443 lines
52 KiB
XML
443 lines
52 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.82527" ID-GBIF-Dataset="4433ecc4-d495-4d4f-a8fb-baebcf8b7d4c" ID-PMC="PMC9046370" ID-Pensoft-Pub="1313-2970-1097-65" ID-Pensoft-UUID="8FCFCC1D8266501EA79AAE732D44FF0B" ID-PubMed="35837584" ID-ZooBank="6FC8EE30904F4E8D9EF9996F81F30693" ModsDocID="1313-2970-1097-65" checkinTime="1650499969150" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Gao, Zhizhong & Zhang, Feng" docDate="2022" docId="42A9FC8DEA135DFD8418A156A6B47383" docLanguage="en" docName="ZooKeys 1097: 65-83" docOrigin="ZooKeys 1097" docPubDate="2022-04-20" docSource="http://dx.doi.org/10.3897/zookeys.1097.82527" docTitle="Lagynochthonius laoxueyanensis Hou & Gao & Zhang 2022, sp. nov." docType="treatment" docUuid="4BD3052B-5217-4D93-B6FF-77DA99011228" docUuidSource="ZooBank" docVersion="4" id="8FCFCC1D8266501EA79AAE732D44FF0B" lastPageNumber="65" masterDocId="8FCFCC1D8266501EA79AAE732D44FF0B" masterDocTitle="Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China" masterLastPageNumber="83" masterPageNumber="65" pageNumber="65" updateTime="1668151692745" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Xinzhou Teachers University, Xinzhou 034000, Shanxi Province, China</mods:affiliation>
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<mods:nameIdentifier type="email">gaozhizhong1987@126.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zhang, Feng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1097</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>65</mods:start>
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<mods:end>83</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1097.82527</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1097.82527</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1097-65</mods:identifier>
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<subSection pageId="0" pageNumber="65" type="taxonomy">
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<treatment ID-GBIF-Taxon="194685363" LSID="urn:lsid:zoobank.org:act:4BD3052B-5217-4D93-B6FF-77DA99011228" httpUri="http://treatment.plazi.org/id/42A9FC8DEA135DFD8418A156A6B47383" lastPageNumber="65" pageId="0" pageNumber="65">
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<subSubSection pageId="0" pageNumber="65" type="nomenclature">
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<paragraph pageId="0" pageNumber="65">
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<taxonomicName LSID="http://zoobank.org/4BD3052B-5217-4D93-B6FF-77DA99011228" authority="Hou & Gao & Zhang, 2022" authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis" status="sp. nov.">Lagynochthonius laoxueyanensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="65">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="description">
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<paragraph pageId="0" pageNumber="65">
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lagynochthonius laoxueyanensis sp. nov., female habitus. Photographed when it crawled on stony natural habitat." figureDoi="10.3897/zookeys.1097.82527.figure7" httpUri="https://binary.pensoft.net/fig/674773" pageId="0" pageNumber="65">
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Chinese name.
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<normalizedToken originalValue="老穴岩拉伪蝎">老穴岩拉伪蝎</normalizedToken>
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Figs 7
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</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lagynochthonius laoxueyanensis sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure8" httpUri="https://binary.pensoft.net/fig/674774" pageId="0" pageNumber="65">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="type material">
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<paragraph pageId="0" pageNumber="65">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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(Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China A Biyu Cave, Tyrannochthonius pandus sp. nov. B Laoxueyan Cave, Lagynochthonius laoxueyanensis sp. nov." figureDoi="10.3897/zookeys.1097.82527.figure1" httpUri="https://binary.pensoft.net/fig/674767" pageId="0" pageNumber="65">1B</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Laoxueyan Cave, type locality of Lagynochthonius laoxueyanensis sp. nov. A entrance B the only access to the deepest part of the cave C inside the cave entrance D area where L. laoxueyanensis specimens were collected." figureDoi="10.3897/zookeys.1097.82527.figure11" httpUri="https://binary.pensoft.net/fig/674777" pageId="0" pageNumber="65">11</figureCitation>
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)
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Holotype</emphasis>
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: China • ♂; Yunnan Province, Yanshan County, Zhela Township, Liuzhao Village, Laoxueyan Cave;
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<geoCoordinate degrees="23" direction="north" minutes="39.03" orientation="latitude" precision="9" value="23.6505">23°39.03'N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="35.74" orientation="longitude" precision="9" value="104.595665">104°35.74'E</geoCoordinate>
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; 1665 m a.s.l.; 17 Oct. 2021; Zegang Feng, Yanmeng Hou, Lu Zhang and Liu Fu leg.; dark zone; Ps.-
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<emphasis bold="true" pageId="0" pageNumber="65">MHBU</emphasis>
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-HBUARA#2021-445-01.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Paratypes</emphasis>
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: • 2♀; the same data as the holotype; Ps.-
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<emphasis bold="true" pageId="0" pageNumber="65">MHBU</emphasis>
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-HBUARA#2021-445-02, Ps.-
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<emphasis bold="true" pageId="0" pageNumber="65">MSWU</emphasis>
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-HBUARA#2021-445-03.
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</paragraph>
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<caption doi="10.3897/zookeys.1097.82527.figure7" httpUri="https://binary.pensoft.net/fig/674773" pageId="0" pageNumber="65" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="65">
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<emphasis bold="true" pageId="0" pageNumber="65">Figure 7.</emphasis>
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<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
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<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
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</taxonomicName>
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sp. nov., female habitus. Photographed when it crawled on stony natural habitat.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="diagnosis">
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<paragraph pageId="0" pageNumber="65">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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Moderately sized troglomorphic species with elongate appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulate, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites I-II with two setae. Pedipalps slender, femur 8.54 times longer than broad; chela 7.71 times longer than broad; chela fingers gently curved in dorsal view and fixed finger with a modified accessory tooth on dorso-antiaxial face (
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<emphasis italics="true" pageId="0" pageNumber="65">td</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="etymology">
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<paragraph pageId="0" pageNumber="65">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="65">Latinized adjective derived from the name of the type locality, Laoxueyan Cave, Yunnan Province, China.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="65" type="description">
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<paragraph pageId="0" pageNumber="65">Description.</paragraph>
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<paragraph pageId="0" pageNumber="65">
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<emphasis bold="true" pageId="0" pageNumber="65">Adult male</emphasis>
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(Figs
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lagynochthonius laoxueyanensis sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure8" httpUri="https://binary.pensoft.net/fig/674774" pageId="0" pageNumber="65">8A</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9A-D</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Color</emphasis>
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: generally pale yellow, chelicera, pedipalps and tergites slightly darker, soft parts pale (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lagynochthonius laoxueyanensis sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure8" httpUri="https://binary.pensoft.net/fig/674774" pageId="0" pageNumber="65">8A</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Cephalothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9B</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10A</figureCitation>
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): carapace 1.02 times longer than broad, gently narrowed posteriorly, surface smooth, without furrows but with 1 small lyrifissure; anterior margin thin, finely denticulate; without any traces of eyes; epistome very pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10A</figureCitation>
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). Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/3 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10C</figureCitation>
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); intercoxal tubercle absent; without sub-oral seta.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Chelicera</emphasis>
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(Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9C</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10B</figureCitation>
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): large, about same length as carapace, 2.37 times longer than broad; 5 setae on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation both ventral and dorsal side. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 10 retrorse continuous small teeth; galea weakly raised, keel-like (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10B</figureCitation>
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). Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig.
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10D</figureCitation>
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).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10E-G</figureCitation>
|
||
): surface of palpal segments smooth; chelal palm gradually constricted towards fingers; setae generally long and acuminate; femur 8.54, patella 2.73, chela 7.71, hand 3.10 times longer than broad; movable finger 1.45 times longer than hand and 0.58 times longer than chela, apodeme complex of movable finger strongly sclerotized. Femur and dorsal hand without tactile setae but with 1 lyrifissure distally at patella (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10E</figureCitation>
|
||
). Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">ib</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">isb</emphasis>
|
||
situated close together, submedially on dorsum of chelal hand;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">eb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">esb</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">ist</emphasis>
|
||
forming a straight oblique row at base of fixed chelal finger;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">it</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">et</emphasis>
|
||
slightly near to tip of fixed finger, very close to chelal teeth;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">dx</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">et</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">sb</emphasis>
|
||
situated closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">st</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">t</emphasis>
|
||
situated subdistally,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
situated at same level as
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="65">t</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
and at same level as
|
||
<emphasis italics="true" pageId="0" pageNumber="65">it</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10F</figureCitation>
|
||
). Microsetae (chemosensory setae) absent on hand and both chelal fingers. Sensilla absent but with 1 lyrifissure between
|
||
<emphasis italics="true" pageId="0" pageNumber="65">t</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">b</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="65">it</emphasis>
|
||
and
|
||
<emphasis italics="true" pageId="0" pageNumber="65">est</emphasis>
|
||
, respectively (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10G</figureCitation>
|
||
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller distally and proximally: fixed finger with 24 large, erect, well-spaced teeth, without intercalary teeth; movable finger with 10 small (slightly smaller than the teeth on fixed finger), retrorse, serrated and well-spaced teeth; fixed finger also with a modified accessory tooth on dorso-antiaxial face (
|
||
<emphasis italics="true" pageId="0" pageNumber="65">td</emphasis>
|
||
) (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10F, G</figureCitation>
|
||
). Chelal fingers gently curved in dorsal view.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Opisthosoma</emphasis>
|
||
: generally typical; pleural membrane finely granulate. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 2: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0; tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IV-XII: 13: 11: 9: 9: 9: 9: 9: 0: 2; sternites IV-X with unpaired median seta. Anterior genital operculum with 9 setae, genital opening slit-like, with 15 marginal setae on each side (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9D</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Lagynochthonius laoxueyanensis sp. nov., holotype male A carapace (dorsal view) with a detail of anterior margin B left chelicera (dorsal view) with details of teeth and tip of movable finger C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods) G finger tips of chela (lateral view) with detail of modified tooth H leg I without trochanter (lateral view) I leg IV (lateral view). Scale bars: 0.25 mm (A, B, E, F, H, I); 0.10 mm (C, D, G)." figureDoi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65">10H, I</figureCitation>
|
||
): generally typical, long and slender. Fine granulation present on anterodorsal faces of trochanter IV and patella IV. Femur of leg I 1.88 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.23 times longer than tibia. Femoropatella of leg IV 3.76 times longer than deep; tibia 6.33 times longer than deep; with tactile setae on both tarsal segments: basitarsus 3.57 times longer than deep, with basal tactile setae (TS = 0.24), telotarsus 12.80 times longer than deep and 2.56 times longer than basitarsus (TS = 0.41). Setae of leg I (trochanter to tibia) 3: 11: 11: 14, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 14: 10. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="65">Dimensions of male holotype</emphasis>
|
||
(length/width or, in the case of the legs, length/depth in mm; ratios in parentheses). Body length 1.78. Pedipalps: trochanter 0.29/0.16 (1.81), femur 1.11/0.13 (8.54), patella 0.41/0.15 (2.73), chela 1.62/0.21 (7.71), hand length 0.65/0.21 (3.10), movable finger length 0.94. Chelicera 0.64/0.27 (2.37), movable finger length 0.34. Carapace 0.59/0.58 (1.02). Leg I: trochanter 0.17/0.11 (1.55), femur 0.60/0.07 (8.57), patella 0.32/0.07 (4.57), tibia 0.30/0.05 (6.00), tarsus 0.67/0.04 (16.75). Leg IV: trochanter 0.21/0.14 (1.50), femoropatella 0.79/0.21 (3.76), tibia 0.57/0.09 (6.33), basitarsus 0.25/0.07 (3.57), telotarsus 0.64/0.05 (12.80).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure8" httpUri="https://binary.pensoft.net/fig/674774" pageId="0" pageNumber="65" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 8.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
holotype male, dorsal view
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
paratype female, dorsal view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Adult females</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Lagynochthonius laoxueyanensis sp. nov., female habitus. Photographed when it crawled on stony natural habitat." figureDoi="10.3897/zookeys.1097.82527.figure7" httpUri="https://binary.pensoft.net/fig/674773" pageId="0" pageNumber="65">7</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Lagynochthonius laoxueyanensis sp. nov. A holotype male, dorsal view B paratype female, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.82527.figure8" httpUri="https://binary.pensoft.net/fig/674774" pageId="0" pageNumber="65">8B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Lagynochthonius laoxueyanensis sp. nov., holotype male (A-D), paratype female (E) A chelal fingers (lateral view) B carapace (dorsal view) C left chelicera (dorsal view) D male genital area (ventral view) E female genital area (ventral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65">9E</figureCitation>
|
||
). Mostly same as males. Anterior genital operculum with 9 setae plus 10-12 setae on posterior margin. Body length 2.00-2.05. Pedipalps: trochanter 0.30-0.32/0.17-0.18 (1.76-1.78), femur 1.17/0.14-0.15 (7.80-8.36), patella 0.40-0.41/0.17 (2.35-2.41), chela 1.65-1.66/0.23-0.24 (6.88-7.22), hand length 0.66-0.70/0.23-0.24 (2.75-3.04), movable finger length 0.95-0.98. Chelicera 0.68-0.70/0.29-0.30 (2.33-2.34), movable finger length 0.36-0.37. Carapace 0.62/0.61 (1.02). Leg I: trochanter 0.18-0.20/0.13 (1.38-1.54), femur 0.61/0.07 (8.71), patella 0.32-0.33/0.07 (4.57-4.71), tibia 0.30/0.06 (5.00), tarsus 0.68-0.69/0.05 (13.60-13.80). Leg IV: trochanter 0.22-0.24/0.13-0.14 (1.69-1.71), femoropatella 0.81-0.82/0.21 (3.86-3.90), tibia 0.60-0.61/0.09-0.10 (6.10-6.67), basitarsus 0.26-0.27/0.06 (4.33-4.50), telotarsus 0.69/0.05 (13.80).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure9" httpUri="https://binary.pensoft.net/fig/674775" pageId="0" pageNumber="65" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 9.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A-D</emphasis>
|
||
), paratype female (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
chelal fingers (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
left chelicera (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="65" type="remarks">
|
||
<paragraph pageId="0" pageNumber="65">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
Of all
|
||
<taxonomicName authorityName="Beier" authorityYear="1951" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius</emphasis>
|
||
</taxonomicName>
|
||
species found in hypogean environments around the world, only three species,
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
Li, Liu & Shi, 2019 (from China),
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. typhlus" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="typhlus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. typhlus</emphasis>
|
||
</taxonomicName>
|
||
Muchmore, 1991 (from Jamaica) and
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. curvidigitatus" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="curvidigitatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. curvidigitatus</emphasis>
|
||
</taxonomicName>
|
||
Mahnert, 1997 (from Spain), have no eyes, and are all highly troglomorphic species.
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. typhlus" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="typhlus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. typhlus</emphasis>
|
||
</taxonomicName>
|
||
in having only 2 setae on tergites I-II, but the latter has intercalary teeth on the chelal fingers and a smaller body size (1.28 vs. 2.00-2.05 mm in females).
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily separated from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
by its smaller body size (
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
1.78 mm in male, 2.00-2.05 mm in females; while
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
is 2.55-2.92 mm in males, 2.72-2.95 mm in females), the number of setae on the anterior tergites (tergites I-II with 2 setae in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
, but 4 in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
), the shape of epistome (pointed and small in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
, but obtuse and inconspicuous in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
) and the number of setae on the pedipalpal coxa (3 setae in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
, but 5 in
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="bailongtanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. bailongtanensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily separated from
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. curvidigitatus" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="curvidigitatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. curvidigitatus</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a pair of curved chelal fingers in the latter and the number of setae on tergites I-II (
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
with 2 and 2 setae, respectively, but
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. curvidigitatus" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="curvidigitatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. curvidigitatus</emphasis>
|
||
</taxonomicName>
|
||
with 3 and 4 setae, respectively) (
|
||
<bibRefCitation DOI="https://doi.org/10.2307/3495247" author="Muchmore, WB" journalOrPublisher="The Florida Entomologist" pageId="0" pageNumber="65" pagination="110 - 121" refId="B21" refString="Muchmore, WB, 1991. Pseudoscorpions from Florida and the Caribbean area. 14. New species of Tyrannochthonius and Lagynochthonius from caves in Jamacia [sic], with discussion of the genera (Chthoniidae). The Florida Entomologist 74 (1): 110 - 121, DOI: https://doi.org/10.2307/3495247" title="Pseudoscorpions from Florida and the Caribbean area. 14. New species of Tyrannochthonius and Lagynochthonius from caves in Jamacia [sic], with discussion of the genera (Chthoniidae)." url="https://doi.org/10.2307/3495247" volume="74" year="1991">Muchmore 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.80011" author="Mahnert, V" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="65" pagination="559 - 585" refId="B15" refString="Mahnert, V, 1997. New species and records of pseudoscorpions (Arachnida, Pseudoscorpiones) from the Canary Islands. Revue Suisse de Zoologie 104: 559 - 585, DOI: https://doi.org/10.5962/bhl.part.80011" title="New species and records of pseudoscorpions (Arachnida, Pseudoscorpiones) from the Canary Islands." url="https://doi.org/10.5962/bhl.part.80011" volume="104" year="1997">Mahnert 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1071/IS07025" author="Edward, KL" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="65" pagination="259 - 293" refId="B4" refString="Edward, KL, Harvey, MS, 2008. Short-range endemism in hypogean environments: the pseudoscorpion genera Tyrannochthonius and Lagynochthonius (Pseudoscorpiones: Chthoniidae) in the semiarid zone of Western Australia. Invertebrate Systematics 22 (2): 259 - 293, DOI: https://doi.org/10.1071/IS07025" title="Short-range endemism in hypogean environments: the pseudoscorpion genera Tyrannochthonius and Lagynochthonius (Pseudoscorpiones: Chthoniidae) in the semiarid zone of Western Australia." url="https://doi.org/10.1071/IS07025" volume="22" year="2008">Edward and Harvey 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Mahnert, V" journalOrPublisher="Revista Iberica de Aracnologia" pageId="0" pageNumber="65" pagination="27 - 45" refId="B18" refString="Mahnert, V, 2011. A nature's treasury: Pseudoscorpion diversity of the Canary Islands, with the description of nine new species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) and new records. Revista Iberica de Aracnologia 19: 27 - 45" title="A nature's treasury: Pseudoscorpion diversity of the Canary Islands, with the description of nine new species (Pseudoscorpiones, Chthoniidae, Cheiridiidae) and new records." volume="19" year="2011">Mahnert 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4571.1.2" author="Li, YC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="65" pagination="28 - 34" refId="B14" refString="Li, YC, Liu, H, Shi, AM, 2019. A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa 4571 (1): 28 - 34, DOI: https://doi.org/10.11646/zootaxa.4571.1.2" title="A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China." url="https://doi.org/10.11646/zootaxa.4571.1.2" volume="4571" year="2019">Li et al. 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure10" httpUri="https://binary.pensoft.net/fig/674776" pageId="0" pageNumber="65" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 10.</emphasis>
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
carapace (dorsal view) with a detail of anterior margin
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
left chelicera (dorsal view) with details of teeth and tip of movable finger
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
coxal spines on coxae II (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
rallum
|
||
<emphasis bold="true" pageId="0" pageNumber="65">E</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">F</emphasis>
|
||
left chela (lateral view) with details of teeth and with trichobothrial pattern (abbreviations explained in Material and methods)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">G</emphasis>
|
||
finger tips of chela (lateral view) with detail of modified tooth
|
||
<emphasis bold="true" pageId="0" pageNumber="65">H</emphasis>
|
||
leg I without trochanter (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="65">I</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.25 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A, B, E, F, H, I</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C, D, G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="65" type="distribution">
|
||
<paragraph pageId="0" pageNumber="65">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="65">
|
||
This species is only known from the type locality, Laoxueyan Cave (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China A Biyu Cave, Tyrannochthonius pandus sp. nov. B Laoxueyan Cave, Lagynochthonius laoxueyanensis sp. nov." figureDoi="10.3897/zookeys.1097.82527.figure1" httpUri="https://binary.pensoft.net/fig/674767" pageId="0" pageNumber="65">1B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Laoxueyan Cave, type locality of Lagynochthonius laoxueyanensis sp. nov. A entrance B the only access to the deepest part of the cave C inside the cave entrance D area where L. laoxueyanensis specimens were collected." figureDoi="10.3897/zookeys.1097.82527.figure11" httpUri="https://binary.pensoft.net/fig/674777" pageId="0" pageNumber="65">11</figureCitation>
|
||
) which is located about 4 km southeast of Liuzhao Village (Yanshan County). The entrance of the cave is slit-shaped (18 m high and 4 m wide), and the total length of the cave is 88.5 m, and the vertical height of the cave is about 30 m. A descent access leads to the bottom of the cave. The bottom of the cave is a large space, covered with gravel, temperature around 13 °C, humidity over 90%. All of the specimens were collected from ground crevices near the end of the cave.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1097.82527.figure11" httpUri="https://binary.pensoft.net/fig/674777" pageId="0" pageNumber="65" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="65">
|
||
<emphasis bold="true" pageId="0" pageNumber="65">Figure 11.</emphasis>
|
||
Laoxueyan Cave, type locality of
|
||
<taxonomicName authorityName="Hou & Gao & Zhang" authorityYear="2022" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius laoxueyanensis" order="Pseudoscorpiones" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">Lagynochthonius laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="65">A</emphasis>
|
||
entrance
|
||
<emphasis bold="true" pageId="0" pageNumber="65">B</emphasis>
|
||
the only access to the deepest part of the cave
|
||
<emphasis bold="true" pageId="0" pageNumber="65">C</emphasis>
|
||
inside the cave entrance
|
||
<emphasis bold="true" pageId="0" pageNumber="65">D</emphasis>
|
||
area where
|
||
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="65" phylum="Arthropoda" rank="species" species="laoxueyanensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="65">L. laoxueyanensis</emphasis>
|
||
</taxonomicName>
|
||
specimens were collected.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |