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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.816.29738" ID-GBIF-Dataset="31bce27d-7094-467b-95ae-009a27e00cf5" ID-PMC="PMC6341056" ID-Pensoft-Pub="1313-2970-816-1" ID-PubMed="30686917" ID-ZBK="51CEEF2E1E1040A8A6731140426ED5A7" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-816-1" ModsDocOrigin="ZooKeys 816" ModsDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" checkinTime="1547561011487" checkinUser="pensoft" docAuthor="Hunting, Wesley &amp; Yang, Man-Miao" docDate="2019" docId="16C33D0D2240E23023FA8B0A11211DAD" docLanguage="en" docName="ZooKeys 816: 1-164" docOrigin="ZooKeys 816" docSource="http://dx.doi.org/10.3897/zookeys.816.29738" docTitle="Amphimenes beichatiensis Hunting &amp; Yang, 2019, sp. n." docType="treatment" docUuid="9296F080-66B8-42A0-B244-A59A5B9EACE1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="17" masterDocId="AA26FF84157BFFAE490E7F133744FF98" masterDocTitle="A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)" masterLastPageNumber="164" masterPageNumber="1" pageNumber="16" updateTime="1668166739332" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A taxonomic review of the pericaline ground-beetles in Taiwan, with descriptions of new species (Coleoptera, Carabidae, Lebiini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunting, Wesley</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Man-Miao</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>816</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>164</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.816.29738</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.816.29738</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-816-1</mods:identifier>
<mods:identifier type="ZBK">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
<mods:identifier type="ZooBank">51CEEF2E1E1040A8A6731140426ED5A7</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="154125992" LSID="urn:lsid:zoobank.org:act:9296F080-66B8-42A0-B244-A59A5B9EACE1" httpUri="http://treatment.plazi.org/id/16C33D0D2240E23023FA8B0A11211DAD" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection pageId="15" pageNumber="16" type="nomenclature">
<paragraph pageId="15" pageNumber="16">
<taxonomicName LSID="http://zoobank.org/9296F080-66B8-42A0-B244-A59A5B9EACE1" class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes beichatiensis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beichatiensis">Amphimenes beichatiensis</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="16">sp. n.</taxonomicNameLabel>
Figs 6, 7
<normalizedToken originalValue="AC">A-C</normalizedToken>
, 8A, 12
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="specific epithet">
<paragraph pageId="15" pageNumber="16">Specific epithet.</paragraph>
<paragraph pageId="15" pageNumber="16">The name of this species refers to the locality, Beichatien Mountain (北插天山), where the single specimen was collected.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="type material">
<paragraph pageId="15" pageNumber="16">Types and other material examined.</paragraph>
<paragraph pageId="15" pageNumber="16">
Holotype (male) labeled
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
[circular, ringed with red]; &quot;TAIWAN: Taoyuan Co./Fuxing township,/Beichatien Mountain/24.7906N, 121.4410E&quot;; &quot;hand collecting, night/~1320m,/Acc. Ti-96b /June 17, 2011/Coll. W. M. Hunting&quot;; &quot; NCHU/100698&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="type locality">
<paragraph pageId="15" pageNumber="16">Type locality.</paragraph>
<paragraph pageId="15" pageNumber="16">Beichatien Mountain (北插天山).</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="16" type="diagnosis">
<paragraph pageId="15" pageNumber="16">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="16">
This species is distinguished from other Taiwanese
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Amphimenes</taxonomicName>
by a combination of elytra with a black suture in apical half and a disc with randomly scattered punctures on entire dorsal surface of, +/- irregularly shaped in basal half.
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph pageId="15" pageNumber="16">Description.</paragraph>
<paragraph pageId="15" pageNumber="16">OBL 6.3 mm. Length: head 0.64, pronotum 1.20, elytra 3.50, metepisternum 0.84 mm; width: head 1.12, pronotum 1.56, elytra 2.5, metepisternum 0.52 mm.</paragraph>
<paragraph pageId="15" pageNumber="16">Body proportions. HW/HL 1.75; PWM/PL 1.30; EL/EW 1.40; ML/MW 1.62.</paragraph>
<paragraph pageId="15" pageNumber="16">Color. Fig. 6. Dorsum of head brunneo-piceous; clypeus and labrum rufo-brunneous; pronotum nearly piceous with margins rufo-brunneous; proepipleuron rufo-brunneous; antennae rufo-brunneous; palpi rufo-brunneous; elytral disc nearly piceous, lateral margins and sutural margin rufo-brunneous, translucent; elytral epipleura rufo-brunneous; thoracic sclerites brunneo-piceous; abdominal sterna brunneo-piceous medially and nearly piceous at lateral margins; legs with trochanter and femora rufo-testaceous, tibia rufo-brunneous to brunneo-piceous.</paragraph>
<caption pageId="15" pageNumber="16">
<paragraph pageId="15" pageNumber="16">
Figure 6. Dorsal habitus and color pattern of
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes beichatiensis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beichatiensis">Amphimenes beichatiensis</taxonomicName>
sp. n.. (OBL 6.30 mm).
</paragraph>
</caption>
<paragraph pageId="15" pageNumber="16">
Microsculpture. Dorsum of head and pronotum with isodiametric mesh pattern easily visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification; elytra with transverse sculpticells forming elongate mesh, faintly visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification.
</paragraph>
<paragraph pageId="15" pageNumber="16">Macrosculpture and pilosity. Elytra randomly scattered punctures on intervals, more or less evenly spaced throughout length, -/+ irregularly shaped in basal half; striae faintly punctate along length.</paragraph>
<paragraph pageId="15" pageNumber="16">
Pilosity. Elytra with scattered punctures visible, each bearing a small seta not visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
magnification.
</paragraph>
<paragraph pageId="15" pageNumber="16">Luster. Elytra moderately glossy; ventral thoracic sterna and abdominal sterna moderately glossy.</paragraph>
<paragraph pageId="15" pageNumber="16">Head. Fig. 8A. Labrum with indentation along apical margin, somewhat bilobed; eyes moderately convex.</paragraph>
<paragraph pageId="15" pageNumber="16">Pronotum. Lateral margins narrow; posterio-lateral margins obtuse, almost right-angled; basal lobe present.</paragraph>
<paragraph pageId="15" pageNumber="16">Elytra. Humeri broadly rounded; striae moderately impressed; lateral margins somewhat rounded; elytral apices truncate.</paragraph>
<paragraph pageId="15" pageNumber="16">Hind wings. Macropterous.</paragraph>
<paragraph pageId="15" pageNumber="16">Legs. Meso-tibia with several shallow notches from mid-way to base along ventral surface.</paragraph>
<paragraph pageId="15" pageNumber="16">
Male genitalia. Fig. 7
<normalizedToken originalValue="AC">A-C</normalizedToken>
. Length 1.60 mm. Phallus with several carina from base (bpc), longer to shorter from center to right of center when viewed ventrally; ventral carina from median of shaft almost to ostium margin; endophallus with small internal endophallic sclerite near apex.
</paragraph>
<caption pageId="15" pageNumber="16">
<paragraph pageId="15" pageNumber="16">
Figure 7. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes beichatiensis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beichatiensis">Amphimenes beichatiensis</taxonomicName>
sp. n.. A right lateral aspect B ventral aspect C left lateral aspect. Legend: bpc basal phallic carina.
</paragraph>
</caption>
<caption pageId="15" pageNumber="16">
<paragraph pageId="15" pageNumber="16">
Figure 8. Line drawing and comparison of menta of A
<taxonomicName class="Insecta" family="Carabidae" genus="Amphimenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphimenes beichatiensis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="beichatiensis">Amphimenes beichatiensis</taxonomicName>
sp. n. (with tooth) B
<taxonomicName class="Insecta" family="Carabidae" genus="Brachichila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Brachichila hypocrita" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hypocrita">Brachichila hypocrita</taxonomicName>
Chaudoir (without tooth).
</paragraph>
</caption>
<paragraph pageId="16" pageNumber="17">
<pageBreakToken pageId="16" pageNumber="17" start="start">Female</pageBreakToken>
genitalia. Female unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="habitat">
<paragraph pageId="16" pageNumber="17">Habitat, habits, and seasonal occurrence.</paragraph>
<paragraph pageId="16" pageNumber="17">The single specimen was collected from a live tree trunk in a montane mixed secondary forest. It was hand collected one night in June, at an elevation of 1320 meters.</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="distribution">
<paragraph pageId="16" pageNumber="17">Geographical distribution.</paragraph>
<paragraph pageId="16" pageNumber="17">Known only from type locality. See Figure 12. This locality was not sampled well or often so it is possible that many more specimens will be collected in the near future.</paragraph>
</subSubSection>
</treatment>
</document>