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<document id="87699F2E8C4C906AD5A057C74713B547" ID-DOI="10.5281/zenodo.175670" ID-GBIF-Dataset="7f25f9fd-d335-42e5-b89a-6ea67e8ed713" ID-ISSN="1175-5326" ID-Zenodo-Dep="175670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459816256441" checkinUser="plazi" docAuthor="Wheeler, Terry A." docDate="2007" docId="E80487F0DE03385BFF7909929ECCC489" docLanguage="en" docName="zt01413p053.pdf" docOrigin="Zootaxa 1413" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Oscinicita Wheeler" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="143DFF88DE00385FFFEE0F419E0BC276" masterDocTitle="Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica" masterLastPageNumber="53" masterPageNumber="47" pageNumber="50" updateTime="1698221377221" updateUser="plazi">
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<mods:title id="5D99136FFD67FD996652C9DD3A15B169">Two new genera of oscinelline Chloropidae (Diptera) from Costa Rica</mods:title>
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<mods:namePart id="3398C7B7BACCFD7383636328407082D2">Wheeler, Terry A.</mods:namePart>
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<mods:date id="EBC6216843BF98E11C76192872889683">2007</mods:date>
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<treatment id="E80487F0DE03385BFF7909929ECCC489" ID-DOI="http://doi.org/10.5281/zenodo.6248560" ID-GBIF-Taxon="119354740" ID-Zenodo-Dep="6248560" LSID="urn:lsid:plazi:treatment:E80487F0DE03385BFF7909929ECCC489" httpUri="http://treatment.plazi.org/id/E80487F0DE03385BFF7909929ECCC489" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
<subSubSection id="28B7656DDE03385CFF7909929FEBC49B" box="[151,480,1747,1773]" pageId="3" pageNumber="50" type="nomenclature">
<paragraph id="601236E6DE03385CFF7909929FEBC49B" blockId="3.[151,480,1747,1773]" box="[151,480,1747,1773]" pageId="3" pageNumber="50">
<heading id="3B5A818ADE03385CFF7909929FEBC49B" bold="true" box="[151,480,1747,1773]" fontSize="11" level="1" pageId="3" pageNumber="50" reason="1">
<emphasis id="52D9EAF4DE03385CFF7909929FEBC49B" bold="true" box="[151,480,1747,1773]" pageId="3" pageNumber="50">
<taxonomicName id="A7AD4D65DE03385CFF7909929F8AC49B" ID-CoL="63GMX" authority="Wheeler" authorityName="Wheeler" box="[151,385,1747,1773]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE03385CFF7909929F1AC49B" bold="true" box="[151,273,1747,1773]" italics="true" pageId="3" pageNumber="50">Oscinicita</emphasis>
Wheeler
</taxonomicName>
, gen. n.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="28B7656DDE03385BFF7908579ECCC489" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="materials_examined">
<paragraph id="601236E6DE03385CFF7908579CF0C55D" blockId="3.[151,763,1814,1835]" box="[151,763,1814,1835]" pageId="3" pageNumber="50">
<typeStatus id="BF168844DE03385CFF7908579ECCC55D" box="[151,199,1814,1835]" pageId="3" pageNumber="50">Type</typeStatus>
species:
<taxonomicName id="A7AD4D65DE03385CFEF008579FD8C55D" box="[286,467,1814,1835]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="hansoni" status="sp. nov.">
<emphasis id="52D9EAF4DE03385CFEF008579FD8C55D" box="[286,467,1814,1835]" italics="true" pageId="3" pageNumber="50">Oscinicita hansoni</emphasis>
</taxonomicName>
<emphasis id="52D9EAF4DE03385CFE3708579C04C55D" bold="true" box="[473,527,1814,1835]" pageId="3" pageNumber="50">
<taxonomicNameLabel id="49EA578FDE03385CFE3708579C04C55D" box="[473,527,1814,1835]" pageId="3" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(by present designation)
</paragraph>
<paragraph id="601236E6DE03385CFF7908189C81C5B0" blockId="3.[151,1436,1881,2034]" pageId="3" pageNumber="50">
Diagnosis: Minute (
<quantity id="A7559B03DE03385CFE6608189C17C505" box="[392,540,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="50" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
)
<taxonomicName id="A7AD4D65DE03385CFDDE08189CC9C505" box="[560,706,1881,1907]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
with antennae in deep foveae, incurved vibrissal angle and gena, deep postgena, strongly convex scutum, and reduced wing venation (dm-cu absent, CuA1 straight and incomplete, other longitudinal veins short)
</paragraph>
<paragraph id="601236E6DE03385BFF2808999A50C01C" blockId="3.[151,1436,1881,2034]" lastBlockId="4.[151,1437,152,1791]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
Description:
<taxonomicName id="A7AD4D65DE03385CFEB008999FF9C584" box="[350,498,2008,2034]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
,
<taxonomicName id="A7AD4D65DE03385CFE1008999C9EC584" box="[510,661,2008,2034]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
. Head higher than long in lateral view, with frons strongly sloping toward lunule (
<figureCitation id="F8962A63DE04385BFEA70FD99F84C2C4" box="[329,399,152,178]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 8</figureCitation>
), wider than long in dorsal view, with occipital margin concave; frontal triangle distinct, bare except for row of setulae along margin, ocellar tubercle large relative to size of frons; frons with dense interfrontal setulae, fronto-orbital setae in long row barely distinguished from interfrontal setulae, vertical, ocellar and postocellar setae short and weak, ocellar setae reclinate, convergent, postocellar setae convergent; eye rounded, with short ommatrichia; gena incurved ventrally, sparsely setulose, with bare shining strip anteroventral to eye, continuous with shining parafacial; postgena deep and swollen posteroventral to eye, densely tomentose (
<figureCitation id="F8962A63DE04385BFE910EC99FCEC3D4" box="[383,453,392,418]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 8</figureCitation>
); vibrissal angle broadly rounded, strongly incurved toward midline of face (
<figureCitation id="F8962A63DE04385BFAA80EC99B86C3D4" box="[1350,1421,392,418]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 9</figureCitation>
), barely visible in lateral view, vibrissa short but distinct; face short, scarcely extending beyond ventral margin of first flagellomere, oral margin high and U-shaped (
<figureCitation id="F8962A63DE04385BFCE00E999D5EC384" box="[782,853,472,498]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 9</figureCitation>
), face narrow because of incurved vibrissal angle and parafacial, most of face forming deep antennal foveae, facial carina small and narrow in ventral part of antennal foveae; antenna small, only slightly projecting in lateral view, scape and pedicel small, first flagellomere small, rounded, arista short, pubescent; proboscis short; palpus short.
</paragraph>
<paragraph id="601236E6DE04385BFF280D399D0AC1A1" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Scutum strongly convex, postpronotum small; scutal setulae evenly distributed, 12 postpronotal, 1 anterior and 2 posterior notopleural, 1 postsutural supra-alar, 1 postsutural intra-alar, 1 dorsocentral seta apparently present (setae missing or damaged to some degree in all
<typeStatus id="BF168844DE04385BFC400D899DFAC094" box="[942,1009,712,738]" pageId="4" pageNumber="51">types</typeStatus>
); scutellum short, convex, densely tomentose, with short apical, subapical and dorsal setae; subscutellum swollen; thoracic pleurites bare. Wing hyaline, alula small, anal angle well-developed, R2+3 and R4+5 upcurved distally toward C, second costal sector longer than third; M1+2 ending near apex of wing, CuA1 straight, without trace of kink near midpoint, ending well before margin of wing, crossvein r-m present, crossvein dm-cu absent (
<figureCitation id="F8962A63DE04385BFBF30C2C9A7DC1F1" box="[1053,1142,877,903]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 10</figureCitation>
). Legs without outstanding setae or spurs; femoral organ not evident in
<typeStatus id="BF168844DE04385BFD0A0CD49D1CC1D9" box="[740,791,917,943]" pageId="4" pageNumber="51">type</typeStatus>
specimens; tibial organ may be present as indistinct, small oval patch (not obvious in all
<typeStatus id="BF168844DE04385BFDD20CFC9C64C1A1" box="[572,623,957,983]" pageId="4" pageNumber="51">type</typeStatus>
specimens).
</paragraph>
<paragraph id="601236E6DE04385BFF280CA49FE6C639" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Abdominal tergites with sparse setae laterally and posteriorly, evenly covered with tiny setulae; sternites relatively broad, with sparse setae, abdominal membrane densely spinulose; male spiracles
<quantity id="A7559B03DE04385BFB230B4C9B12C651" box="[1229,1305,1037,1063]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="4" pageNumber="51" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">35 in</quantity>
membrane near lateral margin of tergite.
</paragraph>
<paragraph id="601236E6DE04385BFF280B1F9B3DC71E" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Male postabdomen: dorsal pregenital sclerite narrow; spiracles 6 and
<quantity id="A7559B03DE04385BFBFA0B1F9A4FC60E" box="[1044,1092,1118,1144]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="51" unit="in" value="7.0">7 in</quantity>
membrane ventral to lateral margin of dorsal sclerite; epandrium simple, convex, with sparse setae; surstylus simple, broadly rounded, freely articulated with epandrium; hypandrium open posteriorly; pregonite fused with postgonite, weakly sclerotized; postgonite with ventral seta and campaniform sensillae; basiphallus elongate, sclerotized, distiphallus short, membranous (
<figureCitation id="F8962A63DE04385BFE100BBF9C9AC76E" box="[510,657,1278,1304]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Figs. 1112</figureCitation>
); phallapodeme simple, phallic guide sclerotized, rounded; cerci small, with triangular ventral points broadly separated across midline, setulose (
<figureCitation id="F8962A63DE04385BFBD30A679A9FC736" box="[1085,1172,1318,1344]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Fig. 12</figureCitation>
); subepandrial sclerite well-developed, with sclerotized projection extending dorsally above base of surstylus (
<figureCitation id="F8962A63DE04385BFB750A0F9B22C71E" box="[1179,1321,1358,1384]" captionStart="FIGURES 8 13" captionStartId="5.[151,269,1898,1922]" captionTargetBox="[190,1326,526,1861]" captionTargetId="figure@5.[190,1397,526,1938]" captionTargetPageId="5" captionText="FIGURES 8 13. Oscinicita hansoni. 8. head, lateral. 9. head, anterior. 10. wing. 11. male genitalia, lateral. 12. male genitalia, posterior. 13. male genitalia, ventral. Abbreviations: fov — antennal fovea; mem — membranous region ventral to facial margin; ses — subepandrial sclerite. Scale bars = 0.4 mm (Fig. 10), 0.1 mm (Figs. 11 13)." httpUri="https://zenodo.org/record/175672/files/figure.png" pageId="4" pageNumber="51">Figs. 1112</figureCitation>
).
</paragraph>
<paragraph id="601236E6DE04385BFF280A379F92C7E6" blockId="4.[151,1437,152,1791]" box="[198,409,1398,1424]" pageId="4" pageNumber="51">Female unknown.</paragraph>
<paragraph id="601236E6DE04385BFF280AE39F92C47A" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Etymology: The generic name is derived from
<emphasis id="52D9EAF4DE04385BFD070AE39D2CC7CD" box="[745,807,1442,1467]" italics="true" pageId="4" pageNumber="51">oscin</emphasis>
- a common generic root in the
<taxonomicName id="A7AD4D65DE04385BFB780AE39B2CC7CA" box="[1174,1319,1442,1468]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
and
<emphasis id="52D9EAF4DE04385BFAB10AE39B9EC7CD" box="[1375,1429,1442,1467]" italics="true" pageId="4" pageNumber="51">-cita</emphasis>
, a Spanish term widely used as a diminutive in
<collectingCountry id="18BA7676DE04385BFD5B0A8B9D3CC792" box="[693,823,1482,1508]" name="Costa Rica" pageId="4" pageNumber="51">Costa Rica</collectingCountry>
, referring to the minute size of these flies. The name is treated as feminine.
</paragraph>
<paragraph id="601236E6DE04385BFF28095C9ECCC489" blockId="4.[151,1437,152,1791]" pageId="4" pageNumber="51">
Remarks: Although the autapomorphic character states of
<taxonomicName id="A7AD4D65DE04385BFC9E095F9DE2C441" box="[880,1001,1566,1591]" class="Insecta" family="Chloropidae" genus="Oscinicita" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="52D9EAF4DE04385BFC9E095F9DE2C441" box="[880,1001,1566,1591]" italics="true" pageId="4" pageNumber="51">Oscinicita</emphasis>
</taxonomicName>
make these flies easily recognizable, they do not resolve the phylogenetic relationships of the genus. The head characters (genal structure, incurved vibrissal angle, short face with distinct antennal foveae) are not informative about relationships. The reduction in wing venation, especially loss of dm-cu, is a homoplasious state, related to small size, in
<taxonomicName id="A7AD4D65DE04385BFB0C09D49B7DC4D9" box="[1250,1398,1685,1711]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
, as well as other acalyptrates, although it occurs more frequently in the subfamily
<taxonomicName id="A7AD4D65DE04385BFBD909FC9AC3C4A1" box="[1079,1224,1725,1751]" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Chloropinae">Chloropinae</taxonomicName>
than in
<taxonomicName id="A7AD4D65DE04385BFAC709FC9EC8C489" class="Insecta" family="Chloropidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="51" phylum="Arthropoda" rank="subFamily" subFamily="Oscinellinae">Oscinellinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>