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<document id="CD1F7054F1D524A311ABD57A19893079" ID-CLB-Dataset="27343" ID-DOI="10.11646/zootaxa.4585.1.13" ID-GBIF-Dataset="168fa4f3-250e-41a7-b4a5-78c76f2501c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="2636181" ID-ZooBank="F655970E-887C-480A-879E-0C11E8536CE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554964583492" checkinUser="plazi" docAuthor="Yoshitake, Hiraku, Bollino, Maurizio &amp; Sandel, Franco" docDate="2019" docId="C06087E4887B3167FF1CFA99FE05399B" docLanguage="en" docName="zootaxa.4585.1.13.pdf" docOrigin="Zootaxa 4585 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyrhynchus phaleratus Waterhouse 1841" docType="treatment" docVersion="7" lastPageNumber="198" masterDocId="3C59FF9C887B3166FF8BFF87FFF13D24" masterDocTitle="Pachyrhynchus elenae Rukmane, 2016, a new synonym of Pachyrhynchus phaleratus Waterhouse, 1841 (Coleoptera: Curculionidae: Entiminae)" masterLastPageNumber="200" masterPageNumber="197" pageNumber="197" updateTime="1698719481126" updateUser="ExternalLinkService">
<mods:mods id="769D86BFEA2D36B58AA0F755FD1343E9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3351FC6AEF7033F106B8243F41C7219F">Pachyrhynchus elenae Rukmane, 2016, a new synonym of Pachyrhynchus phaleratus Waterhouse, 1841 (Coleoptera: Curculionidae: Entiminae)</mods:title>
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<mods:namePart id="C1994F75128EA1A1C4081D42A77BEC56">Bollino, Maurizio</mods:namePart>
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<mods:date id="225B0744FB53822AE82F40C3FE280F94">2019</mods:date>
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<subSubSection id="00D36579887B3166FF1CFA99FD33381C" box="[151,706,1310,1336]" pageId="0" pageNumber="197" type="nomenclature">
<paragraph id="487636F2887B3166FF1CFA99FD33381C" blockId="0.[151,1436,1310,2014]" box="[151,706,1310,1336]" pageId="0" pageNumber="197">
<heading id="133E819E887B3166FF1CFA99FD33381C" bold="true" box="[151,706,1310,1336]" fontSize="11" level="1" pageId="0" pageNumber="197" reason="1">
<taxonomicName id="8FC94D71887B3166FF1CFA99FD33381C" ID-CoL="6THRZ" authority="Waterhouse, 1841" authorityName="Waterhouse" authorityYear="1841" box="[151,706,1310,1336]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887B3166FF1CFA99FD33381C" bold="true" box="[151,706,1310,1336]" pageId="0" pageNumber="197">
<emphasis id="7ABDEAE0887B3166FF1CFA99FE2D381C" bold="true" box="[151,476,1310,1336]" italics="true" pageId="0" pageNumber="197">Pachyrhynchus phaleratus</emphasis>
<bibRefCitation id="2C584B03887B3166FE68FA99FD33381C" author="Waterhouse, G. R." box="[483,706,1310,1336]" pageId="0" pageNumber="197" pagination="1841 - 1846" refId="ref2172" refString="Waterhouse, G. R. (1841) Untitled [Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands.] Journal of Proceedings of the Entomological Society of London, March 1, 1841, 23 - 26. [1841 - 1846]" type="book chapter" year="1841">Waterhouse, 1841</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<taxonomicName id="8FC94D71887B3166FF1CFAC6FD9D3872" ID-CoL="6THRZ" authority="Waterhouse 1841: 24" authorityName="Waterhouse" authorityPageNumber="24" authorityYear="1841" box="[151,620,1345,1366]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887B3166FF1CFAC6FE663872" box="[151,407,1345,1366]" italics="true" pageId="0" pageNumber="197">Pachyrhynchus phaleratus</emphasis>
<treatmentCitation id="C96810E3887B3166FE14FAC6FD9D3872" author="Waterhouse, G. R." box="[415,620,1345,1366]" page="24" pageId="0" pageNumber="197" year="1841">
<bibRefCitation id="2C584B03887B3166FE14FAC6FDBB3872" author="Waterhouse, G. R." box="[415,586,1345,1366]" pageId="0" pageNumber="197" pagination="1841 - 1846" refId="ref2172" refString="Waterhouse, G. R. (1841) Untitled [Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands.] Journal of Proceedings of the Entomological Society of London, March 1, 1841, 23 - 26. [1841 - 1846]" type="book chapter" year="1841">Waterhouse 1841</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="97728850887B3166FDF1FAC6FD543872" box="[634,677,1345,1366]" pageId="0" pageNumber="197">type</typeStatus>
locality: “the
<collectingCountry id="30DE7662887B3166FCBBFAC6FC133872" box="[816,994,1345,1366]" name="Philippines" pageId="0" pageNumber="197">Philippine Islands</collectingCountry>
”;
<typeStatus id="97728850887B3166FC7DFAC6FBEE3872" box="[1014,1055,1345,1366]" pageId="0" pageNumber="197">type</typeStatus>
depository: BMNH); 1842, 218; 1843, 320.—
<treatmentCitation id="C96810E3887B3166FE8DFAD8FE2D3850" author="Schoenherr, C. J." box="[262,476,1375,1396]" page="385" pageId="0" pageNumber="197" year="1845">
<bibRefCitation id="2C584B03887B3166FE8DFAD8FE413850" author="Schoenherr, C. J." box="[262,432,1375,1396]" pageId="0" pageNumber="197" refId="ref2017" refString="Schoenherr, C. J. (1845) Genera et species Curculionidum, cum synonymia hujus familiae. Species novae aut hactenus minus cognitae, descriptionibus a Dom. Leonardo Gyllenhal, C. H. Boheman, et entomologis aliis illustratae. Tomus octavus. Pars secunda. Supplementum continens. Roret, Parisiis and Fleischer, Lipsiae, viii + 504 pp., 27 pls. [Synopsis geographica]" type="book" year="1845">Schoenherr 1845</bibRefCitation>
: 385
</treatmentCitation>
(reprinting of the original description; “
<emphasis id="7ABDEAE0887B3166FCD5FAD8FC9B3850" box="[862,874,1375,1396]" italics="true" pageId="0" pageNumber="197">P</emphasis>
. Ins.
<collectingCountry id="30DE7662887B3166FC2BFAD8FBE53850" box="[928,1044,1375,1396]" name="Philippines" pageId="0" pageNumber="197">Philippines</collectingCountry>
.”).
</treatmentCitationGroup>
</paragraph>
<paragraph id="487636F2887B3166FF1CFAFBFB9F3B22" blockId="0.[151,1436,1310,2014]" pageId="0" pageNumber="197">
<treatmentCitationGroup id="68D911DC887B3166FF1CFAFBFB9F3B22" infered="true" pageId="0" pageNumber="197">
<emphasis id="7ABDEAE0887B3166FF1CFAFBFEC538B5" box="[151,308,1404,1425]" italics="true" pageId="0" pageNumber="197">Pachyrrhynchus</emphasis>
<taxonomicName id="8FC94D71887B3166FEB7FAFBFDA938B5" ID-CoL="6THRZ" authority=": Heller 1912: 309" authorityName=": Heller" authorityPageNumber="309" authorityYear="1912" box="[316,600,1404,1425]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887B3166FEB7FAFBFE5B38B5" box="[316,426,1404,1425]" italics="true" pageId="0" pageNumber="197">phaleratus:</emphasis>
<treatmentCitation id="C96810E3887B3166FE39FAFBFDA938B5" author="Heller, K. M." box="[434,600,1404,1425]" page="309" pageId="0" pageNumber="197" year="1912">
<bibRefCitation id="2C584B03887B3166FE39FAFBFDDB38B5" author="Heller, K. M." box="[434,554,1404,1425]" pageId="0" pageNumber="197" pagination="295 - 346" refId="ref1943" refString="Heller, K. M. (1912) Philippinische Russelkafer. Philippine Journal of Science, (D), Manila, 7 (5), 295 - 346." type="journal article" year="1912">Heller 1912</bibRefCitation>
: 309
</treatmentCitation>
</taxonomicName>
(placed in “GRUPPE V”; in key), 321 (in diagnosis of
<emphasis id="7ABDEAE0887B3166FBFFFAFBFAA238B5" box="[1140,1363,1404,1425]" italics="true" pageId="0" pageNumber="197">Pachyrrhynchus stellio</emphasis>
<bibRefCitation id="2C584B03887B3166FAD0FAFBFF09388A" author="Heller, K. M." pageId="0" pageNumber="197" pagination="295 - 346" refId="ref1943" refString="Heller, K. M. (1912) Philippinische Russelkafer. Philippine Journal of Science, (D), Manila, 7 (5), 295 - 346." type="journal article" year="1912">Heller, 1912</bibRefCitation>
).—
<treatmentCitation id="C96810E3887B3166FE90FA1EFE2D388A" author="Schultze, W." box="[283,476,1433,1454]" page="578" pageId="0" pageNumber="197" year="1922">
<bibRefCitation id="2C584B03887B3166FE90FA1EFE5E388A" author="Schultze, W." box="[283,431,1433,1454]" pageId="0" pageNumber="197" pagination="569 - 596" refId="ref2091" refString="Schultze, W. (1922) X. Beitrag zur Coleopteren Fauna der Philippinen. Philippine Journal of Science, Manila, 21 (6), 569 - 596, 4 pls." type="book chapter" year="1922">Schultze, 1922</bibRefCitation>
: 578
</treatmentCitation>
(in diagnosis of
<emphasis id="7ABDEAE0887B3166FDF6FA1EFC60388A" box="[637,913,1433,1454]" italics="true" pageId="0" pageNumber="197">Pachyrrhynchus halconensis</emphasis>
<bibRefCitation id="2C584B03887B3166FC1CFA1EFBD8388A" author="Schultze, W." box="[919,1065,1433,1454]" pageId="0" pageNumber="197" pagination="569 - 596" refId="ref2091" refString="Schultze, W. (1922) X. Beitrag zur Coleopteren Fauna der Philippinen. Philippine Journal of Science, Manila, 21 (6), 569 - 596, 4 pls." type="book chapter" year="1922">Schultze, 1922</bibRefCitation>
), 593 (male aedeagal structures), pl. 3, fig. 13 (male aedeagus in dorsal and lateral views); 1923: 636 (placed in “Group I”; in key), 646 (redescription; “
<collectingRegion id="8A0DF810887B3166FA89FA30FA6738EF" box="[1282,1430,1463,1484]" country="Philippines" name="Catanduanes" pageId="0" pageNumber="197">CATANDUANES</collectingRegion>
, Virac”), pl. 2, fig. 30 (lateral habitus, female), pl. 3, fig. 24 (male aedeagus in dorsal and lateral views), pl. 5, fig. 19 (dorsal habitus, female; “
<collectingRegion id="8A0DF810887B3166FEE4FA76FE1C3B22" box="[367,493,1521,1542]" country="Philippines" name="Catanduanes" pageId="0" pageNumber="197">Catanduanes</collectingRegion>
, Virac”).—
<treatmentCitation id="C96810E3887B3166FDD1FA76FCA83B22" author="Dalla Torre &amp; Emden, M. van &amp; Emden, F. van" box="[602,857,1521,1542]" page="35" pageId="0" pageNumber="197" year="1931">
<bibRefCitation id="2C584B03887B3166FDD1FA76FCC93B22" author="Dalla Torre &amp; Emden, M. van &amp; Emden, F. van" box="[602,824,1521,1542]" pageId="0" pageNumber="197" refId="ref1831" refString="Dalla Torre †, K. W. von, Emden, M. van &amp; Emden, F. van (1931) Coleopterorum Catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Pars 119. Curculionidae: Brachyderinae: Pachyrrhynchini. Junk, Berlin, 44 pp." type="book" year="1931">
Dalla Torre
<emphasis id="7ABDEAE0887B3166FD45FA76FD0E3B22" box="[718,767,1521,1542]" italics="true" pageId="0" pageNumber="197">et al.</emphasis>
1931
</bibRefCitation>
: 35
</treatmentCitation>
(cataloged; “
<collectingRegion id="8A0DF810887B3166FC52FA76FBAB3B22" box="[985,1114,1521,1542]" country="Philippines" name="Catanduanes" pageId="0" pageNumber="197">Catanduanes</collectingRegion>
”).
</treatmentCitationGroup>
</paragraph>
<paragraph id="487636F2887B3166FF1CF988FAD33B65" blockId="0.[151,1436,1310,2014]" pageId="0" pageNumber="197">
<treatmentCitationGroup id="68D911DC887B3166FF1CF988FAD33B65" infered="true" pageId="0" pageNumber="197">
<taxonomicName id="8FC94D71887B3166FF1CF988FDDC3B00" ID-CoL="8JBJ3" authority="Rukmane 2016: 82" authorityName="Rukmane" authorityPageNumber="82" authorityYear="2016" box="[151,557,1551,1572]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887B3166FF1CF988FE9E3B00" box="[151,367,1551,1572]" italics="true" pageId="0" pageNumber="197">Pachyrhynchus elenae</emphasis>
<treatmentCitation id="C96810E3887B3166FEFEF988FDDC3B00" author="Rukmane, A." box="[373,557,1551,1572]" page="82" pageId="0" pageNumber="197" year="2016">
<bibRefCitation id="2C584B03887B3166FEFEF988FDFA3B00" author="Rukmane, A." box="[373,523,1551,1572]" pageId="0" pageNumber="197" pagination="77 - 89" refId="ref1975" refString="Rukmane, A. (2016) Six new species of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae) from the Philippines. Acta Biologica Universitatis Daugavpiliensis, Ilgas, 16 (1), 77 - 89." type="journal article" year="2016">Rukmane 2016</bibRefCitation>
: 82
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="97728850887B3166FDB1F988FD943B00" box="[570,613,1551,1572]" pageId="0" pageNumber="197">type</typeStatus>
locality: “Mindanao Isl.,
<collectingRegion id="8A0DF810887B3166FCD7F988FC443B00" box="[860,949,1551,1572]" country="Philippines" name="South Cotabato" pageId="0" pageNumber="197">Cotabato</collectingRegion>
, Mt. Apo”—mislabeled;
<typeStatus id="97728850887B3166FB22F988FB233B00" box="[1193,1234,1551,1572]" pageId="0" pageNumber="197">type</typeStatus>
depository: DUBC), fig. 4A, B (dorsal and lateral habitus, male), 4E, F (male aedeagus in dorsal and lateral views). NEW SYNONYMY.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="00D36579887B3167FF1CF9CDFD4A3DD0" lastPageId="1" lastPageNumber="198" pageId="0" pageNumber="197" type="materials_examined">
<paragraph id="487636F2887B3166FF1CF9CDFA6D3B45" blockId="0.[151,1436,1310,2014]" box="[151,1436,1609,1633]" pageId="0" pageNumber="197">
<emphasis id="7ABDEAE0887B3166FF1CF9CDFE433B45" bold="true" box="[151,434,1610,1633]" pageId="0" pageNumber="197">Type material examined.</emphasis>
Male
<typeStatus id="97728850887B3166FE7DF9CDFDA23B45" box="[502,595,1610,1633]" pageId="0" pageNumber="197" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8FC94D71887B3166FDF3F9CEFC653B45" authorityName=": Heller" authorityYear="1912" box="[632,916,1609,1633]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="197" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887B3166FDF3F9CEFC653B45" box="[632,916,1609,1633]" italics="true" pageId="0" pageNumber="197">Pachyrhynchus phaleratus</emphasis>
</taxonomicName>
(BMNH). The
<typeStatus id="97728850887B3166FBB0F9CDFB693B45" box="[1083,1176,1610,1633]" pageId="0" pageNumber="197" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="8FC94D71887B3166FB73F9CEFA6D3B45" authorityName="Rukmane" authorityYear="2016" box="[1272,1436,1609,1633]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887B3166FB73F9CEFA6D3B45" box="[1272,1436,1609,1633]" italics="true" pageId="0" pageNumber="197">Pachyrhynchus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="487636F2887B3166FF1CF9ECFA6A3BA7" blockId="0.[151,1436,1310,2014]" box="[151,1435,1643,1667]" pageId="0" pageNumber="197">
<emphasis id="7ABDEAE0887B3166FF1CF9ECFF2C3BA7" box="[151,221,1643,1667]" italics="true" pageId="0" pageNumber="197">elenae</emphasis>
(DUBC) was not examined, but the images of habitus and aedeagus published along with the original description
</paragraph>
<paragraph id="487636F2887B3166FF1CF908FDEE3B82" blockId="0.[151,1436,1310,2014]" box="[151,543,1679,1702]" pageId="0" pageNumber="197">clearly support the above synonymy.</paragraph>
<paragraph id="487636F2887B3166FF4CF935FA6A3BED" blockId="0.[151,1436,1310,2014]" box="[199,1435,1714,1737]" pageId="0" pageNumber="197">
<emphasis id="7ABDEAE0887B3166FF4CF935FE4C3BED" bold="true" box="[199,445,1714,1737]" pageId="0" pageNumber="197">Specimens examined.</emphasis>
<materialsCitation id="F8A13CAF887B3166FE4CF935FA6A3BED" ID-GBIF-Occurrence="2235579703" box="[455,1435,1714,1737]" collectingDate="2010-06" collectionCode="CFS" country="Philippines" location="Aurora" pageId="0" pageNumber="197" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Aurora">
<collectingCountry id="30DE7662887B3166FE4CF935FD993BED" box="[455,616,1714,1737]" name="Philippines" pageId="0" pageNumber="197">PHILIPPINES</collectingCountry>
: Luzon. Sierra Madre,
<collectingRegion id="8A0DF810887B3166FCE2F935FC483BED" box="[873,953,1714,1737]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
:
<specimenCount id="5ECFFD7B887B3166FC4DF935FBEA3BED" box="[966,1051,1714,1737]" pageId="0" pageNumber="197" type="male">3 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FBDDF935FB323BED" box="[1110,1219,1714,1737]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FB5BF935FAC33BED" box="[1232,1330,1714,1737]" pageId="0" pageNumber="197" value="2010-06">
<collectingDate id="2C33E9DA887B3166FB5BF935FAC33BED" box="[1232,1330,1714,1737]" pageId="0" pageNumber="197" value="2010-06">VI. 2010</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FACFF935FA893BED" box="[1348,1400,1714,1737]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
); 1
</materialsCitation>
</paragraph>
<paragraph id="487636F2887B3166FF1CF953FA6D3BCF" blockId="0.[151,1436,1310,2014]" box="[151,1436,1748,1771]" pageId="0" pageNumber="197">
female,
<date id="3C771032887B3166FF66F953FECE3BCF" box="[237,319,1748,1771]" pageId="0" pageNumber="197" value="2012-05">V. 2012</date>
(NIAES).
<materialsCitation id="F8A13CAF887B3166FE38F953FBC43BCF" ID-GBIF-Occurrence="2235579687" box="[435,1077,1748,1771]" collectingDate="2012-06" collectionCode="NIAES" location="Dingalan" pageId="0" pageNumber="197" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Aurora">
<location id="4D166029887B3166FE38F953FDE63BCF" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887B3166FE38F953FDE63BCF" box="[435,535,1748,1771]" name="Dingalan" pageId="0" pageNumber="197" stateProvince="Aurora">Dingalan</location>
,
<collectingRegion id="8A0DF810887B3166FDA9F953FD833BCF" box="[546,626,1748,1771]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
:
<specimenCount id="5ECFFD7B887B3166FDF7F953FD353BCF" box="[636,708,1748,1771]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FD7CF953FC933BCF" box="[759,866,1748,1771]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FCE7F953FC3B3BCF" box="[876,970,1748,1771]" pageId="0" pageNumber="197" value="2012-06">
<collectingDate id="2C33E9DA887B3166FCE7F953FC3B3BCF" box="[876,970,1748,1771]" pageId="0" pageNumber="197" value="2012-06">VI. 2012</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FC52F953FBDC3BCF" box="[985,1069,1748,1771]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="0" pageNumber="197">NIAES</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FBB4F953FACF3BCF" ID-GBIF-Occurrence="2235579708" box="[1087,1342,1748,1771]" collectingDate="2012-09" collectionCode="CFS" isTailEnumeration="true" location="Dingalan" pageId="0" pageNumber="197" specimenCount="1" specimenCount-male="1" stateProvince="Aurora">
<specimenCount id="5ECFFD7B887B3166FBB4F953FB7B3BCF" box="[1087,1162,1748,1771]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
,
<date id="3C771032887B3166FB1FF953FB023BCF" box="[1172,1267,1748,1771]" pageId="0" pageNumber="197" value="2012-09">
<collectingDate id="2C33E9DA887B3166FB1FF953FB023BCF" box="[1172,1267,1748,1771]" pageId="0" pageNumber="197" value="2012-09">IX. 2012</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FA89F953FAC73BCF" box="[1282,1334,1748,1771]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
; 22 exs.,
</paragraph>
<paragraph id="487636F2887B3166FF1CF970FA6A3A2A" blockId="0.[151,1436,1310,2014]" box="[151,1435,1783,1806]" pageId="0" pageNumber="197">
<materialsCitation id="F8A13CAF887B3166FF1CF970FDAC3A2A" ID-GBIF-Occurrence="2235579701" box="[151,605,1783,1806]" collectingDate="2013-03" collectionCode="MBLI" pageId="0" pageNumber="197" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<date id="3C771032887B3166FF1CF970FF073A2A" box="[151,246,1783,1806]" pageId="0" pageNumber="197" value="2013-03">
<collectingDate id="2C33E9DA887B3166FF1CF970FF073A2A" box="[151,246,1783,1806]" pageId="0" pageNumber="197" value="2013-03">III. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FE8CF970FEBE3A2A" box="[263,335,1783,1806]" pageId="0" pageNumber="197">MBLI</collectionCode>
);
<specimenCount id="5ECFFD7B887B3166FEEFF970FE493A2A" box="[356,440,1783,1806]" pageId="0" pageNumber="197" type="male">2 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FE7AF970FDAC3A2A" box="[497,605,1783,1806]" pageId="0" pageNumber="197" type="female">2 females</specimenCount>
</materialsCitation>
,
<materialsCitation id="F8A13CAF887B3166FDE1F970FC883A2A" ID-GBIF-Occurrence="2235579694" box="[618,889,1783,1806]" collectingDate="2013-04" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="2" specimenCount-male="2">
<date id="3C771032887B3166FDE1F970FD363A2A" box="[618,711,1783,1806]" pageId="0" pageNumber="197" value="2013-04">
<collectingDate id="2C33E9DA887B3166FDE1F970FD363A2A" box="[618,711,1783,1806]" pageId="0" pageNumber="197" value="2013-04">IV. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FD53F970FCFD3A2A" box="[728,780,1783,1806]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
);
<specimenCount id="5ECFFD7B887B3166FCA9F970FC883A2A" box="[802,889,1783,1806]" pageId="0" pageNumber="197" type="male">2 males</specimenCount>
</materialsCitation>
,
<materialsCitation id="F8A13CAF887B3166FC0DF970FADE3A2A" ID-GBIF-Occurrence="2235579705" box="[902,1327,1783,1806]" collectingDate="2013-07" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<date id="3C771032887B3166FC0DF970FC013A2A" box="[902,1008,1783,1806]" pageId="0" pageNumber="197" value="2013-07">
<collectingDate id="2C33E9DA887B3166FC0DF970FC013A2A" box="[902,1008,1783,1806]" pageId="0" pageNumber="197" value="2013-07">VII. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FB8AF970FBC43A2A" box="[1025,1077,1783,1806]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
);
<specimenCount id="5ECFFD7B887B3166FBC0F970FB643A2A" box="[1099,1173,1783,1806]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FB46F970FADE3A2A" box="[1229,1327,1783,1806]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
</materialsCitation>
,
<date id="3C771032887B3166FAB7F970FA6A3A2A" box="[1340,1435,1783,1806]" pageId="0" pageNumber="197" value="2014-03">III. 2014</date>
</paragraph>
<paragraph id="487636F2887B3166FF1CF89DFA6A3A15" blockId="0.[151,1436,1310,2014]" box="[151,1435,1817,1841]" pageId="0" pageNumber="197">
(CFS);
<materialsCitation id="F8A13CAF887B3166FF6DF89DFD7C3A15" ID-GBIF-Occurrence="2235579690" box="[230,653,1818,1841]" collectingDate="2014-09" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="6" specimenCount-female="3" specimenCount-male="3">
<specimenCount id="5ECFFD7B887B3166FF6DF89DFEC83A15" box="[230,313,1818,1841]" pageId="0" pageNumber="197" type="male">3 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FEE5F89DFE293A15" box="[366,472,1818,1841]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FE69F89DFDB03A15" box="[482,577,1818,1841]" pageId="0" pageNumber="197" value="2014-09">
<collectingDate id="2C33E9DA887B3166FE69F89DFDB03A15" box="[482,577,1818,1841]" pageId="0" pageNumber="197" value="2014-09">IX. 2014</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FDDAF89DFD743A15" box="[593,645,1818,1841]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FD13F89DFA753A15" ID-GBIF-Occurrence="2235579704" box="[664,1412,1817,1841]" collectingDate="2013-03-09" collectingDateMax="2013-03-12" collectingDateMin="2013-03-09" collectionCode="NIAES" collectorName="M. Maruyama" pageId="0" pageNumber="197" specimenCount="10" specimenCount-female="3" specimenCount-male="7">
<specimenCount id="5ECFFD7B887B3166FD13F89DFD1A3A15" box="[664,747,1818,1841]" pageId="0" pageNumber="197" type="male">7 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FC94F89DFC7B3A15" box="[799,906,1818,1841]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FC1EF89DFBC93A15" box="[917,1080,1818,1841]" pageId="0" pageNumber="197" value="2013-03-09" valueMax="2013-03-12" valueMin="2013-03-09">
<collectingDate id="2C33E9DA887B3166FC1EF89DFBC93A15" box="[917,1080,1818,1841]" pageId="0" pageNumber="197" value="2013-03-09" valueMax="2013-03-12" valueMin="2013-03-09">9–12. III. 2013</collectingDate>
</date>
,
<collectorName id="E53C5324887B3166FBC8F89DFB2A3A15" box="[1091,1243,1818,1841]" pageId="0" pageNumber="197">M. Maruyama</collectorName>
<emphasis id="7ABDEAE0887B3166FB68F89DFAE83A15" box="[1251,1305,1817,1841]" italics="true" pageId="0" pageNumber="197">et al.</emphasis>
(
<collectionCode id="2ED8AE37887B3166FAA3F89DFA8D3A15" box="[1320,1404,1818,1841]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="0" pageNumber="197">NIAES</collectionCode>
)
</materialsCitation>
; 1
</paragraph>
<paragraph id="487636F2887B3166FF1CF8BBFA6A3A77" blockId="0.[151,1436,1310,2014]" box="[151,1435,1852,1875]" pageId="0" pageNumber="197">
male and
<materialsCitation id="F8A13CAF887B3166FF76F8BBFDFF3A77" ID-GBIF-Occurrence="2235579695" box="[253,526,1852,1875]" collectingDate="2015-03" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="1" specimenCount-female="1">
<specimenCount id="5ECFFD7B887B3166FF76F8BBFEAD3A77" box="[253,348,1852,1875]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FEECF8BBFE323A77" box="[359,451,1852,1875]" pageId="0" pageNumber="197" value="2015-03">
<collectingDate id="2C33E9DA887B3166FEECF8BBFE323A77" box="[359,451,1852,1875]" pageId="0" pageNumber="197" value="2015-03">III. 2015</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FE5AF8BBFDF43A77" box="[465,517,1852,1875]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FD93F8BBFCBB3A77" ID-GBIF-Occurrence="2235579702" box="[536,842,1852,1875]" collectingDate="2015-11" collectionCode="NIAES" pageId="0" pageNumber="197" specimenCount="1" specimenCount-female="1">
<specimenCount id="5ECFFD7B887B3166FD93F8BBFD863A77" box="[536,631,1852,1875]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FD0AF8BBFD2E3A77" box="[641,735,1852,1875]" pageId="0" pageNumber="197" value="2015-11">
<collectingDate id="2C33E9DA887B3166FD0AF8BBFD2E3A77" box="[641,735,1852,1875]" pageId="0" pageNumber="197" value="2015-11">XI. 2015</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FD66F8BBFCB03A77" box="[749,833,1852,1875]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="0" pageNumber="197">NIAES</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FCDFF8BBFBBE3A77" ID-GBIF-Occurrence="2235579688" box="[852,1103,1852,1875]" collectingDate="2016-03" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="1" specimenCount-male="1">
<specimenCount id="5ECFFD7B887B3166FCDFF8BBFC6C3A77" box="[852,925,1852,1875]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
,
<date id="3C771032887B3166FC23F8BBFBF53A77" box="[936,1028,1852,1875]" pageId="0" pageNumber="197" value="2016-03">
<collectingDate id="2C33E9DA887B3166FC23F8BBFBF53A77" box="[936,1028,1852,1875]" pageId="0" pageNumber="197" value="2016-03">III. 2016</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FB98F8BBFBB63A77" box="[1043,1095,1852,1875]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<specimenCount id="5ECFFD7B887B3166FBD1F8BBFB503A77" box="[1114,1185,1852,1875]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FB5FF8BBFAC23A77" box="[1236,1331,1852,1875]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FAB6F8BBFA6A3A77" box="[1341,1435,1852,1875]" pageId="0" pageNumber="197" value="2017-09">IX. 2017</date>
</paragraph>
<paragraph id="487636F2887B3166FF1CF8D8FA6D3A52" blockId="0.[151,1436,1310,2014]" box="[151,1436,1887,1910]" pageId="0" pageNumber="197">
(CFS);
<materialsCitation id="F8A13CAF887B3166FF60F8D8FDF63A52" ID-GBIF-Occurrence="2235579693" box="[235,519,1887,1910]" collectingDate="2018-02" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="1" specimenCount-female="1">
<specimenCount id="5ECFFD7B887B3166FF60F8D8FEBE3A52" box="[235,335,1887,1910]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FED5F8D8FE463A52" box="[350,439,1887,1910]" pageId="0" pageNumber="197" value="2018-02">
<collectingDate id="2C33E9DA887B3166FED5F8D8FE463A52" box="[350,439,1887,1910]" pageId="0" pageNumber="197" value="2018-02">II. 2018</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FE40F8D8FE0E3A52" box="[459,511,1887,1910]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FD9CF8D8FC2A3A52" ID-GBIF-Occurrence="2235579689" box="[535,987,1887,1910]" collectingDate="2018-03" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="8" specimenCount-female="5" specimenCount-male="3">
<specimenCount id="5ECFFD7B887B3166FD9CF8D8FD9F3A52" box="[535,622,1887,1910]" pageId="0" pageNumber="197" type="male">3 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FD20F8D8FCEB3A52" box="[683,794,1887,1910]" pageId="0" pageNumber="197" type="female">5 females</specimenCount>
,
<date id="3C771032887B3166FCA2F8D8FC7A3A52" box="[809,907,1887,1910]" pageId="0" pageNumber="197" value="2018-03">
<collectingDate id="2C33E9DA887B3166FCA2F8D8FC7A3A52" box="[809,907,1887,1910]" pageId="0" pageNumber="197" value="2018-03">III. 2018</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FC15F8D8FC233A52" box="[926,978,1887,1910]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FC60F8D8FA693A52" ID-GBIF-Occurrence="2235579706" box="[1003,1432,1887,1910]" collectingDate="2018-04" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="5ECFFD7B887B3166FC60F8D8FBC93A52" box="[1003,1080,1887,1910]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FBFEF8D8FB293A52" box="[1141,1240,1887,1910]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FB63F8D8FAB93A52" box="[1256,1352,1887,1910]" pageId="0" pageNumber="197" value="2018-04">
<collectingDate id="2C33E9DA887B3166FB63F8D8FAB93A52" box="[1256,1352,1887,1910]" pageId="0" pageNumber="197" value="2018-04">IV. 2018</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FAD7F8D8FA613A52" box="[1372,1424,1887,1910]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="487636F2887B3166FF1CF805FA6A3ABD" blockId="0.[151,1436,1310,2014]" box="[151,1435,1922,1945]" pageId="0" pageNumber="197">
Ditumabo,
<collectingRegion id="8A0DF810887B3166FE85F805FEAF3ABD" box="[270,350,1922,1945]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
:
<specimenCount id="5ECFFD7B887B3166FEE3F805FE433ABD" box="[360,434,1922,1945]" pageId="0" pageNumber="197" type="male">1 male</specimenCount>
,
<date id="3C771032887B3166FE37F805FDEB3ABD" box="[444,538,1922,1945]" pageId="0" pageNumber="197" value="2018-06">VI. 2018</date>
(CFS). Labuyo,
<collectingRegion id="8A0DF810887B3166FD45F805FCEF3ABD" box="[718,798,1922,1945]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
:
<specimenCount id="5ECFFD7B887B3166FCACF805FC883ABD" box="[807,889,1922,1945]" pageId="0" pageNumber="197" type="male">3 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FC27F805FBFA3ABD" box="[940,1035,1922,1945]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FB9EF805FB903ABD" box="[1045,1121,1922,1945]" pageId="0" pageNumber="197" value="2009-01">I. 2009</date>
(NIAES);
<specimenCount id="5ECFFD7B887B3166FB5EF805FAC53ABD" box="[1237,1332,1922,1945]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FAB5F805FA6A3ABD" box="[1342,1435,1922,1945]" pageId="0" pageNumber="197" value="2010-03">III. 2010</date>
</paragraph>
<paragraph id="487636F2887B3166FF1CF823FA6D3A9F" blockId="0.[151,1436,1310,2014]" box="[151,1436,1956,1979]" pageId="0" pageNumber="197">
(NIAES);
<materialsCitation id="F8A13CAF887B3166FE8FF823FDEE3A9F" ID-GBIF-Occurrence="2235579699" box="[260,543,1956,1979]" collectingDate="2011-06" collectionCode="CFS" pageId="0" pageNumber="197" specimenCount="3" specimenCount-female="3">
<specimenCount id="5ECFFD7B887B3166FE8FF823FE9F3A9F" box="[260,366,1956,1979]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FEF3F823FE253A9F" box="[376,468,1956,1979]" pageId="0" pageNumber="197" value="2011-06">
<collectingDate id="2C33E9DA887B3166FEF3F823FE253A9F" box="[376,468,1956,1979]" pageId="0" pageNumber="197" value="2011-06">VI. 2011</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FE68F823FDE63A9F" box="[483,535,1956,1979]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887B3166FDA1F823FC2C3A9F" ID-GBIF-Occurrence="2235579709" box="[554,989,1956,1979]" collectingDate="2013-05" collectionCode="NIAES" pageId="0" pageNumber="197" specimenCount="9" specimenCount-female="3" specimenCount-male="6">
<specimenCount id="5ECFFD7B887B3166FDA1F823FD8A3A9F" box="[554,635,1956,1979]" pageId="0" pageNumber="197" type="male">6 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FD25F823FCE63A9F" box="[686,791,1956,1979]" pageId="0" pageNumber="197" type="female">3 females</specimenCount>
,
<date id="3C771032887B3166FCAAF823FC833A9F" box="[801,882,1956,1979]" pageId="0" pageNumber="197" value="2013-05">
<collectingDate id="2C33E9DA887B3166FCAAF823FC833A9F" box="[801,882,1956,1979]" pageId="0" pageNumber="197" value="2013-05">V. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FC0BF823FC253A9F" box="[896,980,1956,1979]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="0" pageNumber="197">NIAES</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F8A13CAF887B3166FC6DF823FA6D3A9F" ID-GBIF-Occurrence="2235579696" box="[998,1436,1956,1979]" collectingDate="2012-03" collectionCode="CFS" location="San Luis" pageId="0" pageNumber="197" specimenCount="1" stateProvince="Aurora">
<location id="4D166029887B3166FC6DF823FBB73A9F" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887B3166FC6DF823FBB73A9F" box="[998,1094,1956,1979]" name="San Luis" pageId="0" pageNumber="197" stateProvince="Aurora">San Luis</location>
,
<collectingRegion id="8A0DF810887B3166FBDBF823FB513A9F" box="[1104,1184,1956,1979]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
: 22 exs.,
<date id="3C771032887B3166FA8BF823FAAC3A9F" box="[1280,1373,1956,1979]" pageId="0" pageNumber="197" value="2012-03">
<collectingDate id="2C33E9DA887B3166FA8BF823FAAC3A9F" box="[1280,1373,1956,1979]" pageId="0" pageNumber="197" value="2012-03">III. 2012</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FAE0F823FA6D3A9F" box="[1387,1436,1956,1979]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="0" pageNumber="197">CFS</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="487636F2887B3166FF1CF840FA6A3AFA" blockId="0.[151,1436,1310,2014]" box="[151,1435,1991,2014]" pageId="0" pageNumber="197">
&amp;
<materialsCitation id="F8A13CAF887B3166FF3CF840FD253AFA" ID-GBIF-Occurrence="2235579692" box="[183,724,1991,2014]" collectionCode="MBLI" location="San Luis" municipality="Barangay Dimanayat" pageId="0" pageNumber="197" specimenCount="1" stateProvince="Aurora">
<collectionCode id="2ED8AE37887B3166FF3CF840FF0F3AFA" box="[183,254,1991,2014]" pageId="0" pageNumber="197">MBLI</collectionCode>
).
<collectingMunicipality id="A812AC88887B3166FE9EF840FE0E3AFA" box="[277,511,1991,2014]" pageId="0" pageNumber="197">Barangay Dimanayat</collectingMunicipality>
,
<location id="4D166029887B3166FD84F840FD853AFA" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887B3166FD84F840FD853AFA" box="[527,628,1991,2014]" municipality="Barangay Dimanayat" name="San Luis" pageId="0" pageNumber="197" stateProvince="Aurora">San Luis</location>
,
<collectingRegion id="8A0DF810887B3166FD0FF840FD253AFA" box="[644,724,1991,2014]" country="Philippines" name="Aurora" pageId="0" pageNumber="197">Aurora</collectingRegion>
</materialsCitation>
:
<materialsCitation id="F8A13CAF887B3166FD68F840FBEE3AFA" ID-GBIF-Occurrence="2235579707" box="[739,1055,1991,2014]" collectingDate="2012-05" collectionCode="NIAES" pageId="0" pageNumber="197" specimenCount="1" specimenCount-female="1" stateProvince="Aurora">
<specimenCount id="5ECFFD7B887B3166FD68F840FCB63AFA" box="[739,839,1991,2014]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FCDCF840FC5F3AFA" box="[855,942,1991,2014]" pageId="0" pageNumber="197" value="2012-05">
<collectingDate id="2C33E9DA887B3166FCDCF840FC5F3AFA" box="[855,942,1991,2014]" pageId="0" pageNumber="197" value="2012-05">V. 2012</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887B3166FC49F840FBE73AFA" box="[962,1046,1991,2014]" country="Japan" httpUri="http://grbio.org/cool/h0ih-38y8" name="National Institute for Agro-Environmental Sciences" pageId="0" pageNumber="197">NIAES</collectionCode>
)
</materialsCitation>
;
<specimenCount id="5ECFFD7B887B3166FBA5F840FB743AFA" box="[1070,1157,1991,2014]" pageId="0" pageNumber="197" type="male">4 males</specimenCount>
and
<specimenCount id="5ECFFD7B887B3166FB48F840FAD63AFA" box="[1219,1319,1991,2014]" pageId="0" pageNumber="197" type="female">1 female</specimenCount>
,
<date id="3C771032887B3166FAB3F840FA6A3AFA" box="[1336,1435,1991,2014]" pageId="0" pageNumber="197" value="2012-11">XI. 2012</date>
</paragraph>
<paragraph id="487636F2887A3167FF1CFF1FFD4A3DD0" blockId="1.[151,1436,152,1215]" pageId="1" pageNumber="198">
(NIAES).
<materialsCitation id="F8A13CAF887A3167FE8FFF1FFD0F3D8B" ID-GBIF-Occurrence="2235579700" box="[260,766,152,175]" collectingDate="2013-06" collectionCode="CFS" location="Sierra Madre" pageId="1" pageNumber="198" specimenCount="1" specimenCount-male="1" stateProvince="Quirino">
<location id="4D166029887A3167FE8FFF1FFE623D8B" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887A3167FE8FFF1FFE623D8B" box="[260,403,152,175]" name="Sierra Madre" pageId="1" pageNumber="198" stateProvince="Quirino">Sierra Madre</location>
,
<collectingRegion id="8A0DF810887A3167FE15FF1FFE043D8B" box="[414,501,152,175]" country="Philippines" name="Quirino" pageId="1" pageNumber="198">Quirino</collectingRegion>
:
<specimenCount id="5ECFFD7B887A3167FE74FF1FFDBB3D8B" box="[511,586,152,175]" pageId="1" pageNumber="198" type="male">1 male</specimenCount>
,
<date id="3C771032887A3167FDDFFF1FFD423D8B" box="[596,691,152,175]" pageId="1" pageNumber="198" value="2013-06">
<collectingDate id="2C33E9DA887A3167FDDFFF1FFD423D8B" box="[596,691,152,175]" pageId="1" pageNumber="198" value="2013-06">VI. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887A3167FD4AFF1FFD043D8B" box="[705,757,152,175]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="1" pageNumber="198">CFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F8A13CAF887A3167FC82FF1FFBEC3D8B" ID-GBIF-Occurrence="2235579698" box="[777,1053,152,175]" collectingDate="2013-06" collectionCode="CFS" isTailEnumeration="true" location="Sierra Madre" pageId="1" pageNumber="198" specimenCount="1" specimenCount-female="1" stateProvince="Quirino">
<specimenCount id="5ECFFD7B887A3167FC82FF1FFC993D8B" box="[777,872,152,175]" pageId="1" pageNumber="198" type="female">1 female</specimenCount>
,
<date id="3C771032887A3167FCF8FF1FFC203D8B" box="[883,977,152,175]" pageId="1" pageNumber="198" value="2013-06">
<collectingDate id="2C33E9DA887A3167FCF8FF1FFC203D8B" box="[883,977,152,175]" pageId="1" pageNumber="198" value="2013-06">VI. 2013</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887A3167FC6BFF1FFBE53D8B" box="[992,1044,152,175]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="1" pageNumber="198">CFS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F8A13CAF887A3167FBACFF1FFE7D3DF5" ID-GBIF-Occurrence="2235579697" collectingDate="2014-11" collectionCode="CFS" location="Nagtipunan" municipality="Disimugal" pageId="1" pageNumber="198" specimenCount="2" specimenCount-male="2" stateProvince="Quirino">
<collectingMunicipality id="A812AC88887A3167FBACFF1FFB683D8B" box="[1063,1177,152,175]" pageId="1" pageNumber="198">Disimugal</collectingMunicipality>
,
<location id="4D166029887A3167FB2FFF1FFAD23D8B" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887A3167FB2FFF1FFAD23D8B" box="[1188,1315,152,175]" municipality="Disimugal" name="Nagtipunan" pageId="1" pageNumber="198" stateProvince="Quirino">Nagtipunan</location>
,
<collectingRegion id="8A0DF810887A3167FAA6FF1FFA753D8B" box="[1325,1412,152,175]" country="Philippines" name="Quirino" pageId="1" pageNumber="198">Quirino</collectingRegion>
:
<specimenCount id="5ECFFD7B887A3167FA05FF1FFF293DF5" pageId="1" pageNumber="198" type="male">2 males</specimenCount>
,
<date id="3C771032887A3167FF69FF3DFEB13DF5" box="[226,320,186,209]" pageId="1" pageNumber="198" value="2014-11">
<collectingDate id="2C33E9DA887A3167FF69FF3DFEB13DF5" box="[226,320,186,209]" pageId="1" pageNumber="198" value="2014-11">XI. 2014</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887A3167FEDBFF3DFE753DF5" box="[336,388,186,209]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="1" pageNumber="198">CFS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="F8A13CAF887A3167FE1DFF3DFC6D3DF5" ID-GBIF-Occurrence="2235579691" box="[406,924,186,209]" collectingDate="2011-09" collectionCode="CFS" location="Kayapa" pageId="1" pageNumber="198" specimenCount="1" specimenCount-male="1" stateProvince="Nueva Vizcaya">
<location id="4D166029887A3167FE1DFF3DFE1B3DF5" LSID="urn:lsid:plazi:treatment:C06087E4887B3167FF1CFA99FE05399B:4D166029887A3167FE1DFF3DFE1B3DF5" box="[406,490,186,209]" name="Kayapa" pageId="1" pageNumber="198" stateProvince="Nueva Vizcaya">Kayapa</location>
,
<collectingRegion id="8A0DF810887A3167FE7EFF3DFD6A3DF5" box="[501,667,186,209]" country="Philippines" name="Nueva Vizcaya" pageId="1" pageNumber="198">Nueva Vizcaya</collectingRegion>
:
<specimenCount id="5ECFFD7B887A3167FD2FFF3DFD1F3DF5" box="[676,750,186,209]" pageId="1" pageNumber="198" type="male">1 male</specimenCount>
,
<date id="3C771032887A3167FD72FF3DFCAA3DF5" box="[761,859,186,209]" pageId="1" pageNumber="198" value="2011-09">
<collectingDate id="2C33E9DA887A3167FD72FF3DFCAA3DF5" box="[761,859,186,209]" pageId="1" pageNumber="198" value="2011-09">IX. 2011</collectingDate>
</date>
(
<collectionCode id="2ED8AE37887A3167FCE9FF3DFC653DF5" box="[866,916,186,209]" country="Canada" httpUri="http://grbio.org/cool/4rrx-qe60" name="Canadian Forest Service, Pacific Forest Research Centre" pageId="1" pageNumber="198">CFS</collectionCode>
)
</materialsCitation>
. In addition, a series of specimens of both sexes from Virac,
<collectingRegion id="8A0DF810887A3167FE93FF5AFE513DD0" box="[280,416,221,244]" country="Philippines" name="Catanduanes" pageId="1" pageNumber="198">Catanduanes</collectingRegion>
were examined at SMTD.
</paragraph>
</subSubSection>
<subSubSection id="00D36579887A3167FF4CFE87FC423CC3" pageId="1" pageNumber="198" type="distribution">
<paragraph id="487636F2887A3167FF4CFE87FC423CC3" blockId="1.[151,1436,152,1215]" pageId="1" pageNumber="198">
<emphasis id="7ABDEAE0887A3167FF4CFE87FEA83C33" bold="true" box="[199,345,256,279]" pageId="1" pageNumber="198">Distribution.</emphasis>
<collectingCountry id="30DE7662887A3167FEE8FE87FE2A3C33" box="[355,475,256,279]" name="Philippines" pageId="1" pageNumber="198">Philippines</collectingCountry>
(“
<collectingCountry id="30DE7662887A3167FE7EFE87FD303C33" box="[501,705,256,279]" name="Philippines" pageId="1" pageNumber="198">Philippine Islands</collectingCountry>
”; Luzon:
<collectingRegion id="8A0DF810887A3167FCA4FE87FC8C3C33" box="[815,893,256,279]" country="Philippines" name="Aurora" pageId="1" pageNumber="198">Aurora</collectingRegion>
,
<collectingRegion id="8A0DF810887A3167FC00FE87FC103C33" box="[907,993,256,279]" country="Philippines" name="Quirino" pageId="1" pageNumber="198">Quirino</collectingRegion>
, and
<collectingRegion id="8A0DF810887A3167FB94FE87FB563C33" box="[1055,1191,256,279]" country="Philippines" name="Catanduanes" pageId="1" pageNumber="198">Catanduanes</collectingRegion>
Island). Although the precise type locality was not specified (
<bibRefCitation id="2C584B03887A3167FDBCFEA5FD053C1D" author="Waterhouse, G. R." box="[567,756,290,313]" pageId="1" pageNumber="198" pagination="1841 - 1846" refId="ref2172" refString="Waterhouse, G. R. (1841) Untitled [Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands.] Journal of Proceedings of the Entomological Society of London, March 1, 1841, 23 - 26. [1841 - 1846]" type="book chapter" year="1841">Waterhouse 1841</bibRefCitation>
,
<bibRefCitation id="2C584B03887A3167FD75FEA5FCC73C1D" author="Waterhouse, G. R." box="[766,822,290,313]" pageId="1" pageNumber="198" pagination="217 - 220" refId="ref2235" refString="Waterhouse, G. R. (1842) Untitled [Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands.] Annals and Magazine of Natural History, London, Series 1, 8 (50), 217 - 220." type="journal article" year="1842">1842</bibRefCitation>
,
<bibRefCitation id="2C584B03887A3167FCCAFEA5FC883C1D" author="Waterhouse, G. R." box="[833,889,290,313]" pageId="1" pageNumber="198" pagination="310 - 327" refId="ref2295" refString="Waterhouse, G. R. (1843) Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands. Transactions of the Royal Entomological Society of London, 3 (4), 310 - 327. [1841 / 1843]" type="journal article" year="1843">1843</bibRefCitation>
), it is known that Hugh Cuming, during his journey to the
<collectingCountry id="30DE7662887A3167FF57FEC2FEA63C78" box="[220,343,325,348]" name="Philippines" pageId="1" pageNumber="198">Philippines</collectingCountry>
, did not visit
<collectingRegion id="8A0DF810887A3167FE66FEC2FD843C78" box="[493,629,325,348]" country="Philippines" name="Catanduanes" pageId="1" pageNumber="198">Catanduanes</collectingRegion>
nor the provinces of
<collectingRegion id="8A0DF810887A3167FCEBFEC2FC5A3C78" box="[864,939,325,348]" country="Philippines" name="Aurora" pageId="1" pageNumber="198">Aurora</collectingRegion>
or
<collectingRegion id="8A0DF810887A3167FC59FEC2FBD43C78" box="[978,1061,325,348]" country="Philippines" name="Quirino" pageId="1" pageNumber="198">Quirino</collectingRegion>
(
<bibRefCitation id="2C584B03887A3167FBBFFEC2FB2D3C78" author="Broderip, W. J." box="[1076,1244,325,348]" pageId="1" pageNumber="198" pagination="83 - 91" refId="ref1794" refString="Broderip, W. J. (1840) Description of shells collected in the Philippine Islands by Hugh Cuming, Esq. Proceedings of the Zoological Society of London, 8, 83 - 91." type="journal article" year="1840">Broderip, 1840</bibRefCitation>
), while he visited the neighboring province of
<collectingRegion id="8A0DF810887A3167FE50FEEFFD953C5B" box="[475,612,360,383]" country="Philippines" name="Nueva Ecija" pageId="1" pageNumber="198">Nueva Ecija</collectingRegion>
(
<bibRefCitation id="2C584B03887A3167FDFCFEEFFCF93C5B" author="Dance, S. P." box="[631,776,360,383]" pageId="1" pageNumber="198" pagination="477 - 501" refId="ref1885" refString="Dance, S. P. (1980) Hugh Cuming (1791 - 1865) prince of collectors. Journal of the Society for the Bibliography of Natural History, Edinburgh, 9 (4), 477 - 501. https: // doi. org / 10.3366 / jsbnh. 1980.9. PART _ 4.477" type="journal article" year="1980">Dance, 1980</bibRefCitation>
), possibly the actual collecting locality of the
<typeStatus id="97728850887A3167FA97FEEFFA883C5B" box="[1308,1401,360,383]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8FC94D71887A3167FF1CFE0EFE473C85" authorityName=": Heller" authorityYear="1912" box="[151,438,393,417]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FF1CFE0EFE473C85" box="[151,438,393,417]" italics="true" pageId="1" pageNumber="198">Pachyrhynchus phaleratus</emphasis>
</taxonomicName>
. In addition, we studied a single male labeled “
<collectingRegion id="8A0DF810887A3167FC42FE0DFB873C85" box="[969,1142,394,417]" country="Philippines" name="Nueva Vizcaya" pageId="1" pageNumber="198">Nueva Vizcaya</collectingRegion>
”, from where we have not obtained further specimens, for which we exclude the presence of the species in this province at the moment, considering how easy it is for traded or commercially sold specimens to be mislabeled.
</paragraph>
</subSubSection>
<subSubSection id="00D36579887A3167FF4CFE75FE05399B" pageId="1" pageNumber="198" type="discussion">
<paragraph id="487636F2887A3167FF4CFE75FE05399B" blockId="1.[151,1436,152,1215]" pageId="1" pageNumber="198">
<emphasis id="7ABDEAE0887A3167FF4CFE75FEB83F2D" bold="true" box="[199,329,498,521]" pageId="1" pageNumber="198">Comments.</emphasis>
<bibRefCitation id="2C584B03887A3167FEDBFE75FDB83F2D" author="Rukmane, A." box="[336,585,498,521]" pageId="1" pageNumber="198" pagination="77 - 89" refId="ref1975" refString="Rukmane, A. (2016) Six new species of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae) from the Philippines. Acta Biologica Universitatis Daugavpiliensis, Ilgas, 16 (1), 77 - 89." type="journal article" year="2016">Rukmane (2016, p. 83)</bibRefCitation>
provided dorsal and lateral images of the habitus (
<figureCitation id="D0F22A77887A3167FBE4FE75FB123F2D" box="[1135,1251,498,521]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Fig. 4A, B</figureCitation>
) and of the male aedeagus (
<figureCitation id="D0F22A77887A3167FE87FD92FE723F08" box="[268,387,533,556]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Fig. 4E, F</figureCitation>
) of the
<typeStatus id="97728850887A3167FE6EFD92FDB33F08" box="[485,578,533,556]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
and noted morphological differences between
<taxonomicName id="8FC94D71887A3167FBDEFD93FABB3F08" authorityName="Rukmane" authorityYear="2016" box="[1109,1354,532,556]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FBDEFD93FABB3F08" box="[1109,1354,532,556]" italics="true" pageId="1" pageNumber="198">Pachyrhynchus elenae</emphasis>
</taxonomicName>
and
<taxonomicName id="8FC94D71887A3167FA03FD92FEF83F6B" authorityName=": Heller" authorityYear="1912" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FA03FD92FA643F08" box="[1416,1429,533,556]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FF1CFDB0FEF83F6B" box="[151,265,567,591]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
as follows: “
<taxonomicName id="8FC94D71887A3167FE1CFDB0FD763F6B" authority="Yoshitake &amp; Bollino &amp; Sandel, 2019" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[407,647,567,591]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae" status="sp. nov.">
<emphasis id="7ABDEAE0887A3167FE1CFDB0FD763F6B" box="[407,647,567,591]" italics="true" pageId="1" pageNumber="198">Pachyrhynchus elenae</emphasis>
</taxonomicName>
<emphasis id="7ABDEAE0887A3167FD04FDBFFD3A3F6B" bold="true" box="[655,715,568,591]" pageId="1" pageNumber="198">
<taxonomicNameLabel id="618E579B887A3167FD04FDBFFD3A3F6B" box="[655,715,568,591]" pageId="1" pageNumber="198" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar in general appearance to
<taxonomicName id="8FC94D71887A3167FBC3FDBFFA6D3F6B" authority="Waterhouse, 1841" authorityName="Waterhouse" authorityYear="1841" box="[1096,1436,567,591]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FBC3FDBFFBA43F6B" box="[1096,1109,568,591]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FBE8FDB0FB243F6B" box="[1123,1237,567,591]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
<bibRefCitation id="2C584B03887A3167FB57FDBFFA6D3F6B" author="Waterhouse, G. R." box="[1244,1436,568,591]" pageId="1" pageNumber="198" pagination="1841 - 1846" refId="ref2172" refString="Waterhouse, G. R. (1841) Untitled [Descriptions of the species of the curculionideous genus Pachyrhynchus, Sch., collected by H. Cuming, Esq., in the Philippine Islands.] Journal of Proceedings of the Entomological Society of London, March 1, 1841, 23 - 26. [1841 - 1846]" type="book chapter" year="1841">Waterhouse, 1841</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D0F22A77887A3167FF14FDDDFEE43F55" box="[159,277,602,625]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Fig. 4C, D</figureCitation>
). The new species is easily distinguished from
<taxonomicName id="8FC94D71887A3167FC9FFDDDFC533F55" authorityName="Waterhouse" authorityYear="1841" box="[788,930,601,625]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FC9FFDDDFCD03F55" box="[788,801,602,625]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FCBBFDDEFC533F55" box="[816,930,601,625]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
by the shape of aedeagus, which is longer and less curved lateraly [sic!] and by the bigger number of longitudinal lines on each elytron (
<figureCitation id="D0F22A77887A3167FBE7FDFAFB123FB0" box="[1132,1251,637,660]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Fig. 4G, H</figureCitation>
). Pronotum of
<taxonomicName id="8FC94D71887A3167FA03FDFAFF2C3F93" authority="Yoshitake &amp; Bollino &amp; Sandel, 2019" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae" status="sp. nov.">
<emphasis id="7ABDEAE0887A3167FA03FDFAFA643FB0" box="[1416,1429,637,660]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FF1CFD18FF2C3F93" box="[151,221,671,695]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
<emphasis id="7ABDEAE0887A3167FF6EFD27FED23F93" bold="true" box="[229,291,672,695]" pageId="1" pageNumber="198">
<taxonomicNameLabel id="618E579B887A3167FF6EFD27FED23F93" box="[229,291,672,695]" pageId="1" pageNumber="198" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
widest before the midian [sic!] part (pronotum of
<taxonomicName id="8FC94D71887A3167FCC4FD27FC2F3F93" authorityName="Waterhouse" authorityYear="1841" box="[847,990,671,695]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FCC4FD27FCAD3F93" box="[847,860,672,695]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FCE7FD18FC2F3F93" box="[876,990,671,695]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
widest in the middle).” Based on a long series of specimens of
<taxonomicName id="8FC94D71887A3167FE1BFD45FDEE3FFD" authorityName="Waterhouse" authorityYear="1841" box="[400,543,705,729]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FE1BFD45FE6C3FFD" box="[400,413,706,729]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FE26FD46FDEE3FFD" box="[429,543,705,729]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
, it is obvious that markings and body shape are variable (
<figureCitation id="D0F22A77887A3167FB12FD45FAFF3FFD" box="[1177,1294,706,729]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Figs. 1–12</figureCitation>
). Individuals with the ‘
<taxonomicName id="8FC94D71887A3167FE8CFD63FEBD3FD8" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[263,332,740,764]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FE8CFD63FEBD3FD8" box="[263,332,740,764]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
-like’ pattern were collected only from Dingalan and San Luis (
<collectingRegion id="8A0DF810887A3167FBAEFD62FB873FD8" box="[1061,1142,741,764]" country="Philippines" name="Aurora" pageId="1" pageNumber="198">Aurora</collectingRegion>
), while those with typical markings occur in a wide area from
<collectingRegion id="8A0DF810887A3167FDACFC8FFD8B3E3B" box="[551,634,776,799]" country="Philippines" name="Quirino" pageId="1" pageNumber="198">Quirino</collectingRegion>
to the north to
<collectingRegion id="8A0DF810887A3167FCA2FC8FFC403E3B" box="[809,945,776,799]" country="Philippines" name="Catanduanes" pageId="1" pageNumber="198">Catanduanes</collectingRegion>
to the south. However, both localities of the ‘
<taxonomicName id="8FC94D71887A3167FF2BFCAEFF143E65" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[160,229,809,833]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FF2BFCAEFF143E65" box="[160,229,809,833]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
-like’ form overlap with those of the typical form and there are a number of intermediate specimens between the two forms even in the same population. In addition, the prothorax is more or less strongly expanded from the constricted base to its widest point usually situated before the middle, but sometimes precisely so at the middle. With regard to differences in the male genital structures, it is quite clear that the shape of the aedeagus of the
<typeStatus id="97728850887A3167FB39FC15FAFE3E8D" box="[1202,1295,914,937]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8FC94D71887A3167FAB2FC15FA6A3E8D" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[1337,1435,913,937]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FAB2FC15FAB73E8D" box="[1337,1350,914,937]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FADDFC16FA6A3E8D" box="[1366,1435,913,937]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
collapsed probably due to an excessive treatment with KOH solution and subsequent drying, whereas that of the figured
<taxonomicName id="8FC94D71887A3167FF1CFC5FFED53ECB" authorityName="Waterhouse" authorityYear="1841" box="[151,292,983,1007]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FF1CFC5FFF553ECB" box="[151,164,984,1007]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FF39FC50FED53ECB" box="[178,292,983,1007]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
retained its original form (
<bibRefCitation id="2C584B03887A3167FDC8FC5FFC5A3ECB" author="Rukmane, A." box="[579,939,984,1007]" pageId="1" pageNumber="198" pagination="77 - 89" refId="ref1975" refString="Rukmane, A. (2016) Six new species of the genus Pachyrhynchus Germar, 1824 (Coleoptera: Curculionidae) from the Philippines. Acta Biologica Universitatis Daugavpiliensis, Ilgas, 16 (1), 77 - 89." type="journal article" year="2016">
Rukmane 2016: p. 83,
<figureCitation id="D0F22A77887A3167FCB2FC5FFC5A3ECB" box="[825,939,984,1007]" captionStart="FIGURES 1–12" captionStartId="2.[151,258,1938,1960]" captionTargetBox="[216,1386,203,1903]" captionTargetId="figure@2.[211,1386,194,1922]" captionTargetPageId="2" captionText="FIGURES 1–12. Dorsal habitus images of Pachyrhynchus phaleratus Waterhouse, 1843 from Aurora province, Luzon, the Philippines. 1–3. Males from Labuyo. 4–6. Males from San Luis. 7. A female from Dingalan. 8. A female from Sierra Madre. 9, 10. Females from Labuyo. 11, 12. Females from San Luis." httpUri="https://zenodo.org/record/2636183/files/figure.png" pageId="1" pageNumber="198">Fig. 4G, H</figureCitation>
</bibRefCitation>
), a fact that makes a comparison meaningless. Our observations of male genitalia of both forms shows that there are no significant differences to delimit a species boundary, not only in the shape of aedeagus but also in the endophallic structures (
<figureCitation id="D0F22A77887A3167FBB2FB9AFB4E3910" box="[1081,1215,1053,1076]" captionStart="FIGURES 13–18" captionStartId="3.[151,258,1400,1422]" captionTargetBox="[151,1436,192,1378]" captionTargetId="figure@3.[151,1436,191,1378]" captionTargetPageId="3" captionText="FIGURES 13–18. Endophallic structures of Pachyrhynchus phaleratus Waterhouse, 1843.13–15, The typical form (Dingalan, Aurora, Luzon). 16–18, The ‘elenae’ form (San Luis, Aurora, Luzon). 13, 16, Dorsal view. 14, 17, Lateral view. 15, 18, Ventral view." httpUri="https://zenodo.org/record/2636185/files/figure.png" pageId="1" pageNumber="198">Figs. 13–18</figureCitation>
). This demonstrates beyond doubt that
<taxonomicName id="8FC94D71887A3167FED6FBC7FE4C3973" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[349,445,1087,1111]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FED6FBC7FE9B3973" box="[349,362,1088,1111]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FEFCFBB8FE4C3973" box="[375,445,1087,1111]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="8FC94D71887A3167FD1AFBC7FCEF3973" authorityName="Waterhouse" authorityYear="1841" box="[657,798,1087,1111]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FD1AFBC7FD6F3973" box="[657,670,1088,1111]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FD27FBB8FCEF3973" box="[684,798,1087,1111]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
. As far as we know,
<taxonomicName id="8FC94D71887A3167FC70FBC7FB773973" authorityName="Waterhouse" authorityYear="1841" box="[1019,1158,1087,1111]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="phaleratus">
<emphasis id="7ABDEAE0887A3167FC70FBC7FBF93973" box="[1019,1032,1088,1111]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FB9EFBB8FB773973" box="[1045,1158,1087,1111]" italics="true" pageId="1" pageNumber="198">phaleratus</emphasis>
</taxonomicName>
occurs only in Luzon and its adjacent islands. Mount Apo, the type locality of
<taxonomicName id="8FC94D71887A3167FD44FBE5FCC1395D" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[719,816,1121,1145]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FD44FBE5FD2D395D" box="[719,732,1122,1145]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FD60FBE6FCC1395D" box="[747,816,1121,1145]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
, is one of the most famous localities for insect trading in the
<collectingCountry id="30DE7662887A3167FF4AFB02FECD39B8" box="[193,316,1157,1180]" name="Philippines" pageId="1" pageNumber="198">Philippines</collectingCountry>
. The
<typeStatus id="97728850887A3167FEF7FB02FE2839B8" box="[380,473,1157,1180]" pageId="1" pageNumber="198" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8FC94D71887A3167FD88FB02FD9439B8" authorityName="Yoshitake &amp; Bollino &amp; Sandel" authorityYear="2019" box="[515,613,1156,1180]" class="Insecta" family="Curculionidae" genus="Pachyrhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="198" phylum="Arthropoda" rank="species" species="elenae">
<emphasis id="7ABDEAE0887A3167FD88FB02FDE139B8" box="[515,528,1157,1180]" italics="true" pageId="1" pageNumber="198">P</emphasis>
.
<emphasis id="7ABDEAE0887A3167FDABFB03FD9439B8" box="[544,613,1156,1180]" italics="true" pageId="1" pageNumber="198">elenae</emphasis>
</taxonomicName>
, whose collector is unidentified, was most probably mislabeled during the commercial distribution process.
</paragraph>
</subSubSection>
</treatment>
</document>
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