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<document id="83A45010D555E2623F389B76F841EC75" ID-CLB-Dataset="3952" ID-DOI="10.5281/zenodo.1196048" ID-GBIF-Dataset="fed4a06f-7511-4cb0-82de-218f7a6dd132" ID-ISSN="0035-418" ID-Zenodo-Dep="1196048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1634783961719" checkinUser="plazi" docAuthor="Kurbatov, Sergey A., Cuccodoro, Giulio &amp; Sabella, Giorgio" docDate="2018" docId="A7582020FF89FFF6FEE4FEAD52EBEC0A" docLanguage="en" docName="13_Kurbatov_etal.pdf" docOrigin="Revue suisse de Zoologie 125 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Achilia frontalis Jeannel 1962" docType="treatment" docVersion="6" lastPageNumber="179" masterDocId="5B615858FF87FFF8FFFFFFAC5639E96C" masterDocTitle="A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="188" masterPageNumber="165" pageNumber="179" updateTime="1699464010858" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5197DC17BBAF2021AEEA649951319740" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="640AF5A932FB95E1B60512F7DEA0412B">A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="B21FAEEC642AE6284FCDBBFF68F8C0B2" type="personal">
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<mods:namePart id="5F68C0423DD98F7B137E8ED01D4EB399">Kurbatov, Sergey A.</mods:namePart>
<mods:affiliation id="877B6767A3EFB38AF481976DADDCCDFE">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail:</mods:affiliation>
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<mods:namePart id="83E73F6055038A53C627ECB64262E900">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="4A5E09357D2A23DE528003B84A8F340E">Muséum d’histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="619EB7B2B7DD3EAC677B652C2D1510D2" type="email">giulio.cuccodoro@ville-ge.ch</mods:nameIdentifier>
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<mods:namePart id="F9FE4078192A2FA76A13DCCC7B661FA5">Sabella, Giorgio</mods:namePart>
<mods:nameIdentifier id="FA6B0AF5BB808385748044E02E227935" type="ORCID">0000-0002-2234-1901</mods:nameIdentifier>
<mods:affiliation id="C638CB77B8DE6CF7B0DA13C26F82CD39">Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dell’Università - sezione Biologia Animale, via</mods:affiliation>
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<mods:title id="F9CC4ED0E6E7E31888F21AA261F4950F">Revue suisse de Zoologie</mods:title>
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<mods:date id="631D3B49BB418F58B8B5103F2E1BE6C5">2018</mods:date>
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<mods:number id="6B537F4D0DE95B80FBD9340D571CFD6D">2018-03-31</mods:number>
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<treatment id="A7582020FF89FFF6FEE4FEAD52EBEC0A" ID-DOI="http://doi.org/10.5281/zenodo.5589980" ID-GBIF-Taxon="189149716" ID-Zenodo-Dep="5589980" LSID="urn:lsid:plazi:treatment:A7582020FF89FFF6FEE4FEAD52EBEC0A" httpUri="http://treatment.plazi.org/id/A7582020FF89FFF6FEE4FEAD52EBEC0A" lastPageNumber="179" pageId="14" pageNumber="179">
<subSubSection id="67EBC2BDFF89FFF6FEE4FEAD5449E875" box="[283,624,257,281]" pageId="14" pageNumber="179" type="nomenclature">
<paragraph id="2F4E9136FF89FFF6FEE4FEAD5449E875" blockId="14.[238,669,257,313]" box="[283,624,257,281]" pageId="14" pageNumber="179">
<heading id="7406265AFF89FFF6FEE4FEAD5449E875" bold="true" box="[283,624,257,281]" centered="true" level="2" pageId="14" pageNumber="179" reason="2">
<taxonomicName id="E8F1EAB5FF89FFF6FEE4FEAD5449E875" ID-CoL="977T" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" box="[283,624,257,281]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FEE4FEAD5414E875" bold="true" box="[283,557,257,281]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FEE4FEAD57F6E875" bold="true" box="[283,463,257,281]" italics="true" pageId="14" pageNumber="179">Achilia frontalis</emphasis>
<bibRefCitation id="4B60ECC7FF89FFF6FE29FEAE5414E875" author="Jeannel R." box="[470,557,258,281]" pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel</bibRefCitation>
</emphasis>
,
<emphasis id="1D854D24FF89FFF6FDC4FEAE5449E875" bold="true" box="[571,624,258,281]" pageId="14" pageNumber="179">1962</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FF11FE8E54A4E855" box="[238,669,290,313]" pageId="14" pageNumber="179" type="description">
<paragraph id="2F4E9136FF89FFF6FF11FE8E54A4E855" blockId="14.[238,669,257,313]" box="[238,669,290,313]" pageId="14" pageNumber="179">
<figureCitation id="B7CA8DB3FF89FFF6FF11FE8E5779E855" box="[238,320,290,313]" captionStart="Figs 24-29" captionStartId="4.[151,192,1401,1422]" captionTargetBox="[216,1403,263,1340]" captionTargetId="figure-245@4.[162,1413,254,1361]" captionTargetPageId="4" captionText="Figs 24-29. Aedeagi of Achilia. (24) A.caracolana, specimen from Hualpèn, prov. Concepción. (25) A. auriculata, specimen from Peniquillo, prov. Concepción. (26) A. ovallensis, specimen from Ovalle, prov. Limarí [paratype]. (27) A. frontalis, specimen from Aguas Calientes to Puyehue National Park, prov. Osorno. (28) A. trauco, specimen from 34 km WNW La Union, prov. Ranco [paratype]. (29) A. longispina, specimen from Mina de la Disputada, prov. Valparaíso." figureDoi="http://doi.org/10.5281/zenodo.5589962" httpUri="https://zenodo.org/record/5589962/files/figure.png" pageId="14" pageNumber="179">Figs 27</figureCitation>
,
<figureCitation id="B7CA8DB3FF89FFF6FEB4FE8E5751E855" box="[331,360,290,313]" captionStart="Figs 30-34" captionStartId="5.[151,192,1388,1409]" captionTargetBox="[189,1397,257,1335]" captionTargetId="figure-284@5.[148,1409,254,1359]" captionTargetPageId="5" captionText="Figs 30-34. Male antennae of Achilia. (30) A. caracolana. (31) A. auriculata. (32) A. ovallensis. (33) A. frontalis. (34) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589966" httpUri="https://zenodo.org/record/5589966/files/figure.png" pageId="14" pageNumber="179">33</figureCitation>
,
<figureCitation id="B7CA8DB3FF89FFF6FE8CFE8E57E5E855" box="[371,476,290,313]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="14" pageNumber="179">37, 38-40</figureCitation>
,
<figureCitation id="B7CA8DB3FF89FFF6FE18FE8E5415E855" box="[487,556,290,313]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="14" pageNumber="179">45, 50</figureCitation>
, 71-76, 84
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FF68FEE35704E8F2" pageId="14" pageNumber="179" type="reference_group">
<paragraph id="2F4E9136FF89FFF6FF68FEE354DBE809" blockId="14.[151,756,335,414]" box="[151,738,335,357]" pageId="14" pageNumber="179">
<treatmentCitationGroup id="0FE1B618FF89FFF6FF68FEE354DBE809" box="[151,738,335,357]" pageId="14" pageNumber="179">
<treatmentCitation id="AE50B727FF89FFF6FF68FEE357C9E809" author="Jeannel R." box="[151,496,335,357]" page="409" pageId="14" pageNumber="179" year="1962">
<taxonomicName id="E8F1EAB5FF89FFF6FF68FEE357C9E809" ID-CoL="977T" authority="Jeannel, 1962: 409" authorityName="Jeannel" authorityPageNumber="409" authorityYear="1962" box="[151,496,335,357]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FF68FEE3570AE808" box="[151,307,335,356]" italics="true" pageId="14" pageNumber="179">Achilia frontalis</emphasis>
<bibRefCitation id="4B60ECC7FF89FFF6FEC6FEFC57C9E809" author="Jeannel R." box="[313,496,336,357]" pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 409</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 413 fig. 161 (aedeagus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F4E9136FF89FFF6FF68FEC05704E8F2" blockId="14.[151,756,335,414]" pageId="14" pageNumber="179">
<treatmentCitation id="AE50B727FF89FFF6FF68FEC0543AE8ED" author="Jeannel R." box="[151,515,364,385]" page="409" pageId="14" pageNumber="179" year="1962">
<taxonomicName id="E8F1EAB5FF89FFF6FF68FEC0543AE8ED" ID-CoL="977S" authority="Jeannel, 1962: 409" authorityName="Jeannel" authorityPageNumber="409" authorityYear="1962" box="[151,515,364,385]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="foveifrons">
<emphasis id="1D854D24FF89FFF6FF68FEC0577AE8ED" box="[151,323,364,385]" italics="true" pageId="14" pageNumber="179">Achilia foveifrons</emphasis>
<bibRefCitation id="4B60ECC7FF89FFF6FEB5FEC0543AE8ED" author="Jeannel R." box="[330,515,364,385]" pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 409</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 414 fig. 165 (aedeagus)
<emphasis id="1D854D24FF89FFF6FF1CFE25570FE8F2" bold="true" box="[227,310,393,414]" pageId="14" pageNumber="179">syn. nov</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FF68FE1B54A7EB91" pageId="14" pageNumber="179" type="materials_examined">
<paragraph id="2F4E9136FF89FFF6FF68FE1B5476EBA2" blockId="14.[151,757,439,718]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FF68FE1B579EE8A2" bold="true" box="[151,423,439,462]" pageId="14" pageNumber="179">
Type material (
<specimenCount id="39F75ABFFF89FFF6FEA8FE1B57A0E8A2" box="[343,409,439,462]" pageId="14" pageNumber="179" type="generic">8 ex.</specimenCount>
):
</emphasis>
<materialsCitation id="9F999B6BFF89FFF6FE49FE1B5798EB42" ID-GBIF-Occurrence="3391021323" collectingDate="1954-09-20" collectionCode="MHNS, MNHN" collectorName="G. Kuschel" country="Chile" county="Llanquihue prov." latitude="-36.9" location="Frutillar" municipality="Frutillar" pageId="14" pageNumber="179" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Los Lagos" typeStatus="holotype">
CENTRAL
<collectingCountry id="57E6D1A6FF89FFF6FDBFFE1B54ADE8A2" box="[576,660,439,462]" name="Chile" pageId="14" pageNumber="179">CHILE</collectingCountry>
:
<collectingRegion id="ED355FD4FF89FFF6FD59FE1B572BE882" country="Chile" name="Los Lagos" pageId="14" pageNumber="179">Región Los Lagos</collectingRegion>
:
<collectingCounty id="C62FE9BAFF89FFF6FEDEFE7B57DAE882" box="[289,483,471,494]" pageId="14" pageNumber="179">Llanquihue prov.</collectingCounty>
:
<collectionCode id="49E009F3FF89FFF6FE0DFE7B547AE882" box="[498,579,471,494]" pageId="14" pageNumber="179">MHNS</collectionCode>
;
<specimenCount id="39F75ABFFF89FFF6FDACFE7B54B8E881" box="[595,641,471,494]" pageId="14" pageNumber="179" type="male">1 ♂</specimenCount>
(
<typeStatus id="F04A2F94FF89FFF6FD6BFE7B54CDE882" box="[660,756,471,494]" pageId="14" pageNumber="179" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E8F1EAB5FF89FFF6FF45FE5A57B9EB62" authorityName="Jeannel" authorityYear="1962" box="[186,384,502,526]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="foveifrons">
<emphasis id="1D854D24FF89FFF6FF45FE5A57B9EB62" box="[186,384,502,526]" italics="true" pageId="14" pageNumber="179">Achilia foveifrons</emphasis>
</taxonomicName>
n° 1777);
<collectingMunicipality id="CF2A0B4CFF89FFF6FDFAFE5B5458EB62" box="[517,609,503,526]" pageId="14" pageNumber="179">Frutillar</collectingMunicipality>
;
<date id="5B4FB7F6FF89FFF6FD8DFE5B54C9EB62" box="[626,752,503,526]" pageId="14" pageNumber="179" value="1954-09-20">
<collectingDate id="4B0B4E1EFF89FFF6FD8DFE5B54C9EB62" box="[626,752,503,526]" pageId="14" pageNumber="179" value="1954-09-20">20.IX.1954</collectingDate>
</date>
;
<collectorName id="8204F4E0FF89FFF6FF68FDBB5726EB42" box="[151,287,535,558]" pageId="14" pageNumber="179">G. Kuschel.</collectorName>
–
<collectionCode id="49E009F3FF89FFF6FEB7FDBB57A4EB42" box="[328,413,535,558]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="179">MNHN</collectionCode>
;
</materialsCitation>
<materialsCitation id="9F999B6BFF89FFF6FE50FDBB54E4EB02" ID-GBIF-Occurrence="3391021319" collectingDate="1954-09-20" collectionCode="MHNS, MNHN" collectorName="G. Kuschel" country="Chile" county="Llanquihue prov." latitude="-36.9" location="Frutillar" municipality="Frutillar" pageId="14" pageNumber="179" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Los Lagos" typeStatus="paratype">
<specimenCount id="39F75ABFFF89FFF6FE50FDBB57E6EB41" box="[431,479,535,558]" pageId="14" pageNumber="179" type="male">1 ♂</specimenCount>
and
<specimenCount id="39F75ABFFF89FFF6FDDDFDBB5468EB41" box="[546,593,535,558]" pageId="14" pageNumber="179" type="female">1 ♀</specimenCount>
(
<typeStatus id="F04A2F94FF89FFF6FD99FDBB54F6EB42" box="[614,719,535,558]" pageId="14" pageNumber="179" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="E8F1EAB5FF89FFF6FF68FD9A5759EB22" authorityName="Jeannel" authorityYear="1962" box="[151,352,566,590]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="foveifrons">
<emphasis id="1D854D24FF89FFF6FF68FD9A5759EB22" box="[151,352,566,590]" italics="true" pageId="14" pageNumber="179">Achilia foveifrons</emphasis>
</taxonomicName>
);
<location id="2A2EC7EDFF89FFF6FE7EFD9B57E4EB22" LSID="urn:lsid:plazi:treatment:A7582020FF89FFF6FEE4FEAD52EBEC0A:2A2EC7EDFF89FFF6FE7EFD9B57E4EB22" box="[385,477,567,590]" country="Chile" county="Llanquihue prov." latitude="-36.9" municipality="Frutillar" name="Frutillar" pageId="14" pageNumber="179" stateProvince="Los Lagos">Frutillar</location>
; 41° 08’S;
<date id="5B4FB7F6FF89FFF6FD8EFD9B54D6EB22" box="[625,751,567,590]" pageId="14" pageNumber="179" value="1954-09-20">
<collectingDate id="4B0B4E1EFF89FFF6FD8EFD9B54D6EB22" box="[625,751,567,590]" pageId="14" pageNumber="179" value="1954-09-20">20.IX.1954</collectingDate>
</date>
;
<collectorName id="8204F4E0FF89FFF6FF68FDFB5723EB02" box="[151,282,599,622]" pageId="14" pageNumber="179">G. Kuschel.</collectorName>
–
<collectorName id="8204F4E0FF89FFF6FEC6FDFB57D3EB02" box="[313,490,599,622]" pageId="14" pageNumber="179">Región Bío Bío</collectorName>
:
<location id="2A2EC7EDFF89FFF6FE08FDFB5445EB02" LSID="urn:lsid:plazi:treatment:A7582020FF89FFF6FEE4FEAD52EBEC0A:2A2EC7EDFF89FFF6FE08FDFB5445EB02" box="[503,636,599,622]" country="Chile" county="Llanquihue prov." latitude="-36.9" municipality="Frutillar" name="Nuble prov." pageId="14" pageNumber="179" stateProvince="Los Lagos">Ñuble prov.</location>
:
<collectionCode id="49E009F3FF89FFF6FD76FDFB54E0EB02" box="[649,729,599,622]" pageId="14" pageNumber="179">MHNS</collectionCode>
;
</materialsCitation>
<materialsCitation id="9F999B6BFF89FFF6FD18FDFB5472EBA2" ID-GBIF-Occurrence="3391021325" collectingDate="1954-09-20" collectionCode="MHNS, MNHN" collectorName="P. Germain" country="Chile" county="Llanquihue prov." latitude="-36.9" location="Frutillar" municipality="Frutillar" pageId="14" pageNumber="179" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Los Lagos" typeStatus="holotype">
<specimenCount id="39F75ABFFF89FFF6FD18FDFB5692EBE1" pageId="14" pageNumber="179" type="male">1 ♂</specimenCount>
(
<typeStatus id="F04A2F94FF89FFF6FF41FDDB5727EBE2" box="[190,286,631,654]" pageId="14" pageNumber="179" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E8F1EAB5FF89FFF6FEB4FDDA57C7EBE2" authorityName="Jeannel" authorityYear="1962" box="[331,510,630,654]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FEB4FDDA57C7EBE2" box="[331,510,630,654]" italics="true" pageId="14" pageNumber="179">Achilia frontalis</emphasis>
</taxonomicName>
n° 1703);
<collectorName id="8204F4E0FF89FFF6FD80FDDB54EBEBE2" box="[639,722,631,654]" pageId="14" pageNumber="179">Chillàn</collectorName>
;
<collectorName id="8204F4E0FF89FFF6FD1EFDDB56C5EBC2" pageId="14" pageNumber="179">P. Germain.</collectorName>
–
<collectionCode id="49E009F3FF89FFF6FEE5FD3B5756EBC2" box="[282,367,663,686]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="179">MNHN</collectionCode>
;
<specimenCount id="39F75ABFFF89FFF6FE84FD3B579FEBC1" box="[379,422,663,686]" pageId="14" pageNumber="179" type="male">1 ♂</specimenCount>
and
<specimenCount id="39F75ABFFF89FFF6FE22FD3B543EEBC1" box="[477,519,663,686]" pageId="14" pageNumber="179" type="female">3 ♀</specimenCount>
(
<typeStatus id="F04A2F94FF89FFF6FDE8FD3B5446EBC2" box="[535,639,663,686]" pageId="14" pageNumber="179" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="E8F1EAB5FF89FFF6FD58FD3A56C9EBA2" authorityName="Jeannel" authorityYear="1962" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FD58FD3A56C9EBA2" italics="true" pageId="14" pageNumber="179">Achilia frontalis</emphasis>
</taxonomicName>
);
<collectorName id="8204F4E0FF89FFF6FEF7FD1B5763EBA2" box="[264,346,695,718]" pageId="14" pageNumber="179">Chillàn</collectorName>
; 36° 54’S;
<collectorName id="8204F4E0FF89FFF6FE2EFD1B5472EBA2" box="[465,587,695,718]" pageId="14" pageNumber="179">P. Germain</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="2F4E9136FF89FFF6FF68FD4A54A7EB91" blockId="14.[151,670,742,765]" box="[151,670,742,765]" pageId="14" pageNumber="179">
<heading id="7406265AFF89FFF6FF68FD4A54A7EB91" box="[151,670,742,765]" fontSize="10" level="3" pageId="14" pageNumber="179" reason="3">
<emphasis id="1D854D24FF89FFF6FF68FD4A57D1EB91" bold="true" box="[151,488,742,765]" pageId="14" pageNumber="179">
Additional material (
<specimenCount id="39F75ABFFF89FFF6FE79FD4A57E3EB91" box="[390,474,742,765]" pageId="14" pageNumber="179" type="generic">150 ex.</specimenCount>
):
</emphasis>
See Appendix 1.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FF68FCB9531CEBE4" pageId="14" pageNumber="179" type="description">
<paragraph id="2F4E9136FF89FFF6FF68FCB95404ED00" blockId="14.[151,756,789,2060]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FF68FCB9571EEA40" bold="true" box="[151,295,789,812]" pageId="14" pageNumber="179">Description:</emphasis>
Body
<quantity id="E8093CD3FF89FFF6FE76FCB95409EA40" box="[393,560,789,812]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.55" metricValueMin="1.35" pageId="14" pageNumber="179" unit="mm" value="1.45" valueMax="1.55" valueMin="1.35">1.35-1.55 mm</quantity>
long, generally bicolored with head, pronotum and abdomen blackish, elytra, antennae, and legs reddish, and palpi yellowish, or entirely reddish with darker abdomen, sometimes also entirely reddish with darker head. Head with eyes slightly longer than temples. Pronotum slightly wider than head, with maximal width on anterior half; median antebasal fovea as large as lateral ones. First abdominal tergite with basal striae slightly diverging, extending to about one-third of paratergal length, and separated at base by about one-third of tergal width.
</paragraph>
<paragraph id="2F4E9136FF89FFF6FF68FBD852A3E855" blockId="14.[151,756,789,2060]" lastBlockId="14.[831,1437,258,473]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FF68FBD856F7EDE0" box="[151,206,1140,1164]" italics="true" pageId="14" pageNumber="179">Male</emphasis>
: Head as in
<figureCitation id="B7CA8DB3FF89FFF6FE9AFBD957D9EDE0" box="[357,480,1141,1164]" captionStart="Figs 71-76" captionStartId="15.[151,192,1998,2019]" captionTargetBox="[151,1431,257,1958]" captionTargetId="figure-10@15.[151,1431,257,1958]" captionTargetPageId="15" captionText="Figs 71-76. Achilia frontalis. (71, 73, 75) specimen from Chillàn. (72, 74, 76) specimen from Frutillar. Male heads in dorsal (71-72) and lateral (73-74) views. (75-76) male supraocular area in lateral view. Scale bar left for (71, 73) and right for (72, 74)." figureDoi="http://doi.org/10.5281/zenodo.5589984" httpUri="https://zenodo.org/record/5589984/files/figure.png" pageId="14" pageNumber="179">Figs 71-74</figureCitation>
; on each side with deep lateral pit with outer edge forming small tooth at level of eyes (
<figureCitation id="B7CA8DB3FF89FFF6FF0DFB195778EDA0" box="[242,321,1205,1228]" captionStart="Figs 71-76" captionStartId="15.[151,192,1998,2019]" captionTargetBox="[151,1431,257,1958]" captionTargetId="figure-10@15.[151,1431,257,1958]" captionTargetPageId="15" captionText="Figs 71-76. Achilia frontalis. (71, 73, 75) specimen from Chillàn. (72, 74, 76) specimen from Frutillar. Male heads in dorsal (71-72) and lateral (73-74) views. (75-76) male supraocular area in lateral view. Scale bar left for (71, 73) and right for (72, 74)." figureDoi="http://doi.org/10.5281/zenodo.5589984" httpUri="https://zenodo.org/record/5589984/files/figure.png" pageId="14" pageNumber="179">Fig. 75</figureCitation>
), the latter occasionally absent (
<figureCitation id="B7CA8DB3FF89FFF6FD68FB1954DEEDA0" box="[663,743,1205,1228]" captionStart="Figs 71-76" captionStartId="15.[151,192,1998,2019]" captionTargetBox="[151,1431,257,1958]" captionTargetId="figure-10@15.[151,1431,257,1958]" captionTargetPageId="15" captionText="Figs 71-76. Achilia frontalis. (71, 73, 75) specimen from Chillàn. (72, 74, 76) specimen from Frutillar. Male heads in dorsal (71-72) and lateral (73-74) views. (75-76) male supraocular area in lateral view. Scale bar left for (71, 73) and right for (72, 74)." figureDoi="http://doi.org/10.5281/zenodo.5589984" httpUri="https://zenodo.org/record/5589984/files/figure.png" pageId="14" pageNumber="179">Fig. 76</figureCitation>
); frontal protuberance very swollen at base and anteriorly flattened to vertexal sulcus. Antennae (
<figureCitation id="B7CA8DB3FF89FFF6FDA5FB595496EC60" box="[602,687,1269,1292]" captionStart="Figs 30-34" captionStartId="5.[151,192,1388,1409]" captionTargetBox="[189,1397,257,1335]" captionTargetId="figure-284@5.[148,1409,254,1359]" captionTargetPageId="5" captionText="Figs 30-34. Male antennae of Achilia. (30) A. caracolana. (31) A. auriculata. (32) A. ovallensis. (33) A. frontalis. (34) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589966" httpUri="https://zenodo.org/record/5589966/files/figure.png" pageId="14" pageNumber="179">Fig. 33</figureCitation>
) with scape and pedicel longer than wide; antennomeres III- VIII slightly transverse; antennomeres VII-VIII with mesal margin protruding; antennomeres IX-X strongly transverse, with mesal margin protruding; antennomere X wider than IX; antennomere XI elongate, longer than VII-X combined, bearing long and thin subbasal seta inserted in deep depression. Metasternum with ovoidal median impression, posterior edge of the impression pubescent. Legs with trochanters simple; all femora (
<figureCitation id="B7CA8DB3FF89FFF6FEE8F99957B6EF20" box="[279,399,1589,1612]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="14" pageNumber="179">Figs 38-40</figureCitation>
) strongly thickened and sinuate, particularly metafemora; protibiae (
<figureCitation id="B7CA8DB3FF89FFF6FDD8F9F95445EF00" box="[551,636,1621,1644]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="14" pageNumber="179">Fig. 45</figureCitation>
) distinctly thickened on distal half, bearing small subapical tooth; mesotibiae (
<figureCitation id="B7CA8DB3FF89FFF6FEDDF9395740EFC0" box="[290,377,1685,1708]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="14" pageNumber="179">Fig. 50</figureCitation>
) at middle slightly bulging and bearing tuft of short setae; medial edge of metatibiae (
<figureCitation id="B7CA8DB3FF89FFF6FF60F97956D7EF80" box="[159,238,1749,1772]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="14" pageNumber="179">Fig. 37</figureCitation>
) densely pubescent on apical half and distinctly narrowed subapically. Abdominal tergites unmodified; first abdominal ventrite impressed at middle, remaining ventrites flattened at middle; ventrite I with median carina extending from posterior margin to posterior edge of median impression. Aedeagus (
<figureCitation id="B7CA8DB3FF89FFF6FE19F8D9540DEEE0" box="[486,564,1909,1932]" captionStart="Figs 24-29" captionStartId="4.[151,192,1401,1422]" captionTargetBox="[216,1403,263,1340]" captionTargetId="figure-245@4.[162,1413,254,1361]" captionTargetPageId="4" captionText="Figs 24-29. Aedeagi of Achilia. (24) A.caracolana, specimen from Hualpèn, prov. Concepción. (25) A. auriculata, specimen from Peniquillo, prov. Concepción. (26) A. ovallensis, specimen from Ovalle, prov. Limarí [paratype]. (27) A. frontalis, specimen from Aguas Calientes to Puyehue National Park, prov. Osorno. (28) A. trauco, specimen from 34 km WNW La Union, prov. Ranco [paratype]. (29) A. longispina, specimen from Mina de la Disputada, prov. Valparaíso." figureDoi="http://doi.org/10.5281/zenodo.5589962" httpUri="https://zenodo.org/record/5589962/files/figure.png" pageId="14" pageNumber="179">Fig. 27</figureCitation>
)
<quantity id="E8093CD3FF89FFF6FDBCF8D954CAEEE0" box="[579,755,1909,1932]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" metricValueMax="3.35" metricValueMin="3.25" pageId="14" pageNumber="179" unit="mm" value="0.33" valueMax="0.335" valueMin="0.325">
<geoCoordinate id="4AC5F7F1FF89FFF6FDBCF8D954B9EEE0" box="[579,640,1909,1932]" degrees="0.325" direction="north" orientation="latitude" pageId="14" pageNumber="179" precision="55" value="0.325">0.325</geoCoordinate>
<geoCoordinate id="4AC5F7F1FF89FFF6FD7FF8D954FAEEE0" box="[640,707,1909,1932]" degrees="0.335" direction="west" orientation="longitude" pageId="14" pageNumber="179" precision="55" value="-0.335">-0.335</geoCoordinate>
mm
</quantity>
long; dorsal plate ovoid with dorsal longitudinal struts divergent; copulatory pieces consisting of pair of long medial recurved sclerites strongly sclerotised basally and apically pointed. Parameres wide with long seta on poorly developed outer lobe; tips recurved posteriorly and bearing wide subapical seta.
</paragraph>
<paragraph id="2F4E9136FF89FFF6FCC0FEED55F9E8B5" blockId="14.[831,1437,258,473]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FCC0FEED55B6E835" box="[831,911,321,345]" italics="true" pageId="14" pageNumber="179">Female</emphasis>
: Similar to male except: head with occipital protuberance less developed and lacking lateral pits; antennomere XI shorter than VII-X combined; disc of pronotum less convex; metasternum, abdomen, and legs unmodified.
</paragraph>
<paragraph id="2F4E9136FF89FFF6FCC0FE5D531CEBE4" blockId="14.[831,1436,497,648]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FCC0FE5D55C3EB64" bold="true" box="[831,1018,497,520]" pageId="14" pageNumber="179">Collecting data:</emphasis>
Collected from November to March, mainly in mixed forests at elevations ranging from
<quantity id="E8093CD3FF89FFF6FCC0FD9D55B3EB24" box="[831,906,561,584]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="179" unit="m" value="100.0">100 m</quantity>
to
<specimenCount id="39F75ABFFF89FFF6FC46FD9D5232EB24" box="[953,1035,561,584]" pageId="14" pageNumber="179" type="generic">
<quantity id="E8093CD3FF89FFF6FC46FD9D523FEB24" box="[953,1030,561,584]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="179" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
Most specimens came from sifted samples of leaf and log litter, but some were also collected by flight intercept and carrion traps.
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FCC0FD0C5595EB9B" pageId="14" pageNumber="179" type="distribution">
<paragraph id="2F4E9136FF89FFF6FCC0FD0C5595EB9B" blockId="14.[831,1437,672,759]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FCC0FD0C55EDEBDB" bold="true" box="[831,980,672,695]" pageId="14" pageNumber="179">Distribution:</emphasis>
<taxonomicName id="E8F1EAB5FF89FFF6FC21FD0C52B4EBD4" authorityName="Jeannel" authorityYear="1962" box="[990,1165,672,696]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FC21FD0C52B4EBD4" box="[990,1165,672,696]" italics="true" pageId="14" pageNumber="179">Achilia frontalis</emphasis>
</taxonomicName>
is distributed in Central
<collectingCountry id="57E6D1A6FF89FFF6FCC0FD6C5540EBBB" box="[831,889,704,727]" name="Chile" pageId="14" pageNumber="179">Chile</collectingCountry>
from Chiloé to Ñuble Provinces (
<figureCitation id="B7CA8DB3FF89FFF6FAF7FD6C5367EBBB" box="[1288,1374,704,727]" captionStart="Fig" captionStartId="17.[831,866,1968,1989]" captionTargetBox="[880,1355,257,1924]" captionTargetId="figure-12@17.[913,1355,257,1924]" captionTargetPageId="17" captionText="Fig. 84. Distribution map. (● red circles) Achilia testacea. (▲ blue triangles) A. frontalis. (◆ green diamonds) A. caracolana. (■ pink squares) A. trauco. (✷ yellow stars) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589993" httpUri="https://zenodo.org/record/5589993/files/figure.png" pageId="14" pageNumber="179">Fig. 84</figureCitation>
: blue triangles).
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF89FFF6FCC0FCA352EBEC0A" pageId="14" pageNumber="179" type="discussion">
<paragraph id="2F4E9136FF89FFF6FCC0FCA352EBEC0A" blockId="14.[831,1437,783,1383]" pageId="14" pageNumber="179">
<emphasis id="1D854D24FF89FFF6FCC0FCA355FDEA4A" bold="true" box="[831,964,783,806]" pageId="14" pageNumber="179">Comments:</emphasis>
According to
<bibRefCitation id="4B60ECC7FF89FFF6FB98FCA35330EA4A" author="Jeannel R." box="[1127,1289,783,806]" pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962)</bibRefCitation>
the males of
<taxonomicName id="E8F1EAB5FF89FFF6FCC0FC8355F2EA2B" authorityName="Jeannel" authorityYear="1962" box="[831,971,815,839]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="foveifrons">
<emphasis id="1D854D24FF89FFF6FCC0FC8355F2EA2B" box="[831,971,815,839]" italics="true" pageId="14" pageNumber="179">A. foveifrons</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="E8F1EAB5FF89FFF6FB40FC835302EA2B" authorityName="Jeannel" authorityYear="1962" box="[1215,1339,815,839]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FB40FC835302EA2B" box="[1215,1339,815,839]" italics="true" pageId="14" pageNumber="179">A. frontalis</emphasis>
</taxonomicName>
by their short and wide head for the anterior region (while being subtriangular in
<taxonomicName id="E8F1EAB5FF89FFF6FBC6FCC3528AEAEB" authorityName="Jeannel" authorityYear="1962" box="[1081,1203,879,903]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FBC6FCC3528AEAEB" box="[1081,1203,879,903]" italics="true" pageId="14" pageNumber="179">A. frontalis</emphasis>
</taxonomicName>
), the anterior part of the occipital protuberance occupying almost the entire width of the frons (while with a comparatively narrower vertex for
<taxonomicName id="E8F1EAB5FF89FFF6FC4EFC635210EA8B" authorityName="Jeannel" authorityYear="1962" box="[945,1065,975,999]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FC4EFC635210EA8B" box="[945,1065,975,999]" italics="true" pageId="14" pageNumber="179">A. frontalis</emphasis>
</taxonomicName>
), and the aedeagus with divergent dorsal longitudinal struts and thickened and sinuate copulatory pieces (while for
<taxonomicName id="E8F1EAB5FF89FFF6FB66FBA35322ED4B" authorityName="Jeannel" authorityYear="1962" box="[1177,1307,1039,1063]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FB66FBA35322ED4B" box="[1177,1307,1039,1063]" italics="true" pageId="14" pageNumber="179">A. frontalis</emphasis>
</taxonomicName>
the dorsal longitudinal struts are attached to each other, and the copulatory pieces are thin and not sinuate). However the aedeagal conformation of their
<typeStatus id="F04A2F94FF89FFF6FB32FBC3533FEDEA" box="[1229,1286,1135,1158]" pageId="14" pageNumber="179">types</typeStatus>
appear to be very similar, and after examination of extensive material it appears that these differences (when really present) have been strongly exaggerated by Jeannel and fall, in our opinion, within the intraspecific variation of this taxon. Therefore we conclude that
<taxonomicName id="E8F1EAB5FF89FFF6FB49FAA3554CEC2A" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="foveifrons">
<emphasis id="1D854D24FF89FFF6FB49FAA35306EC4B" box="[1206,1343,1295,1319]" italics="true" pageId="14" pageNumber="179">A. foveifrons</emphasis>
<bibRefCitation id="4B60ECC7FF89FFF6FAB9FAA3554CEC2A" author="Jeannel R." pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</taxonomicName>
must be considered to be the junior synonym of
<taxonomicName id="E8F1EAB5FF89FFF6FCC0FAE35276EC0A" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" box="[831,1103,1359,1383]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="179" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="1D854D24FF89FFF6FCC0FAE3558FEC0B" box="[831,950,1359,1383]" italics="true" pageId="14" pageNumber="179">A. frontalis</emphasis>
<bibRefCitation id="4B60ECC7FF89FFF6FC42FAE35276EC0A" author="Jeannel R." box="[957,1103,1359,1382]" pageId="14" pageNumber="179" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</taxonomicName>
(
<emphasis id="1D854D24FF89FFF6FBA0FAE352FBEC0A" bold="true" box="[1119,1218,1359,1382]" pageId="14" pageNumber="179">syn. nov.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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