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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.85584" ID-GBIF-Dataset="3d0dc830-b1f9-4173-9ce5-752c63580560" ID-Pensoft-Pub="1860-1324-2-139" ID-Pensoft-UUID="03EFF5CBC2CF53CFA27D350B268657C3" ID-ZooBank="73B92B049B454237B81497A21DB59468" ModsDocID="1860-1324-69-2-139" checkinTime="1658439162963" checkinUser="pensoft" docAuthor="Kobor, Peter" docDate="2022" docId="090787AB85555BB09263F252D6FC9257" docLanguage="en" docName="DeutEntomolZeit 69(2): 139-150" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-07-21" docSource="http://dx.doi.org/10.3897/dez.69.85584" docTitle="Umbrageocoris Kobor 2019" docType="treatment" docVersion="4" id="03EFF5CBC2CF53CFA27D350B268657C3" lastPageNumber="139" masterDocId="03EFF5CBC2CF53CFA27D350B268657C3" masterDocTitle="A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)" masterLastPageNumber="150" masterPageNumber="139" pageNumber="139" updateTime="1732488684203" updateUser="ExternalLinkService">
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<mods:title id="61A03CBE573DA2CC14AD426BAC0A6BB6">A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)</mods:title>
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<mods:namePart id="9AA2B500BD8C8C1EB14E369A6FD2C869">Kobor, Peter</mods:namePart>
<mods:nameIdentifier id="59A637CB9B21EB3A1E36EBC4799DD013" type="ORCID">https://orcid.org/0000-0003-0266-7303</mods:nameIdentifier>
<mods:affiliation id="E692CB08D1587420FD8E81D6FDF99285">Plant Protection Institute, Centre for Agricultural Research, Eoetvoes Lorand Research Network, 15 Herman Otto street, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier id="534F07747EDDF81DD2B472F12A342F08" type="email">kobor.peter@atk.hu</mods:nameIdentifier>
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<mods:title id="D5A26B3C9AE136A76CCD1702E35818BF">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="A852A88D7E2AD0CB6E9593693B447FD4">2022</mods:date>
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<mods:number id="A595B939AC9B63A24C81D56AE3874CCF">69</mods:number>
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<mods:classification id="3AF04D7581C76728A67C62A8C5DC555E">journal article</mods:classification>
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<treatment id="090787AB85555BB09263F252D6FC9257" ID-GBIF-Taxon="197838462" LSID="urn:lsid:plazi:treatment:090787AB85555BB09263F252D6FC9257" httpUri="http://treatment.plazi.org/id/090787AB85555BB09263F252D6FC9257" lastPageNumber="139" pageId="0" pageNumber="139" scope_family="Geocoridae" scope_superFamily="Lygaeoidea">
<subSubSection id="7B437261EAEE8B18303C2B47880C50BE" pageId="0" pageNumber="139" type="nomenclature">
<paragraph id="29952BAD2357EAE5F1D5FDD4968B0C3E" pageId="0" pageNumber="139">
Genus
<taxonomicName id="86965C4234EC64060C9155E85C74DE8A" ID-CoL="8QNXC" LSID="090787AB-8555-5BB0-9263-F252D6FC9257" authority="Kobor, 2019" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
Umbrageocoris
<normalizedToken id="98D35A9FA823F5DABC1463B98330A30D" originalValue="Kóbor">Kobor</normalizedToken>
, 2019
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BAA0B4A21DDB636D25E1A0BA45AD646" pageId="0" pageNumber="139" type="type species">
<paragraph id="A65DA72317FFD49DD6FA1989197BEE75" pageId="0" pageNumber="139">Type species.</paragraph>
<paragraph id="B1B77698BA8DE0A0C6C9D8259EB7B054" pageId="0" pageNumber="139">
<taxonomicName id="EC20C0EE8EEF4588BC17246DFAB8A7B0" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris kondorosyi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="kondorosyi">
<emphasis id="8B2D22C184A0390E3EBF4D4FEFBE611F" italics="true" pageId="0" pageNumber="139">Umbrageocoris kondorosyi</emphasis>
</taxonomicName>
<normalizedToken id="432AB3ED8BB25009C645545DAFB9F721" originalValue="Kóbor">Kobor</normalizedToken>
, 2019, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="F3E2B085F13CC73B18982D612E1D458D" pageId="0" pageNumber="139" type="redescription">
<paragraph id="9FE0A8E405272EFBB84586EACE218AB5" pageId="0" pageNumber="139">Redescription.</paragraph>
<paragraph id="043E1C39EB0290209FE500D0F9314CFA" pageId="0" pageNumber="139">
General habitus ovoid, moderately elongate. Integument shiny, with sparse, silvery pubescence at least on abdominal venter. Head pentagonal; compound eyes large, reniform, slightly stylate; posterior edge of compound eyes touching anterior edge of pronotum. Ocular sulcus complete but shallow, slightly visible. Integument of vertex impunctate, with a thin longitudinal furrow of various length, at least present on clypeus. Antenniferous tubercles minute, not visible in dorsal view. Antennomere I shortest, graniform; antennomere II longest, cylindrical; antennomeres III and IV subequal in length; antennomere III cylindrical, antennomere IV fusiform. Clypeus with margins subparallel and apex rounded, surpassing the mandibular plates. Bucculae pointed, surpassing mandibular plates; ventral margins converging, forming a Y- or cup-shaped labial trough which continues in a suture of variable length towards base of head (Figs
<figureCitation id="CB69C743AB95CFA26EFFCC45478D7BBE" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">1</figureCitation>
,
<figureCitation id="86C9411D58EDC82C2E8E5F6FF7C5CA31" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">2</figureCitation>
). Labiomeres I-II conspicuously stouter than labiomeres III-IV; labiomere I not reaching anterior margin of propleurite, labiomere II shorter than labiomere III, apex of labiomere IV reaching or slightly surpassing metacoxae. Thorax. Pronotum trapeziform with anterior edges and margin broadly convex, lateral margin slightly constricted. Integument shiny, deeply punctate; punctation variably dense. Pronotal callosities and humeral angles at most moderately bulging. Scutellum elongate triangular, basal width less than length; variably punctate except at trifurcate carina. Trifurcate carina of scutellum complete or apically reduced. Apex of scutellum sharply pointed. Submacropterous, brachypterous and coleopterous wing morphs known. Submacropterous morph: hemelytron with margins of clavus converging apically, claval commissure reduced, indistinct; corium with lines of punctures along claval margin, Cu, and costal margin; punctation variably present between M-R and costal margin in the apical half of corium. M-R of corium branching in apical third. Exocorium mostly narrow, sometimes slightly flared in apical half. Membrane at most slightly surpassing apex of abdomen. Metathoracic wing with hamus partly reduced, level of reduction variable (Figs
<figureCitation id="18F509C6B1EBB215B9DDBBD29AB97C8E" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">3</figureCitation>
,
<figureCitation id="F2332CB638EC5148233A0E3B95DC3D28" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">4</figureCitation>
); intervannals present, basally fused. Thoracic pleurites and sternites with dense punctation except prosternal collar, supracoxal lobes and peritreme of metathoracic scent efferent apparatus. Prosternite with a narrow, but conspicuously bulging collar. Peritreme bulbous with dorsal flange protruding, sometimes indented; vestibular scar present, reaching metasternal venter; evaporatorium reduced to extreme surroundings of peritreme (Figs
<figureCitation id="E1ECE293E5F08D931F4688A679541C5A" captionStart="Figures 5–10" captionStartId="F2" captionText="Figures 5 - 10. Exoskeletal characters of Umbrageocoris: peritreme of metathoracic scent efferent apparatus of U. kondorosyi (5), U. maai maai (6), U. boonei sp. nov. (7), U. malipatili sp. nov. (8), U. elegantulus (9), and U. woodwardi (10) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures5-10" httpUri="https://binary.pensoft.net/fig/718125" pageId="0" pageNumber="139">5-10</figureCitation>
). Femora of prothoracic legs more incrassate than those of meso- and metathoracic legs. Legs covered with sparse, decumbent pubescence; fore femora with simple trichobothria in a single line, fore tibia with strong, dense setosity ventrally. Tarsomeres I and III of pro- and mesothoracic legs subequal in length, tarsomere II shortest. Tarsomere III of metathoracic leg conspicuously elongate, longer than combined lengths of tarsomeres I and II. Abdomen. Abdominal tergites III-V rugose medially; sutures of tergites 4/5 and 5/6 strongly curved medially; suture 4/5 elongate nearly reaching suture 5/6 (Fig.
<figureCitation id="31FD422A78FC614857E4100B87BA59E9" captionStart="Figures 15–18" captionStartId="F4" captionText="Figures 15 - 18. Sutures of abdominal tergites 4 / 5 and 5 / 6 in Umbrageocoris kondorosyi (15), Stylogeocoris elongatus (16), Germalus victoriae (17) and Geocoris (Geocoris) willeyi (18) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures15-18" httpUri="https://binary.pensoft.net/fig/718127" pageId="0" pageNumber="139">15</figureCitation>
). Abdominal sternites with rugose spots subdorsally. Abdominal trichobothria on sterintes III-IV with simple bothrium bearing a single sensilla, situated submedially, very close to each other in triangular arrangement (Fig.
<figureCitation id="BAF11E3695AAF912E60AF92EF59F3957" captionStart="Figures 11–14" captionStartId="F3" captionText="Figures 11 - 14. Exoskeletal characters of Umbrageocoris: male pygophore with parameres in situ (11), female ovipositor in situ (12), arrangement of abdominal trichobothria on sternite IV (13) and sternites V-VII (14)" figureDoi="10.3897/dez.69.85584.figures11-14" httpUri="https://binary.pensoft.net/fig/718126" pageId="0" pageNumber="139">11</figureCitation>
); trichobothria on sternites V-VII are subdorsal, either simple bothrium or trichome with microtrichia bearing a single sensilla, arrangement as in Fig.
<figureCitation id="A2A8A2A93D0F776F17F098BE5718EE30" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Exoskeletal characters of Umbrageocoris: 1 - 2. Labial trough (indicated by arrow) of U. kondorosyi (1) and U. malipatili sp. nov. (2), 3 - 4. Metathoracic wing (arrow indicates hamus) of U. kondorosyi (3) and U. elegantulus (4) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139">3B</figureCitation>
. Genitalia. Posterior opening of male pygophore with pointed, short lateral processes, situated dorsolaterally; paramere with trunk moderately stout and blade slender, evenly curved (Fig.
<figureCitation id="80E5216EEF720D3C1768DC6E69E8A072" captionStart="Figures 11–14" captionStartId="F3" captionText="Figures 11 - 14. Exoskeletal characters of Umbrageocoris: male pygophore with parameres in situ (11), female ovipositor in situ (12), arrangement of abdominal trichobothria on sternite IV (13) and sternites V-VII (14)" figureDoi="10.3897/dez.69.85584.figures11-14" httpUri="https://binary.pensoft.net/fig/718126" pageId="0" pageNumber="139">13</figureCitation>
); gonoporal process of aedeagus with 10 coils. Female ovipositor short, bisecting only abdominal sternite VII (Fig.
<figureCitation id="B7CAC3E9BB83002AFA5A7BCFEB21A473" captionStart="Figures 11–14" captionStartId="F3" captionText="Figures 11 - 14. Exoskeletal characters of Umbrageocoris: male pygophore with parameres in situ (11), female ovipositor in situ (12), arrangement of abdominal trichobothria on sternite IV (13) and sternites V-VII (14)" figureDoi="10.3897/dez.69.85584.figures11-14" httpUri="https://binary.pensoft.net/fig/718126" pageId="0" pageNumber="139">14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E84B839AC504C72745AC911A6F4344FB" pageId="0" pageNumber="139" type="comparisons">
<paragraph id="8D04E1BA59169BFEF110932E4431E96E" pageId="0" pageNumber="139">Comparative notes.</paragraph>
<paragraph id="F01352B07A9BC0BC8880BB773C8EF540" pageId="0" pageNumber="139">
<taxonomicName id="D8C48DD12B0513D25FD9872006B3B120" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="E2DA37ABED603EA91E6EDB75E4547912" italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
displays a remarkable similarity to the Australian genus
<taxonomicName id="734DF353D52593E1CCA064EDBEF03E39" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="8110A36C49CC7D006E8311933B32078F" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
in general facies [studied species:
<taxonomicName id="DB34FA2C22F82889783D5769133D5522" lsidName="S. biroi" pageId="0" pageNumber="139" rank="species" species="biroi">
<emphasis id="059E0ACFA98B9D1DEFA8B903D94DB8CA" italics="true" pageId="0" pageNumber="139">S. biroi</emphasis>
</taxonomicName>
Montandon, 1913 (lectotype, HNHM) and
<taxonomicName id="E07277258458973818599C50E3647B1F" lsidName="S. elongatus" pageId="0" pageNumber="139" rank="species" species="elongatus">
<emphasis id="8123040B32A9A7486C7627B750C1F3F3" italics="true" pageId="0" pageNumber="139">S. elongatus</emphasis>
</taxonomicName>
(Distant, 1901) (lectotype and paralectotype, BMNH; non-types, PCPK). However,
<taxonomicName id="AB3F99749A1A13183F5E85269A7A4178" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="EF5F9FC54EF241257ADC832C1F154ABA" italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
can be readily distinguished from the latter genus based on the combination of the following characters: integument of vertex smooth, dorsum of head with a single longitudinal furrow of various extent medially (it might be finely punctate and dorsum of head with arcuate grooves anteriad to ocelli in
<taxonomicName id="60FDBA6A6714A124FAB5FE78B0D8A4C9" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="D7611C3961716FED21C4A4C069553749" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); labial through Y- or cup-shaped (V-shaped in
<taxonomicName id="7190D69E8625F7B5FA1CD4FE8DE3130B" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="D0E50CF70F7D58F4EF1D24611C45163A" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); labiomere III longer than II (labiomeres II and III subequal in length in
<taxonomicName id="135E61654F59A9C03D8CEA062BA0391B" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="B82707658281991D54AC6DE1F94E5486" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); margins of clavus converging gradually towards apex, claval commissure absent (margins of clavus subparallel, claval commissure short but distinct in
<taxonomicName id="949F80D14497CB41616FFED752B143D8" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="8972977D78C0EBF16C1C0BCB9C509B64" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); hamus of hemelytron partly reduced (complete in
<taxonomicName id="D6E89E0FC52008E5609831AE203A13B5" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="25A5B9FC01E6C73A2DCC15ED56FA9951" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); peritreme rounded, evaporatorium reduced to surroundings of peritreme (peritreme auricular, evaporatorium covering most of the metapleurite in
<taxonomicName id="B2D7A680179CBC924910259ED581CB81" authorityName="Montandon" authorityYear="1913" class="Insecta" family="Geocoridae" genus="Stylogeocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Stylogeocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="9696F079948869693F01EF38601836BC" italics="true" pageId="0" pageNumber="139">Stylogeocoris</emphasis>
</taxonomicName>
); sutures of abdominal tergites 4/5 elongate almost reaching margin of 5/6, apex of suture 5/6 obtuse (sutures of abdominal tergites 4/5 and 5/6 not elongate, apex rounded as in Fig.
<figureCitation id="BCE4F5BF7432F079669D836962CFB8ED" captionStart="Figures 15–18" captionStartId="F4" captionText="Figures 15 - 18. Sutures of abdominal tergites 4 / 5 and 5 / 6 in Umbrageocoris kondorosyi (15), Stylogeocoris elongatus (16), Germalus victoriae (17) and Geocoris (Geocoris) willeyi (18) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures15-18" httpUri="https://binary.pensoft.net/fig/718127" pageId="0" pageNumber="139">15</figureCitation>
).
</paragraph>
<caption id="3AF6807D70CBEDF9AB32218B0E4F697D" doi="10.3897/dez.69.85584.figures1-4" httpUri="https://binary.pensoft.net/fig/718124" pageId="0" pageNumber="139" start="Figures 1–4" startId="F1">
<paragraph id="6E4915D1D11F5C177F9673803804C0C8" pageId="0" pageNumber="139">
<emphasis id="1B4DB643D3E80E4BE5AC6613F22808EA" bold="true" pageId="0" pageNumber="139">Figures 1-4.</emphasis>
Exoskeletal characters of
<taxonomicName id="A10CDDEE7C9D793C59775AE3E9BB9CAE" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="41E4E6935BB746D2E3F01B480910AF48" italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
:
<emphasis id="D4DBC407F386962676577E67B425645B" bold="true" pageId="0" pageNumber="139">1-2.</emphasis>
Labial trough (indicated by arrow) of
<taxonomicName id="F41A978568A991D61A5B8823496F3A56" class="Insecta" kingdom="Animalia" lsidName="U. kondorosyi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="kondorosyi">
<emphasis id="43D1E8DC244D6975EAE2CDA4705AB4CC" italics="true" pageId="0" pageNumber="139">U. kondorosyi</emphasis>
</taxonomicName>
(
<emphasis id="0CE907186EE1A820BB379A4956021ABB" bold="true" pageId="0" pageNumber="139">1</emphasis>
) and
<taxonomicName id="9767D75ED07CF57DE429592336B3D1CB" class="Insecta" kingdom="Animalia" lsidName="U. malipatili" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="malipatili">
<emphasis id="049FE61CDF03C996E3D15159F3613F26" italics="true" pageId="0" pageNumber="139">U. malipatili</emphasis>
</taxonomicName>
sp. nov. (
<emphasis id="E0D7817EC479E3ED747D7D116BEC59AC" bold="true" pageId="0" pageNumber="139">2</emphasis>
),
<emphasis id="C3C5029F636B508351299A4860F1D717" bold="true" pageId="0" pageNumber="139">3-4.</emphasis>
Metathoracic wing (arrow indicates hamus) of
<taxonomicName id="C9AAC35D44886A9FBBCC1F6F0712EEE0" class="Insecta" kingdom="Animalia" lsidName="U. kondorosyi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="kondorosyi">
<emphasis id="8570B19668B05016C69F0FBD1712CDB3" italics="true" pageId="0" pageNumber="139">U. kondorosyi</emphasis>
</taxonomicName>
(
<emphasis id="D64A7385B373E798BA8215083F225AE7" bold="true" pageId="0" pageNumber="139">3</emphasis>
) and
<taxonomicName id="1084C3186FDE5610AE1CC9FEBA3FF8E2" class="Insecta" kingdom="Animalia" lsidName="U. elegantulus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="FE93E3010858CD69154D1CECADF0E6E1" italics="true" pageId="0" pageNumber="139">U. elegantulus</emphasis>
</taxonomicName>
(
<emphasis id="B7E73F498207A6B92DB8BF5C8EBBA4B3" bold="true" pageId="0" pageNumber="139">4</emphasis>
) (images not to scale).
</paragraph>
</caption>
<paragraph id="3604BF9A8F2E3ECCF72BB6686FF40F78" pageId="0" pageNumber="139">
<taxonomicName id="D3287552C0D0114ABAED3958EDDB0EB6" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="4E7A99E0EACAC0DDCCE1004ACC6B0BDF" italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
can be distinguished from representatives of genus
<taxonomicName id="29EC5C65C2EE6E6039EAE4D65D8B573C" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="0C91976E3D3E6111D92A28EF8B4451E5" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
distributed in the region [studied species:
<taxonomicName id="1BC58EED354C299BEA1CCDDBED2B7A7C" lsidName="G. ochropterus" pageId="0" pageNumber="139" rank="species" species="ochropterus">
<emphasis id="B0DE00BFD9A736EEE06712BAE6E6397A" italics="true" pageId="0" pageNumber="139">G. ochropterus</emphasis>
</taxonomicName>
(Fieber, 1844) (lectotype, BMNH);
<taxonomicName id="2018A4961451442011A5697CE43123B4" lsidName="G. willeyi" pageId="0" pageNumber="139" rank="species" species="willeyi">
<emphasis id="C79114548E5D1624B1B1B3777E05F9AE" italics="true" pageId="0" pageNumber="139">G. willeyi</emphasis>
</taxonomicName>
Kirkaldy, 1905 (syntype, BMNH)] by the combination of following characters: head pentagonal, posterior edge of compound eyes touching anterior edges of pronotum (head lunulate, posterior edge of compound eyes encompassing anterior edges of pronotum in
<taxonomicName id="B301B382A2EE83595153684FE37AE6D7" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="139206D170DEC1169AEF817C1B308C06" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
); pronotum trapeziform with lateral margins slightly constricted medially (semicircular, lateral margins mostly not constricted in
<taxonomicName id="4CEAF1589B1E704944A23D0143CC0766" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="F7DE96DA465E724777386FAE710F1459" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
); trifurcate carina of scutellum distinct, at most apically reduced (slightly distinct to completely reduced in
<taxonomicName id="AD4DD8995A07215D36FCCB5C0F7C39FD" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="12816F85CA02B31008C2DA69C9E1D507" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
); hamus of metathoracic wing at most partly reduced, intervannals present, fused basally (hamus completely reduced and intervannals absent in
<taxonomicName id="8637D93A3E611ED08226C607490F623B" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="BB492A832E82091ED15BFDCCDFD29CFF" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
); apex of suture of abdominal tergites 5/6 obtuse (rounded in
<taxonomicName id="BBCD942C7E99168B9EA3ED1917D36699" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="1B9C7B38D4C44E23096F72A39C30BB7F" italics="true" pageId="0" pageNumber="139">Geocoris</emphasis>
</taxonomicName>
sp., as in Fig.
<figureCitation id="5235E1D4B1695E9EE0E7BDD8E41FA07D" captionStart="Figures 15–18" captionStartId="F4" captionText="Figures 15 - 18. Sutures of abdominal tergites 4 / 5 and 5 / 6 in Umbrageocoris kondorosyi (15), Stylogeocoris elongatus (16), Germalus victoriae (17) and Geocoris (Geocoris) willeyi (18) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures15-18" httpUri="https://binary.pensoft.net/fig/718127" pageId="0" pageNumber="139">18</figureCitation>
).
</paragraph>
<paragraph id="788586A8593F4ACF7B9AE0EA9C793CED" pageId="0" pageNumber="139">
<taxonomicName id="9B7231A8B96CEBFC17635555E786E6C9" authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="407722385F6CC182AD6F240D531EFF9B" italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
differs from representatives of
<taxonomicName id="D8C272FC0C25BF66B7AB286CE8ECBE4E" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="CEA699C5E40A2E5B6162DFBEFFDCD49F" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
[studied species:
<taxonomicName id="A3D3ADE04D7162037865D32CB56087E2" lsidName="G. coloratus" pageId="0" pageNumber="139" rank="species" species="coloratus">
<emphasis id="E8BF8F39F7389F031919B92B59930701" italics="true" pageId="0" pageNumber="139">G. coloratus</emphasis>
</taxonomicName>
Distant, 1918 (syntype, BMNH);
<taxonomicName id="79AA576742B89353E64FBE5F550C5156" lsidName="G. fuscovittatus" pageId="0" pageNumber="139" rank="species" species="fuscovittatus">
<emphasis id="048268B936D0BE8B927375399BA7C78E" italics="true" pageId="0" pageNumber="139">G. fuscovittatus</emphasis>
</taxonomicName>
Malipatil, 2013 (non-type, BMNH);
<taxonomicName id="237D631D53C874F1CAD93A7C16B4E454" lsidName="G. victoriae" pageId="0" pageNumber="139" rank="species" species="victoriae">
<emphasis id="8084BFB6924AD0C4605A9AD78C1E132E" italics="true" pageId="0" pageNumber="139">G. victoriae</emphasis>
</taxonomicName>
Bergroth, 1895 (lectotype, BMNH)] by the following characters: ocular sulcus complete, but slightly visible (always well-defined in
<taxonomicName id="B52D9859C1E3AC63FACEE27B99CC6D60" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="EAF0D017E5530CAD7CBE2C74200D602A" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); labial through Y- or cup-shaped (U- or V-shaped in
<taxonomicName id="9F146346139F6508EACB4CFFD82EE549" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="D2F4244D966FDD2DC0E79D5B5EF01471" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); labiomere III longer than II (labiomeres II and III subequal in length in
<taxonomicName id="80655EB543E88AF3149F4DBB44C44F4F" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="759AED5C05A2F50273AD8647BAA543FA" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); margins of clavus converging gradually towards apex, claval commissure absent (margins of clavus subparallel, claval commissure well-developed, distinct in
<taxonomicName id="B0ADDD82A15F4054F02AE7F609E669CA" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="EAD411EE502B3F1B9A7EE531CE39DEE6" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); hamus of hemelytron partly reduced (complete in
<taxonomicName id="BD5DA3D33BDA75FB0E31AF9D38FF17DF" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="643D6A309497F126C82E69A9DC9B8D7E" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); peritreme rounded, evaporatorium reduced to surroundings of peritreme (peritreme auricular with dorsal supportive process, evaporatorium covering metapleurite and posterior half of mesopleurite in
<taxonomicName id="2E416C2313CB7B1585CCF884AFBDEA12" authorityName="Stal" authorityYear="1862" class="Insecta" family="Geocoridae" genus="Germalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Germalus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis id="5BC63CCA549EB0780AED7C316328AC0C" italics="true" pageId="0" pageNumber="139">Germalus</emphasis>
</taxonomicName>
); sutures of abdominal tergites 4/5 and 5/6 elongated, suture 4/5 almost reaching margin of 5/6 (sutures of abdominal tergites 4/5 and 5/6 very short, strongly obtuse, as in Fig.
<figureCitation id="47B8E08EC4B732ED47A5E664197625FE" captionStart="Figures 15–18" captionStartId="F4" captionText="Figures 15 - 18. Sutures of abdominal tergites 4 / 5 and 5 / 6 in Umbrageocoris kondorosyi (15), Stylogeocoris elongatus (16), Germalus victoriae (17) and Geocoris (Geocoris) willeyi (18) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures15-18" httpUri="https://binary.pensoft.net/fig/718127" pageId="0" pageNumber="139">17</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="60AD1D0405F98FDFE3C3C708414842C7" pageId="0" pageNumber="139" type="distribution">
<paragraph id="EE43F44A55370FE44C241DF9BCE3C119" pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph id="98B5C869D3FEC8769CEE7B4C6D60F9EC" pageId="0" pageNumber="139">Representatives of the genus are distributed in continental Indomalaya and in the Indo-Australian Archipelago, from Laos to Northeast Australia (Northern Territories, Queensland, New South Wales).</paragraph>
</subSubSection>
</treatment>
</document>
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