273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document id="3EEF3464A9DD2F73E7758FEA7CF8E767" ID-CLB-Dataset="46261" ID-DOI="10.11646/zootaxa.3774.5.3" ID-GBIF-Dataset="a73d4668-01cd-4e4c-8173-e130939a89c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="226487" ID-ZooBank="091256F3-B410-42FD-AFA2-3415BA48DDBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460394050722" checkinUser="plazi" docAuthor="Santos, Bianca Tamires Silva Dos, Nascimento, Agata Tyanne Silva & Fernandes, José Antônio Marin" docDate="2014" docId="5D0E87ABFFD9702DFF4D5E01695DFEDD" docLanguage="en" docName="zt03774p459.pdf" docOrigin="Zootaxa 3774 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessa godmani Distant 1881" docType="treatment" docVersion="9" lastPageNumber="446" masterDocId="A137FFD3FFDA7028FFDA5F736A78FF9D" masterDocTitle="Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)" masterLastPageNumber="459" masterPageNumber="441" pageNumber="444" updateTime="1698298553916" updateUser="ExternalLinkService">
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<mods:mods id="11ADB019554F601D0181FE27D33EA8BB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FD199B494F4F52E2CB0C9ED0D261A38C">
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<mods:title id="A19BFCEB79B6246994899E3882AE1EE3">Proposition of a new species group in Edessa Fabricius, 1803 (Hemiptera: Heteroptera: Pentatomidae: Edessinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2200B3A73EA3C957C04E87D5FCA1BF84" type="personal">
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<mods:role id="4F37CE401F85F386F52AD67CA2E1B2E2">
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<mods:roleTerm id="348CB5BDFB2CDD5B346BCFE6B521A00E">Author</mods:roleTerm>
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<mods:namePart id="337F9375607530939A0564AF023B4828">Santos, Bianca Tamires Silva Dos</mods:namePart>
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</mods:name>
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<mods:roleTerm id="F3F6349FB064D94C1D1D18E9D87D178D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B0032E070E2E712FF7EC6114BF05C8E5">Nascimento, Agata Tyanne Silva</mods:namePart>
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</mods:name>
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<mods:name id="132F8B94786FC960ECE0986A69DB8DAF" type="personal">
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<mods:role id="D12E330837C6065C9CD9B1CC1D2B5668">
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<mods:roleTerm id="D716E3112024D987B3A669844741B233">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="29C410B76A8C2390F166E233849EFFA5">Fernandes, José Antônio Marin</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5C12368198BDA002FED1B8341A1923EE">text</mods:typeOfResource>
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<mods:titleInfo id="128D4BFDFAB6A35043E9620247355C2A">
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<mods:title id="17CCF2166AC6BD3F4CF1C86CCCAB603F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1CC7343813BE24A59A8ABC4966AACDA2">
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<mods:date id="E991A67348E1D752D713D1A400C081B4">2014</mods:date>
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<mods:detail id="FDF6E3319823BAC308E2BB81A25FF620" type="volume">
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<mods:number id="48EF74806588E297E35A022B2770F291">3774</mods:number>
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</mods:detail>
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<mods:detail id="97BF9AACBCAC9E422FC7C493104553BD" type="issue">
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<mods:number id="3A36C6B2E0B103DCCB24F27509B3D9DB">5</mods:number>
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<mods:end id="90F1F22B19E7441D7D55D0005E306953">459</mods:end>
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<mods:classification id="C011326032F770355E2A59AA28522311">journal article</mods:classification>
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<mods:identifier id="418F056A818C5683F5B6EA515A82C55C" type="CLB-Dataset">46261</mods:identifier>
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<mods:identifier id="8DE34FD6AF1F62B27C5EC1B41574E7AD" type="DOI">10.11646/zootaxa.3774.5.3</mods:identifier>
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<treatment id="5D0E87ABFFD9702DFF4D5E01695DFEDD" ID-DOI="http://doi.org/10.5281/zenodo.6133719" ID-GBIF-Taxon="119417649" ID-Zenodo-Dep="6133719" LSID="urn:lsid:plazi:treatment:5D0E87ABFFD9702DFF4D5E01695DFEDD" httpUri="http://treatment.plazi.org/id/5D0E87ABFFD9702DFF4D5E01695DFEDD" lastPageId="5" lastPageNumber="446" pageId="3" pageNumber="444">
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<subSubSection id="9DBD6536FFD9702BFF4D5E016BF3FE33" pageId="3" pageNumber="444" type="nomenclature">
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<paragraph id="D51836BDFFD9702BFF4D5E016879FE11" blockId="3.[151,513,370,430]" box="[151,513,370,396]" pageId="3" pageNumber="444">
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<heading id="8E5081D1FFD9702BFF4D5E016879FE11" bold="true" box="[151,513,370,396]" fontSize="11" level="1" pageId="3" pageNumber="444" reason="1">
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<taxonomicName id="12A74D3EFFD9702BFF4D5E016879FE11" authority="Distant 1881" authorityName="Distant" authorityYear="1881" box="[151,513,370,396]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="godmani">
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||
<emphasis id="E7D3EAAFFFD9702BFF4D5E016879FE11" bold="true" box="[151,513,370,396]" pageId="3" pageNumber="444">
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||
<emphasis id="E7D3EAAFFFD9702BFF4D5E016B25FE11" bold="true" box="[151,349,370,396]" italics="true" pageId="3" pageNumber="444">Edessa godmani</emphasis>
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||
Distant 1881
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="D51836BDFFD9702BFF4D5EE66BF3FE33" blockId="3.[151,513,370,430]" box="[151,395,405,430]" pageId="3" pageNumber="444">
|
||
(
|
||
<figureCitation id="4D9C2A38FFD9702BFF455EE66B7DFE33" box="[159,261,405,430]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Figs.1–6</figureCitation>
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||
; 42; 49; 56)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9DBD6536FFD9702BFF4D5EAE6978FDC1" pageId="3" pageNumber="444" type="reference_group">
|
||
<paragraph id="D51836BDFFD9702BFF4D5EAE6E59FE6E" blockId="3.[151,1057,477,499]" box="[151,1057,477,499]" pageId="3" pageNumber="444">
|
||
<treatmentCitationGroup id="F5B71193FFD9702BFF4D5EAE6E59FE6E" box="[151,1057,477,499]" pageId="3" pageNumber="444">
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||
<taxonomicName id="12A74D3EFFD9702BFF4D5EAE6B97FE6E" authority="Distant 1881: 96" authorityName="Distant" authorityPageNumber="96" authorityYear="1881" box="[151,495,477,499]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="godmani">
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<emphasis id="E7D3EAAFFFD9702BFF4D5EAE6B46FE6E" box="[151,318,477,499]" italics="true" pageId="3" pageNumber="444">Edessa godmani</emphasis>
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Distant 1881: 96
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</taxonomicName>
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;
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<treatmentCitation id="540610ACFFD9702BFE235EAE693DFE6E" author="Lethierry" box="[505,837,477,499]" page="190" pageId="3" pageNumber="444" year="1893">
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<bibRefCitation id="B1364B4CFFD9702BFE235EAE696AFE6E" author="Lethierry" box="[505,786,477,499]" pageId="3" pageNumber="444" refString="Lethierry, L. & Severin, G. (1893) Catalogue General des Hemipteres. Tome I. Heteropteres Pentatomidae. F. Hayez Brussels, X + 286 pp." type="book" year="1893">Lethierry and Severin 1893</bibRefCitation>
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||
: 190
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="540610ACFFD9702BFC8A5EAE6E65FE6E" author="Kirkaldy" box="[848,1053,477,499]" page="158" pageId="3" pageNumber="444" year="1909">
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||
<bibRefCitation id="B1364B4CFFD9702BFC8A5EAE6993FE6E" author="Kirkaldy" box="[848,1003,477,499]" pageId="3" pageNumber="444" refString="Kirkaldy, G. W. (1909) Catalogue of Hemiptera (Heteroptera). Vol. 1. Cimicidae. Felix L. Dames Berlin, XL + 392 pp." type="book" year="1909">Kirkaldy, 1909</bibRefCitation>
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||
: 158
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="D51836BDFFD9702BFF4D5D536901FDA5" blockId="3.[151,1437,543,2009]" box="[151,889,543,568]" pageId="3" pageNumber="444">
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<typeStatus id="0A1C881FFFD9702BFF4D5D536B7FFDA5" box="[151,263,544,568]" pageId="3" pageNumber="444" type="lectotype">Lectotype</typeStatus>
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male:
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<collectingCountry id="ADB0762DFFD9702BFE895D6C6BB7FDA5" box="[339,463,543,568]" name="Guatemala" pageId="3" pageNumber="444">Guatemala</collectingCountry>
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, Capetillo (BMNH). Here designated.
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||
</paragraph>
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||
<paragraph id="D51836BDFFD9702BFF1D5D376978FDC1" blockId="3.[151,1437,543,2009]" box="[199,768,580,605]" pageId="3" pageNumber="444">
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||
<typeStatus id="0A1C881FFFD9702BFF1D5D376B27FDC0" box="[199,351,580,605]" pageId="3" pageNumber="444" type="paralectotype">Paralectotype</typeStatus>
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female with the same data (BMNH).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="9DBD6536FFD9702BFF1D5D146BF9FC3D" pageId="3" pageNumber="444" type="materials_examined">
|
||
<paragraph id="D51836BDFFD9702BFF1D5D146BF9FC3D" blockId="3.[151,1437,543,2009]" pageId="3" pageNumber="444">
|
||
<emphasis id="E7D3EAAFFFD9702BFF1D5D14683BFD1D" bold="true" box="[199,579,615,640]" pageId="3" pageNumber="444">
|
||
Material.
|
||
<collectingCountry id="ADB0762DFFD9702BFE925D146BB8FD1D" box="[328,448,615,640]" name="Mexico" pageId="3" pageNumber="444">MEXICO</collectingCountry>
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||
, Morelos:
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||
</emphasis>
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||
♂ Cuernavaca,
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||
<date id="A119107DFFD9702BFCC15D1B6904FD1D" box="[795,892,616,640]" pageId="3" pageNumber="444" value="1900-09">IX-1900</date>
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||
, van Duzee coll., Comp. W.
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||
<typeStatus id="0A1C881FFFD9702BFAD85D1B6F42FD1D" box="[1282,1338,616,640]" pageId="3" pageNumber="444">Type</typeStatus>
|
||
(
|
||
<collectionCode id="B3B6AE78FFD9702BFA8E5D1B6FE8FD1D" LSID="urn:lsid:biocol.org:col:34700" box="[1364,1424,616,640]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3" pageNumber="444">CAS</collectionCode>
|
||
).
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||
<emphasis id="E7D3EAAFFFD9702BFF4D5DFF6BA3FD38" bold="true" box="[151,475,652,677]" pageId="3" pageNumber="444">
|
||
<collectingCountry id="ADB0762DFFD9702BFF4D5DFF6B28FD38" box="[151,336,652,677]" name="Guatemala" pageId="3" pageNumber="444">GUATEMALA</collectingCountry>
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||
, Zacapa: ♀
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||
</emphasis>
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||
<quantity id="125F9B58FFD9702BFE385DFF684FFD38" box="[482,567,652,677]" metricMagnitude="3" metricUnit="m" metricValue="1.6764" pageId="3" pageNumber="444" unit="ft" value="5500.0">5500 ft</quantity>
|
||
.,
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||
<date id="A119107DFFD9702BFD925DFF68A0FD39" box="[584,728,652,676]" pageId="3" pageNumber="444" value="1948-07-31">31-VII-1948</date>
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||
, Exp. Zool. 1948
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||
<collectionCode id="B3B6AE78FFD9702BFC785DFF6981FD39" LSID="urn:lsid:biocol.org:col:33139" box="[930,1017,652,676]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="3" pageNumber="444">CNHM</collectionCode>
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||
, R. D. Mitchell leg., Santa Clara, int. Valle of Sierra de las Minas, N. of Cabañas in cornfield (
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||
<collectionCode id="B3B6AE78FFD9702BFCCC5DC36909FD55" LSID="urn:lsid:biocol.org:col:33139" box="[790,881,688,712]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="3" pageNumber="444">CNHM</collectionCode>
|
||
).
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||
<emphasis id="E7D3EAAFFFD9702BFC595DDC6984FD55" bold="true" box="[899,1020,687,712]" pageId="3" pageNumber="444">Escuintla:</emphasis>
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||
♀ Finca El Zapote, Zapote,
|
||
<quantity id="125F9B58FFD9702BFAE75DDC6FEBFD55" box="[1341,1427,687,712]" metricMagnitude="3" metricUnit="m" metricValue="1.6764" pageId="3" pageNumber="444" unit="ft" value="5500.0">5500 ft</quantity>
|
||
.,
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||
<date id="A119107DFFD9702BFF4D5DA76B48FD71" box="[151,304,724,748]" pageId="3" pageNumber="444" value="1948-08-10">10-VIII-1948</date>
|
||
, Exp. Zool. 1948
|
||
<collectionCode id="B3B6AE78FFD9702BFDDB5DA76820FD71" LSID="urn:lsid:biocol.org:col:33139" box="[513,600,724,748]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="3" pageNumber="444">CNHM</collectionCode>
|
||
, R. D. Mitchell leg., Santa Clara, int. Valle of Sierra de las Minas, N. of Cabañas (
|
||
<collectionCode id="B3B6AE78FFD9702BFED35D8B6B22FC8D" box="[265,346,760,784]" httpUri="http://grbio.org/cool/nen5-04em" name="Instituto de Biologia" pageId="3" pageNumber="444">UFRG</collectionCode>
|
||
).
|
||
<emphasis id="E7D3EAAFFFD9702BFEAB5D846864FC8D" bold="true" box="[369,540,759,784]" pageId="3" pageNumber="444">Sacatepequez:</emphasis>
|
||
♀ Finca San Rafael,
|
||
<quantity id="125F9B58FFD9702BFCFE5D846906FC8D" box="[804,894,759,784]" metricMagnitude="3" metricUnit="m" metricValue="2.1031199999999997" pageId="3" pageNumber="444" unit="ft" value="6900.0">6900 ft</quantity>
|
||
.,
|
||
<date id="A119107DFFD9702BFC495D8B6E61FC8D" box="[915,1049,760,784]" pageId="3" pageNumber="444" value="1948-06-30">30-VI-1948</date>
|
||
, Exp. Zool. 1948
|
||
<collectionCode id="B3B6AE78FFD9702BFB2F5D8B6F34FC8D" LSID="urn:lsid:biocol.org:col:33139" box="[1269,1356,760,784]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="3" pageNumber="444">CNHM</collectionCode>
|
||
, R. D. Mitchell leg., sweeping herbs in second growth Oaks (
|
||
<collectionCode id="B3B6AE78FFD9702BFD2E5C6F6937FCA9" LSID="urn:lsid:biocol.org:col:33139" box="[756,847,796,820]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33139" name="Cincinnati Museum of Natural History" pageId="3" pageNumber="444">CNHM</collectionCode>
|
||
).
|
||
<emphasis id="E7D3EAAFFFD9702BFCBA5C6F69C5FCA8" bold="true" box="[864,957,796,821]" pageId="3" pageNumber="444">Patzun:</emphasis>
|
||
♂
|
||
<quantity id="125F9B58FFD9702BFC055C6F6E69FCA8" box="[991,1041,796,821]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="3" pageNumber="444" unit="mi" value="4.0">4 mi</quantity>
|
||
SE Patzun, 7100’,
|
||
<date id="A119107DFFD9702BFB3D5C6F6F00FCA9" box="[1255,1400,796,820]" pageId="3" pageNumber="444" value="1974-07-27">27-VII-1974</date>
|
||
, C. and L. O’Brien and
|
||
<collectingCountry id="ADB0762DFFD9702BFE5C5C4C6B92FCC5" box="[390,490,831,856]" name="Marshall Islands" pageId="3" pageNumber="444">Marshall</collectingCountry>
|
||
(
|
||
<collectionCode id="B3B6AE78FFD9702BFE265C33682BFCC5" LSID="urn:lsid:biocol.org:col:34868" box="[508,595,832,856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="3" pageNumber="444">UNSM</collectionCode>
|
||
).
|
||
<emphasis id="E7D3EAAFFFD9702BFDB35C4C69C0FCC5" bold="true" box="[617,952,831,856]" pageId="3" pageNumber="444">
|
||
<collectingCountry id="ADB0762DFFD9702BFDB35C4C694AFCC5" box="[617,818,831,856]" name="El Salvador" pageId="3" pageNumber="444">EL SALVADOR</collectingCountry>
|
||
, Usulután:
|
||
</emphasis>
|
||
♀ Cerro verde, 6800’,
|
||
<date id="A119107DFFD9702BFB0A5C4C6F2EFCC5" box="[1232,1366,831,856]" pageId="3" pageNumber="444" value="1963-06-29">29-VI-1963</date>
|
||
, D.Q. Cavagnaro and M. E. Irwin Coll. (
|
||
<collectionCode id="B3B6AE78FFD9702BFDE35C17680DFCE1" LSID="urn:lsid:biocol.org:col:34700" box="[569,629,868,892]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="3" pageNumber="444">CAS</collectionCode>
|
||
).
|
||
<emphasis id="E7D3EAAFFFD9702BFD575C1769A2FCE0" bold="true" box="[653,986,868,893]" pageId="3" pageNumber="444">
|
||
<collectingCountry id="ADB0762DFFD9702BFD575C176945FCE0" box="[653,829,868,893]" name="Nicaragua" pageId="3" pageNumber="444">NICARAGUA</collectingCountry>
|
||
, Matagalpa:
|
||
</emphasis>
|
||
♂
|
||
<quantity id="125F9B58FFD9702BFBD25C176E2DFCE0" box="[1032,1109,868,893]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="444" unit="km" value="20.0">20 km</quantity>
|
||
N. Matagalpa,
|
||
<date id="A119107DFFD9702BFAC95C176F02FCE1" box="[1299,1402,868,892]" pageId="3" pageNumber="444" value="1989-07">VII-1989</date>
|
||
F. Reinboldt (
|
||
<collectionCode id="B3B6AE78FFD9702BFECE5CFB6B0DFC3D" box="[276,373,904,928]" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="3" pageNumber="444">UFRGS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DBD6536FFD9702BFF1D5CDF6FEAF925" pageId="3" pageNumber="444" type="description">
|
||
<paragraph id="D51836BDFFD9702BFF1D5CDF68D8FC75" blockId="3.[151,1437,543,2009]" pageId="3" pageNumber="444">
|
||
<emphasis id="E7D3EAAFFFD9702BFF1D5CDF6B05FC58" bold="true" box="[199,381,940,965]" pageId="3" pageNumber="444">Measurements.</emphasis>
|
||
Head length (1.2–1.7); head width (3.5–3.7); pronotal length (2–3); pronotal width (6–7.5) total length (10–11); abdominal width (6.5–7).
|
||
</paragraph>
|
||
<paragraph id="D51836BDFFD9702BFF1D5C876F48FAE9" blockId="3.[151,1437,543,2009]" pageId="3" pageNumber="444">
|
||
Body greenish-brown dorsally (
|
||
<figureCitation id="4D9C2A38FFD9702BFDF45C8768FBFB91" box="[558,643,1012,1037]" captionStart="FIGURES 42 – 48" captionStartId="16.[151,264,1877,1899]" captionTargetBox="[253,1358,215,1847]" captionTargetId="figure@16.[214,1372,193,1856]" captionTargetPageId="16" captionText="FIGURES 42 – 48. Photo of the species, dorsal view. 42 - Edessa godmani Distant, 1881; 43 — Edessa punctata sp. nov.; 44 — Edessa picticornis Stål, 1872; 45 — Edessa luteomaculata sp. nov.; 46 — Edessa pudibunda Stål, 1862; 47 — Edessa nigriventris sp. nov.; 48 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226495/files/figure.png" pageId="3" pageNumber="444">Fig. 42</figureCitation>
|
||
), densely punctured, punctures black surrounded by suffused spots on scutellum and corium; some parts with dark spots instead of punctures. Ventral surface dark brown (
|
||
<figureCitation id="4D9C2A38FFD9702BFAED5B646FE8FBAD" box="[1335,1424,1047,1072]" captionStart="FIGURES 49 – 55" captionStartId="17.[151,264,1877,1899]" captionTargetBox="[262,1330,230,1838]" captionTargetId="figure@17.[236,1351,193,1856]" captionTargetPageId="17" captionText="FIGURES 49 – 55. Photo of species, ventral view. 49 - Edessa godmani; 50 — Edessa punctata sp. nov.; 51 — Edessa picticornis; 52 — Edessa luteomaculata sp. nov.; 53 — Edessa pudibunda; 54 — Edessa nigriventris sp. nov.; 55 — Edessa nigridorsata sp. nov." httpUri="https://zenodo.org/record/226496/files/figure.png" pageId="3" pageNumber="444">Fig. 49</figureCitation>
|
||
). Mandibular plates and tylus at the same level, rugulose. Ventral surface of head brown with a tiny dark spot below antenniferous tubercle; not punctured. First three antennal segments brown and densely spotted; basal third of fourth and fifth segments whitish with tiny dark spots densely distributed. Pronotum uniformly greenish-brown, with a median pale line not punctured; anterolateral margin reddish. Apex of scutellum with a yellow spot. Corium with a dark oblique spot. Metasternal process and evaporatorium spotted. Ventral punctures of thorax black. Metasternal process uniformly brown; arms of bifurcation of the metasternal process strong and subparallel. Legs densely dark spotted. Connexivum greenish-brown with dark punctures and a medial small calloused area yellow. Connexivum of segment VII distally dark due to high concentration of dark punctures. Middle ventral surface of abdomen sometimes with sparse punctures. Female without a rounded pair of dark spots on segment VII.
|
||
</paragraph>
|
||
<paragraph id="D51836BDFFD9702BFF1D5A0C6E46F9B5" blockId="3.[151,1437,543,2009]" pageId="3" pageNumber="444">
|
||
Male: Pygophore mostly brown (
|
||
<figureCitation id="4D9C2A38FFD9702BFD9A5A0C68D6FA05" box="[576,686,1407,1432]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Figs. 1–3</figureCitation>
|
||
). Dorsal rim with fine and concolorous punctures concentrated in the middle region. Superior processes of the genital cup pedunculate, distal part oval and concave, swollen margins not reaching dorsal rim (
|
||
<figureCitation id="4D9C2A38FFD9702BFE735AB46B96FA7D" box="[425,494,1479,1504]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Fig. 2</figureCitation>
|
||
). Paramere with margins dark, distal part elliptical; ventral process laterally directed and longer than distal one (
|
||
<figureCitation id="4D9C2A38FFD9702BFE125A9F684CF999" box="[456,564,1516,1541]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Figs. 1–2</figureCitation>
|
||
). Proctiger with two keels; posterior margin rounded and impunctate (
|
||
<figureCitation id="4D9C2A38FFD9702BFA935A9F6FF6F999" box="[1353,1422,1516,1541]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Fig. 2</figureCitation>
|
||
). Ventral rim with dark brown punctures; rounded expansion not swollen (
|
||
<figureCitation id="4D9C2A38FFD9702BFC1F597C6E49F9B5" box="[965,1073,1551,1576]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Figs. 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D51836BDFFD9702BFF1D59476FEAF925" blockId="3.[151,1437,543,2009]" pageId="3" pageNumber="444">
|
||
Female: Gonocoxites 8 strongly convex with dark punctures, excavation of the mesial margins deep, U-shaped (
|
||
<figureCitation id="4D9C2A38FFD9702BFF4559246B71F9ED" box="[159,265,1623,1648]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Fig. 5–6</figureCitation>
|
||
), narrower than gonocoxite 9, rounded distally and leaving visible the smooth gonapophyses 8. Gonocoxite 9 constricted just below punctured part, basal part bent upwards forming a small conical projection. Laterotergites 9 punctured and not grooved, acuminate apices surpassing the band uniting laterotergites 8 (
|
||
<figureCitation id="4D9C2A38FFD9702BFA9A59EC6FFCF925" box="[1344,1412,1695,1720]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1877,1899]" captionTargetBox="[214,1374,218,1830]" captionTargetId="figure@4.[199,1387,193,1856]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Edessa godmani. 1 – 3. Male, pygophore; 1 — dorsal; 2 — posterior; 3 — ventral. 4 — Metasternal process. 5 – 6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc 8 — gonocoxites 8; gc 9 — gonocoxite 9; gp—genital cup process; gp 8 — gonapophyses 8; la 8, laterotergites 8; la 9 — laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm." httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="3" pageNumber="444">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DBD6536FFD9702DFF1D59B76B12FF65" lastPageId="5" lastPageNumber="446" pageId="3" pageNumber="444" type="discussion">
|
||
<paragraph id="D51836BDFFD9702DFF1D59B76B12FF65" blockId="3.[151,1437,543,2009]" lastBlockId="5.[151,1436,151,320]" lastPageId="5" lastPageNumber="446" pageId="3" pageNumber="444">
|
||
<emphasis id="E7D3EAAFFFD9702BFF1D59B76B37F940" bold="true" box="[199,335,1732,1757]" pageId="3" pageNumber="444">Comments.</emphasis>
|
||
The syntypic series is formed by a pair–both were examined and we here designate the male specimen as
|
||
<typeStatus id="0A1C881FFFD9702BFEF759946BEBF89D" box="[301,403,1767,1792]" pageId="3" pageNumber="444" type="lectotype">lectotype</typeStatus>
|
||
of the species. Distant (1881) considered this species close to
|
||
<taxonomicName id="12A74D3EFFD9702BFBAD599A6F6AF89D" box="[1143,1298,1768,1792]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="pudibunda">
|
||
<emphasis id="E7D3EAAFFFD9702BFBAD599A6F6AF89D" box="[1143,1298,1768,1792]" italics="true" pageId="3" pageNumber="444">E. pudibunda</emphasis>
|
||
</taxonomicName>
|
||
, the former having a more linear and less sinuous body, second antennal segment shorter than third, and punctures much closer and coarser. According to measurements presented here Distant inverted the relationship between second and third antennal segments of the species.
|
||
<taxonomicName id="12A74D3EFFD9702BFDEC5827688EF8F1" box="[566,758,1876,1900]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="godmani">
|
||
<emphasis id="E7D3EAAFFFD9702BFDEC5827688EF8F1" box="[566,758,1876,1900]" italics="true" pageId="3" pageNumber="444">Edessa godmani</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="12A74D3EFFD9702BFC4D58266E58F8F1" box="[919,1056,1877,1900]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="punctata">
|
||
<emphasis id="E7D3EAAFFFD9702BFC4D58266E58F8F1" box="[919,1056,1877,1900]" italics="true" pageId="3" pageNumber="444">E. punctata</emphasis>
|
||
</taxonomicName>
|
||
the body densely and coarsely punctured, metasternal process punctured, punctures and color pattern of the connexivum and a double keel on the proctiger.
|
||
<taxonomicName id="12A74D3EFFD9702BFED158EF6BBDF829" box="[267,453,1948,1972]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="godmani">
|
||
<emphasis id="E7D3EAAFFFD9702BFED158EF6BBDF829" box="[267,453,1948,1972]" italics="true" pageId="3" pageNumber="444">Edessa godmani</emphasis>
|
||
</taxonomicName>
|
||
can be separate from
|
||
<taxonomicName id="12A74D3EFFD9702BFD1258EE6933F829" box="[712,843,1949,1972]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="punctata">
|
||
<emphasis id="E7D3EAAFFFD9702BFD1258EE6933F829" box="[712,843,1949,1972]" italics="true" pageId="3" pageNumber="444">E. punctata</emphasis>
|
||
</taxonomicName>
|
||
by fourth antennal segment with a light basal ring, smaller size of the superior process of genital cup, shape of paramere and U-shaped excavation of medial margins of gonocoxites 8. The specimen of CNHM coming from
|
||
<collectingCountry id="ADB0762DFFDF702DFCC05FE469EBFF2D" box="[794,915,151,176]" name="Guatemala" pageId="5" pageNumber="446">Guatemala</collectingCountry>
|
||
has a label stating that it was collected in low vegetation along the secondary forest. That is the only information on habitat possibly used by all species of
|
||
<taxonomicName id="12A74D3EFFDF702DFF4D5FAC6B62FF65" box="[151,282,223,248]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="446" phylum="Arthropoda" rank="species" species="pudibunda">
|
||
<emphasis id="E7D3EAAFFFDF702DFF4D5FAC6B62FF65" bold="true" box="[151,282,223,248]" pageId="5" pageNumber="446">pudibunda</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="81D86635FFDE702CFF4D5826695AF85A" httpUri="https://zenodo.org/record/226488/files/figure.png" pageId="4" pageNumber="445" targetBox="[214,1374,218,1830]" targetPageId="4">
|
||
<paragraph id="D51836BDFFDE702CFF4D5826695AF85A" blockId="4.[151,1435,1877,1991]" pageId="4" pageNumber="445">
|
||
<emphasis id="E7D3EAAFFFDE702CFF4D58266B44F8F6" bold="true" box="[151,316,1877,1899]" pageId="4" pageNumber="445">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="12A74D3EFFDE702CFE9F58266B96F8F6" box="[325,494,1877,1899]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="godmani">
|
||
<emphasis id="E7D3EAAFFFDE702CFE9F58266B96F8F6" box="[325,494,1877,1899]" italics="true" pageId="4" pageNumber="445">Edessa godmani</emphasis>
|
||
</taxonomicName>
|
||
. 1–3. Male, pygophore; 1—dorsal; 2—posterior; 3—ventral. 4—Metasternal process. 5–6, female, genital plates, ventral and posterior respectively. dr—dorsal rim; gc8—gonocoxites 8; gc9—gonocoxite 9; gp—genital cup process; gp8—gonapophyses 8; la8, laterotergites 8; la9—laterotergites 9; pa—paramere; pla—posterolateral angle; vrventral rim; VII—urotergite VII; X—segment X. Scale = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9DBD6536FFDF702DFF1D5E77695DFEDD" pageId="5" pageNumber="446" type="distribution">
|
||
<paragraph id="D51836BDFFDF702DFF1D5E77695DFEDD" blockId="5.[151,1436,151,320]" pageId="5" pageNumber="446">
|
||
<emphasis id="E7D3EAAFFFDF702DFF1D5E776B21FE80" bold="true" box="[199,345,260,285]" pageId="5" pageNumber="446">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="4D9C2A38FFDF702DFEB65E776BCEFE80" box="[364,438,260,285]" captionStart="FIGURE 56" captionStartId="18.[151,250,1159,1181]" captionTargetBox="[168,1417,206,1121]" captionTargetId="figure@18.[151,1436,171,1138]" captionTargetPageId="18" captionText="FIGURE 56. Distribution of the species of the pudibunda group." httpUri="https://zenodo.org/record/226497/files/figure.png" pageId="5" pageNumber="446">Fig.56</figureCitation>
|
||
).
|
||
<collectingCountry id="ADB0762DFFDF702DFE0A5E776852FE80" box="[464,554,260,285]" name="Mexico" pageId="5" pageNumber="446">Mexico</collectingCountry>
|
||
: Morelos.
|
||
<collectingCountry id="ADB0762DFFDF702DFD7D5E77695DFE80" box="[679,805,260,285]" name="Guatemala" pageId="5" pageNumber="446">Guatemala</collectingCountry>
|
||
: Zacapa, Escuintla, Sacapequetez, Patzun.
|
||
<collectingCountry id="ADB0762DFFDF702DFACC5E776FE1FE80" box="[1302,1433,260,285]" name="Costa Rica" pageId="5" pageNumber="446">Costa Rica</collectingCountry>
|
||
: Puntarenas.
|
||
<collectingCountry id="ADB0762DFFDF702DFEFA5E546BE0FEDD" box="[288,408,295,320]" name="Nicaragua" pageId="5" pageNumber="446">Nicaragua</collectingCountry>
|
||
: Matagalpa.
|
||
<collectingCountry id="ADB0762DFFDF702DFDFC5E5468D5FEDD" box="[550,685,295,320]" name="El Salvador" pageId="5" pageNumber="446">El Salvador</collectingCountry>
|
||
: Usulután.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |