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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.445.8370" ID-GBIF-Dataset="3e66a561-c6f9-4de1-baab-5ea3b9d6bf9c" ID-PMC="PMC4205742" ID-Pensoft-Pub="1313-2970-445-97" ID-PubMed="25349509" ID-ZBK="4AA8D97F225A4EEE81F6697D1E1A8723" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-445-97" ModsDocOrigin="ZooKeys 445" ModsDocTitle="On the validity of Epeorella Ulmer, 1939 (Ephemeroptera, Heptageniidae) with general considerations on the Heptageniidae of the Sunda Islands" checkinTime="1451245241452" checkinUser="pensoft" docAuthor="Sartori, Michel" docDate="2014" docId="1B65BF778A0FF3000B2A5F1110FAA751" docLanguage="en" docName="ZooKeys 445: 97-106" docOrigin="ZooKeys 445" docSource="http://dx.doi.org/10.3897/zookeys.445.8370" docTitle="Epeorella borneonia Ulmer 1939" docType="treatment" docVersion="3" lastPageNumber="99" masterDocId="FFCB6872FFC1FFFE0B0FFFEDFFA0FF9E" masterDocTitle="On the validity of Epeorella Ulmer, 1939 (Ephemeroptera, Heptageniidae) with general considerations on the Heptageniidae of the Sunda Islands" masterLastPageNumber="106" masterPageNumber="97" pageNumber="98" updateTime="1668159478256" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On the validity of Epeorella Ulmer, 1939 (Ephemeroptera, Heptageniidae) with general considerations on the Heptageniidae of the Sunda Islands</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sartori, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>445</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>97</mods:start>
<mods:end>106</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.445.8370</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.445.8370</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-445-97</mods:identifier>
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<mods:identifier type="ZooBank">4AA8D97F225A4EEE81F6697D1E1A8723</mods:identifier>
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<treatment ID-GBIF-Taxon="152055822" LSID="urn:lsid:plazi:treatment:1B65BF778A0FF3000B2A5F1110FAA751" httpUri="http://treatment.plazi.org/id/1B65BF778A0FF3000B2A5F1110FAA751" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98">
<subSubSection pageId="1" pageNumber="98" type="multiple">
<paragraph pageId="1" pageNumber="98">Taxon classification Animalia Ephemeroptera Heptageniidae</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="98" type="nomenclature">
<paragraph pageId="1" pageNumber="98">
<taxonomicName authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" class="Insecta" family="Heptageniidae" genus="Epeorella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeorella borneonia" order="Ephemeroptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="borneonia">Epeorella borneonia Ulmer, 1939</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="98" type="reference_group">
<paragraph pageId="1" pageNumber="98">
<taxonomicName class="Insecta" family="Heptageniidae" genus="Epeorella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeorella borneonia" order="Ephemeroptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="borneonia">Epeorella borneonia</taxonomicName>
:
<bibRefCitation author="Ulmer, G" journalOrPublisher="Archiv fuer Hydrobiologie" pageId="7" pageNumber="104" pagination="443 - 692" title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln." volume="16" year="1939">Ulmer 1939</bibRefCitation>
: 579 (male and female imago).
</paragraph>
<paragraph pageId="1" pageNumber="98">
<taxonomicName class="Insecta" family="Heptageniidae" genus="Epeorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeorus borneonia" order="Ephemeroptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="borneonia">Epeorus borneonia</taxonomicName>
:
<bibRefCitation author="Wang, T-Q" journalOrPublisher="Transactions of the American Entomological Society" pageId="7" pageNumber="104" pagination="11 - 45" title="Heptageniidae (Ephemeroptera) of the world. Part I: Phylogenetic higher classification." volume="130" year="2004">Wang and McCafferty 2004</bibRefCitation>
: 21.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="98" type="material examined">
<paragraph pageId="1" pageNumber="98">Material examined.</paragraph>
<paragraph pageId="1" pageNumber="98">
One male imago, one female imago, one female subimago, all bearing the following labels: 1) Type [typewriting on red label], 2) Borneo, Nanga Serawei, 12-18.11.1924 3) Sammelreise Prof. Dr. H. Winckler, ded. 1924-1925 4) Z.M.H. Hamburg 5) G. Ulmer det. 1942 Vers. 13.9.1927. This last label is confusing, and according to Prof. H.
<normalizedToken originalValue="Strümpel">Struempel</normalizedToken>
(in litt.) it is a probable mistake.
</paragraph>
<paragraph pageId="1" pageNumber="98">
The male imago was wrongly mentioned as holotype by
<bibRefCitation author="Weidner, H" journalOrPublisher="Mitteilungen der hamburger zoologischer Museum und Institut" pageId="7" pageNumber="104" pagination="81 - 109" title="Die entomologische Sammlungen des zoologischen Staatsinstituts und zoologischen Museums Hamburg. Insecta I." volume="60" year="1962">Weidner (1962)</bibRefCitation>
. This terminology cannot be accepted because the
<normalizedToken originalValue="“holotype”">&quot;holotype&quot;</normalizedToken>
has not been designated by Ulmer and cannot be ascertained by the presence of a single specimen (see also Recommendation 73F. Avoidance of assumption of holotype,
<bibRefCitation author="Gillies, MT" journalOrPublisher="The international trust for zoological nomenclature, London" pageId="6" pageNumber="103" title="International Code of Zoological Nomenclature." year="1999">ICZN 1999</bibRefCitation>
).
</paragraph>
<paragraph pageId="1" pageNumber="98">
The male imago is accordingly designated as LECTOTYPE of the species
<taxonomicName class="Insecta" family="Heptageniidae" genus="Epeorella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeorella borneonia" order="Ephemeroptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="borneonia">Epeorella borneonia</taxonomicName>
by present designation.
</paragraph>
<paragraph pageId="1" pageNumber="98">
The three specimens have been adequately described by
<bibRefCitation author="Ulmer, G" journalOrPublisher="Archiv fuer Hydrobiologie" pageId="7" pageNumber="104" pagination="443 - 692" title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln." volume="16" year="1939">Ulmer (1939)</bibRefCitation>
. Only significant morphological characters are mentioned here.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="98" type="male imago">
<paragraph pageId="1" pageNumber="98">Male imago.</paragraph>
<paragraph pageId="1" pageNumber="98">Anterior margin of the head not protruding anteriorly (Fig. 1); median depression of mesothoracic furcasternum subparallel, not convergent anteriorly (Fig. 3); mesonotum with a transverse suture (Fig. 2); styliger fig strongly convex, penis lobes minute, rounded and closely tight together (Fig. 4), without apparent sclerites or titillators [a complete analysis of the genitalia will be presented later with the help of non-invasive techniques].</paragraph>
<caption pageId="1" pageNumber="98">
<paragraph pageId="1" pageNumber="98">
Figures 1-6.
<taxonomicName class="Insecta" family="Heptageniidae" genus="Epeorella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeorella borneonia" order="Ephemeroptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="borneonia">Epeorella borneonia</taxonomicName>
Ulmer, 1939 1 Lectotype male imago in dorsal view 2 Details of the mesonotum with transversal suture (arrow) 3 Detail of the mesothoracic furcasternum depression (arrow) 4 Detail of the genitalia in ventral view 5 Female subimago in lateral view 6 Detail of abdominal ridge (arrows) in dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="2" pageNumber="99" type="female imago">
<paragraph pageId="2" pageNumber="99">
<pageBreakToken pageId="2" pageNumber="99" start="start">Female</pageBreakToken>
imago.
</paragraph>
<paragraph pageId="2" pageNumber="99">Abdominal patterns similar to the male. Extracted eggs from the rehydrated specimen were unfortunately not in a satisfactory state for chorionic structure examination through SEM.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="99" type="female subimago">
<paragraph pageId="2" pageNumber="99">Female subimago.</paragraph>
<paragraph pageId="2" pageNumber="99">
Similar to the female, except abdominal terga
<normalizedToken originalValue="VIVIII">VI-VIII</normalizedToken>
(
<normalizedToken originalValue="IVV">IV-V</normalizedToken>
to a lesser extent) present the remains of a longitudinal ridge (Figs 5-6).
</paragraph>
</subSubSection>
</treatment>
</document>