httpUri="https://zenodo.org/record/5780164/files/figure.png" pageId="9" pageNumber="525" startId="9.[136,235,1084,1108]" targetBox="[312,1276,189,1051]" targetPageId="9">
<paragraph blockId="9.[136,1451,1084,1144]" pageId="9" pageNumber="525">
<emphasis bold="true" box="[136,259,1084,1108]" pageId="9" pageNumber="525">FIGURE 8.</emphasis>
<taxonomicName authorityName="Handlirsch" authorityYear="1939" box="[265,414,1084,1108]" pageId="9" pageNumber="517" rank="subFamily" subFamily="Epigambriinae">Epigambriinae</taxonomicName>
gen. et sp. indet., PIN 5552/48.
<emphasis bold="true" box="[736,754,1084,1108]" pageId="9" pageNumber="525">A</emphasis>
, Photograph.
<emphasis bold="true" box="[894,911,1084,1108]" pageId="9" pageNumber="525">B</emphasis>
, Hind wing venation. Scale bar = 1 mm (both to same scale).
</paragraph>
</caption>
<subSubSection pageId="9" pageNumber="525" type="discussion">
<paragraph blockId="9.[136,778,1203,1487]" pageId="9" pageNumber="525">
<emphasis bold="true" box="[189,305,1314,1340]" pageId="9" pageNumber="525">Remarks.</emphasis>
The venation of this hind wing is very similar to that of
<taxonomicName authorityName="Handlirsch" authorityYear="1939" box="[339,614,1350,1376]" class="Insecta" family="Epigambriidae" genus="Epigambria" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="525" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis box="[339,614,1350,1376]" italics="true" pageId="9" pageNumber="525">Epigambria longipennis</emphasis>
</taxonomicName>
and is clearly distinguished from that of
<taxonomicName authorityName="Ansorge" authorityYear="1996" box="[453,699,1387,1413]" class="Insecta" genus="Polyosmylus" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="525" phylum="Arthropoda" rank="species" species="excelsus">
<emphasis box="[453,699,1387,1413]" italics="true" pageId="9" pageNumber="525">Polyosmylus excelsus</emphasis>
</taxonomicName>
by its dichotomously branched M. Judging from its size, this wing might also belong to
<taxonomicName authority="Makarkin &amp; Ansorge &amp; Khramov, 2021" authorityName="Makarkin &amp; Ansorge &amp; Khramov" authorityYear="2021" box="[435,595,1462,1487]" class="Insecta" family="Nymphitidae" genus="Rasnitsyneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="525" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis box="[435,595,1462,1487]" italics="true" pageId="9" pageNumber="525">Rasnitsyneura</emphasis>
</taxonomicName>
<emphasis bold="true" box="[603,708,1461,1487]" pageId="9" pageNumber="525">
<taxonomicNameLabel box="[603,708,1461,1487]" pageId="9" pageNumber="525" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>