1128 lines
109 KiB
XML
1128 lines
109 KiB
XML
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<mods:title id="D768D8D7C0505274EAA6DB981050786C">The missing piece of the puzzle: A new and widespread species of the genus Rhynchocalamus Günther, 1864 (Squamata, Colubridae) from the Arabian Peninsula</mods:title>
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|
||
<figureCitation id="AC0F48692C204A229DCA37823BF41E48" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype (RCU-URN - 93850, sample code FLI 447, bottom left) and paratype (RCU-URN - 94064, sample code FLI 330, bottom right) specimens in life. Lateral and ventral views of their heads (above each life picture). Photo credit: FL." figureDoi="10.3897/zse.100.123441.figure3" httpUri="https://binary.pensoft.net/fig/1059094">3</figureCitation>
|
||
; Table
|
||
<tableCitation id="7738F49FA029BCB26961FDE55F906818" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm) of the type series of Rhynchocalamus hejazicus sp. nov. For abbreviations see the Methods section.">1</tableCitation>
|
||
).
|
||
<specimenCount id="B94298B54D07D59CB45E12AE660270C1" type="adult">Adult male</specimenCount>
|
||
;
|
||
<collectingCountry id="80A55343B3B4A4E84C440C047363010F" name="Saudi Arabia">Kingdom of Saudi Arabia</collectingCountry>
|
||
,
|
||
<collectingRegion id="EBBB9B3639EB1C115E0E0A161CF6A681" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
,
|
||
<location id="39F2DC98C114561F9231E10341EE157F" LSID="urn:lsid:plazi:treatment:7B8E986F6C7D5857A65AE8D79456F576:39F2DC98C114561F9231E10341EE157F" country="Saudi Arabia" latitude="26.85895" longLatPrecision="1" longitude="38.30387" municipality="Adult" name="Shaaran Nature Reserve" stateProvince="Medina Province">Shaaran Nature Reserve</location>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EOVAE" specific-use="{"type":"Point","coordinates":[38.303870,26.858950]}">
|
||
<geoCoordinate id="075A7BF564147E927CF32E0D782BEC61" degrees="26.85895" direction="north" orientation="latitude" precision="1" value="26.85895">26.85895 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="24E8096A7071B5BFBFE9D2848C88AC00" degrees="38.30387" direction="east" orientation="longitude" precision="1" value="38.30387">38.30387 ° E</geoCoordinate>
|
||
</named-content>
|
||
),
|
||
<elevation id="894275B55C8C0822E1F230B2CA106EE6" metricMagnitude="3" metricUnit="m" metricValue="1.035" unit="m" value="1035.0">
|
||
<quantity id="81A92AFD4538DAEACA011FB95CD2B48C" metricMagnitude="3" metricUnit="m" metricValue="1.035" unit="m" value="1035.0">1035 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.; collected by
|
||
<collectorName id="D8C6B112A633A12F8121B16EB365B485">Fulvio Licata</collectorName>
|
||
on
|
||
<date id="2DBAC205C759C22CB14936BE516C9128" value="2023-05-24">
|
||
<collectingDate id="7254CEF8A011558021F14F9CBB72D0DB" value="2023-05-24">May 24, 2023</collectingDate>
|
||
</date>
|
||
. MorphoBank accessions:
|
||
<morphoBankNumber id="859B5670B85A3298C3CF23ED03B40822">
|
||
M
|
||
<ext-link id="B9CBE5D13803A9ED419EA1EDE683B088" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902248" xlink_type="simple">902248</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="147D422585FAD0739ACD7A2D95BABDEE">
|
||
M
|
||
<ext-link id="2260693D611D42D5398B8BF997B12BB1" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902255" xlink_type="simple">902255</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="0956B4D33B9A9DD9CD7BC472B4068212">
|
||
<materialsCitation id="4AAA05984510C4E9DE7B3BFA5BBB293C" collectingDate="2023-05-11" collectorName="Fulvio Licata & Leili Khalatbari" country="Saudi Arabia" elevation="1616" latitude="25.60676" location="Harrat Khaybar" longLatPrecision="1" longitude="39.96734" municipality="Harrat Khaybar" specimenCode="RCU-URN - 94064" specimenCount="1" stateProvince="Medina Province" typeStatus="paratype">
|
||
<emphasis id="7F17EAC4D1C54934EBE08BA422871D49" bold="true">
|
||
<emphasis id="10B10BE95468A5BAAE619E0E05BEDE2F" bold="true" italics="true">
|
||
<typeStatus id="2E1CA1C5C5FC76197FB3572DA179812C" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="0A41F4EDF320884A5D5A08DD2CEF7D23">
|
||
<abbrev content-type="institution" id="ABBRID0EEWAE" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 94064
|
||
</specimenCode>
|
||
(sample code FLI 330; Fig.
|
||
<figureCitation id="5D7FA3B8D739AFD843E50DFE76673718" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype (RCU-URN - 93850, sample code FLI 447, bottom left) and paratype (RCU-URN - 94064, sample code FLI 330, bottom right) specimens in life. Lateral and ventral views of their heads (above each life picture). Photo credit: FL." figureDoi="10.3897/zse.100.123441.figure3" httpUri="https://binary.pensoft.net/fig/1059094">3</figureCitation>
|
||
; Table
|
||
<tableCitation id="62DC92DE2BBDD59B7B543F77355E47B5" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm) of the type series of Rhynchocalamus hejazicus sp. nov. For abbreviations see the Methods section.">1</tableCitation>
|
||
), melanistic
|
||
<specimenCount id="B5D74F8F32F75A2AD10C3843E67B1655" type="adult">adult female</specimenCount>
|
||
collected by
|
||
<collectorName id="AD2DAF49CEDB66D03512373D11EE68C4">Fulvio Licata</collectorName>
|
||
and
|
||
<collectorName id="2A6868B399E6D2B645AABE30B4917572">Leili Khalatbari</collectorName>
|
||
on
|
||
<date id="99401321F7D4321BD99AA2A0A6E08C3F" value="2023-05-11">
|
||
<collectingDate id="B45086DC0F769F6B19D3C692E3A505FA" value="2023-05-11">May 11, 2023</collectingDate>
|
||
</date>
|
||
,
|
||
<abbrev id="ABBRID0ERWAE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="0A45FC2D86B1456493EC40D68169FE99" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
,
|
||
<collectingRegion id="4378EB4A6846A155E62442F085EAB6A7" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="4445726F9917632C99839C1E484A04C7">Harrat Khaybar</collectingMunicipality>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EYWAE" specific-use="{"type":"Point","coordinates":[39.967340,25.606760]}">
|
||
<geoCoordinate id="F61C235D7C7C67D1B178BB9314077312" degrees="25.60676" direction="north" orientation="latitude" precision="1" value="25.60676">25.60676 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="FB378E72ED1C5A7F4B145DC90D4B6076" degrees="39.96734" direction="east" orientation="longitude" precision="1" value="39.96734">39.96734 ° E</geoCoordinate>
|
||
</named-content>
|
||
), elevation
|
||
<quantity id="05623A8FA03FBC13450109F259361AA8" metricMagnitude="3" metricUnit="m" metricValue="1.616" unit="m" value="1616.0">
|
||
<elevation id="A4C0CF6857DB1B36EBBC301383684090" metricMagnitude="3" metricUnit="m" metricValue="1.616" unit="m" value="1616.0">1616 m</elevation>
|
||
</quantity>
|
||
asl, MorphoBank accessions:
|
||
<morphoBankNumber id="2C366D1D211706B21E4D5B427A7B3B3A">
|
||
M
|
||
<ext-link id="C82273170A70CFF32CA56B56DD28FCF7" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902243" xlink_type="simple">902243</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="CD9BE78FF8A7055A720E276A40D6FB27">
|
||
M
|
||
<ext-link id="C5090CDB060E078FDA4C130AC151BC91" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902247" xlink_type="simple">902247</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="BBA6B0A6C28D3E589481E7B8D2ECDF42" collectingDate="2023-11-02" collectorName="Jose Carlos Brito" country="Saudi Arabia" elevation="1277" latitude="26.36263" location="Jabal Al Ward" longLatPrecision="1" longitude="37.3878" specimenCode="RCU-URN - 94065" specimenCount="1" stateProvince="Medina Province">
|
||
<specimenCode id="D768DEA9C149285B48B11C9A1A1EFE2A">
|
||
<abbrev content-type="institution" id="ABBRID0EHXAE" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 94065
|
||
</specimenCode>
|
||
(sample code JCB 222),
|
||
<specimenCount id="568A2162557CD7EE2DAAC200A96850BE" type="juvenile">juvenile male</specimenCount>
|
||
collected by
|
||
<collectorName id="1C066F6CDAB6FE0F6F4FBB21C513BE9B">José Carlos Brito</collectorName>
|
||
on
|
||
<date id="641FE1B01F94C60C5A214EC816F5AA82" value="2023-11-02">
|
||
<collectingDate id="1692539ADFF4C12D1523786C7C00AA2A" value="2023-11-02">November 2, 2023</collectingDate>
|
||
</date>
|
||
,
|
||
<abbrev id="ABBRID0EMXAE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="EF18C7D3D029B7D95C82A1A5610B4B5E" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
,
|
||
<collectingRegion id="69BCD471BF53F4B9200789F0D8586DE0" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
,
|
||
<location id="D39BF85773BB208DB6284142DA704F61" LSID="urn:lsid:plazi:treatment:7B8E986F6C7D5857A65AE8D79456F576:D39BF85773BB208DB6284142DA704F61" country="Saudi Arabia" latitude="26.36263" longLatPrecision="1" longitude="37.3878" name="Jabal Al Ward" stateProvince="Medina Province">Jabal Al Ward</location>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETXAE" specific-use="{"type":"Point","coordinates":[37.387800,26.362630]}">
|
||
<geoCoordinate id="80FF15394CBFC4B4CF5C41CB5FA626F4" degrees="26.36263" direction="north" orientation="latitude" precision="1" value="26.36263">26.36263 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ABE4BBBE8E455BAFF587CA152F559CD7" degrees="37.38780" direction="east" orientation="longitude" precision="1" value="37.3878">37.38780 ° E</geoCoordinate>
|
||
</named-content>
|
||
), elevation
|
||
<quantity id="EC2A3463972504B025347D105D80CA52" metricMagnitude="3" metricUnit="m" metricValue="1.277" unit="m" value="1277.0">
|
||
<elevation id="7CF247C8007C2A8B0AB65597A6B275BE" metricMagnitude="3" metricUnit="m" metricValue="1.277" unit="m" value="1277.0">1277 m</elevation>
|
||
</quantity>
|
||
asl, MorphoBank accessions:
|
||
<morphoBankNumber id="06F823E4459B5AEA60DE5400332D0504">
|
||
M
|
||
<ext-link id="4759E4C024DCC7D80BE75CA9F8AFF642" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902256" xlink_type="simple">902256</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="DB8FBCD3D924A6F6700384C04249EC49">
|
||
M
|
||
<ext-link id="2D70C60D9D71B40093210C8DB82F6718" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902261" xlink_type="simple">902261</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="7807147E17DB468E34553D2F85A978F4" collectingDate="2022-04-22" collectionCode="NMP" collectorName="Damien M. Egan & Lukas Pola" country="Saudi Arabia" elevation="781" latitude="26.602814" location="Al Qurr village" longLatPrecision="1" longitude="37.122513" municipality="Prince Mohammed bin Salman Royal Reserve" specimenCode="NMP 76815" specimenCount="1" stateProvince="Tabuk / Medina Provinces">
|
||
<specimenCode id="DE4DC1369EAACF6B2456FAB3166ECBFB">
|
||
<named-content id="86426C81F9C7A2F260F8BC9154945CBA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-1" xlink_title="National Museum (Prague)">NMP</named-content>
|
||
76815
|
||
</specimenCode>
|
||
(sample code LP 760; Fig.
|
||
<figureCitation id="6FF34C915123711C10391E28A8FA4E2D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Colour variation within R. hejazicus sp. nov. (or its lack, thereof). Top row: two unvouchered specimens from NEOM, Tabuk Province, KSA (photo credit: Euan Ferguson and Neil Rowntree); bottom left: paratype NMP 76815 (sample code LP 760, photo credit: DME); bottom right: paratype MNHN – RA – 2023.0013 (sample code JIR 544, photo credit: AAI)." figureDoi="10.3897/zse.100.123441.figure4" httpUri="https://binary.pensoft.net/fig/1059095">4</figureCitation>
|
||
; Table
|
||
<tableCitation id="F3F100E7DCE02457B2C131B55A27F7D8" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm) of the type series of Rhynchocalamus hejazicus sp. nov. For abbreviations see the Methods section.">1</tableCitation>
|
||
),
|
||
<specimenCount id="6C25149D37460618CCE7107B4CA5B03D" type="juvenile">juvenile male</specimenCount>
|
||
collected by
|
||
<collectorName id="A1850ECE41E940BFD88813E460F59862">Damien M. Egan</collectorName>
|
||
and
|
||
<collectorName id="FDCE327E50FAA5F515B41FF284C9F0CC">Lukáš Pola</collectorName>
|
||
on
|
||
<date id="8312A597A7F2EA6D8808E83223CCB24E" value="2022-04-22">
|
||
<collectingDate id="9B48B181EBFD5C5B2785916918DD8568" value="2022-04-22">April 22, 2022</collectingDate>
|
||
</date>
|
||
,
|
||
<abbrev id="ABBRID0EPYAE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="0D8654E40B2144CDB12A1D9E7FF61F42" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
,
|
||
<collectingRegion id="5CF73EA1E7459FD8DF1B9B006C932A80">Tabuk / Medina Provinces</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="5CD4E8A761BBA5E0D3B6BBAD0D16072F">Prince Mohammed bin Salman Royal Reserve</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="8DB26AFEE02763B15604868CAC5CEB13">
|
||
<quantity id="A669106263F279C78DDF8A5A881EA69F" metricMagnitude="4" metricUnit="m" metricValue="2.6" unit="km" value="26.0">26 km</quantity>
|
||
east of
|
||
</locationDeviation>
|
||
<location id="3FEF735193D30341F2B6890F8B43DA89" LSID="urn:lsid:plazi:treatment:7B8E986F6C7D5857A65AE8D79456F576:3FEF735193D30341F2B6890F8B43DA89" country="Saudi Arabia" latitude="26.602814" longLatPrecision="1" longitude="37.122513" municipality="Prince Mohammed bin Salman Royal Reserve" name="Al Qurr village" stateProvince="Tabuk / Medina Provinces">Al Qurr village</location>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWYAE" specific-use="{"type":"Point","coordinates":[37.122515,26.602814]}">
|
||
<geoCoordinate id="204444464C9C765AF877E8135ECEE759" degrees="26.6028142" direction="north" orientation="latitude" precision="1" value="26.602814">26.6028142 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="DBDFDAF77B006BBDE74709D84353CA1E" degrees="37.1225145" direction="east" orientation="longitude" precision="1" value="37.122513">37.1225145 ° E</geoCoordinate>
|
||
</named-content>
|
||
), elevation
|
||
<quantity id="060727F86CB8ACA7C23B8A172E460AC2" metricMagnitude="2" metricUnit="m" metricValue="7.81" unit="m" value="781.0">
|
||
<elevation id="CF3FE338698D8F9EB08BAAE1B31EB113" metricMagnitude="2" metricUnit="m" metricValue="7.81" unit="m" value="781.0">781 m</elevation>
|
||
</quantity>
|
||
asl, MorphoBank accessions:
|
||
<morphoBankNumber id="DA73DAF5330484FBC907B113436F3EB7">
|
||
M
|
||
<ext-link id="541E6F1A4F8D83B13015CF212A38FBAE" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902293" xlink_type="simple">902293</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="6BB547BF27E1247A217DB2A6E6426D6F">
|
||
M
|
||
<ext-link id="52C04360D921AC64E55C4FD022DF2D2F" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902338" xlink_type="simple">902338</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="5F4C7DDE7E7DEAA35F589160053C7223" collectingDate="2021-07" collectionCode="NMP" collectorName="Ahmed Mohajja AlShammari & Mubarak Al - Aslami" country="Saudi Arabia" elevation="1052" latitude="27.165277" location="An Niayy village" longLatPrecision="1" longitude="42.294445" municipality="Jabal Salma" specimenCode="NMP 76816" specimenCount="1" specimenCount-adult="1" stateProvince="Hail Province">
|
||
<specimenCode id="B2D9F36264C967401BEFC42E8C6E153D">
|
||
<named-content id="C185BF828CFF1CA6ED38891366243E40" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-1" xlink_title="National Museum (Prague)">NMP</named-content>
|
||
76816
|
||
</specimenCode>
|
||
(sample code JIR 551),
|
||
<specimenCount id="E5861496B03049A851B22A2F8C21B86E" type="adult">adult</specimenCount>
|
||
, sex undetermined, collected by
|
||
<collectorName id="2A8D6549F42C42633C433CCDB2A490A8">Ahmed Mohajja AlShammari</collectorName>
|
||
and
|
||
<collectorName id="25204B8B478741080469DBA8B493726A">Mubarak Al – Aslami</collectorName>
|
||
in
|
||
<date id="B1879A692EF536E76A6AC7E9EC971B52" value="2021-07">
|
||
<collectingDate id="A7EB79B43C5C7B55D3ED3DB50580DC3F" value="2021-07">July 2021</collectingDate>
|
||
</date>
|
||
,
|
||
<abbrev id="ABBRID0EKZAE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="7D382BCD2890A7C172DB0587D93C2E18" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
,
|
||
<collectingRegion id="924F7DB233096886454A66939543F77D" country="Saudi Arabia" name="Ha'il">Hail Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="AA60D04ACB452AD8D6D9832F8E99CDEB">Jabal Salma</collectingMunicipality>
|
||
, south of
|
||
<location id="5ED0365F9E1F4C1058E8225F76BB8DD1" LSID="urn:lsid:plazi:treatment:7B8E986F6C7D5857A65AE8D79456F576:5ED0365F9E1F4C1058E8225F76BB8DD1" country="Saudi Arabia" latitude="27.165277" longLatPrecision="1" longitude="42.294445" municipality="Jabal Salma" name="An Niayy village" stateProvince="Hail Province">An Niayy village</location>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERZAE" specific-use="{"type":"Point","coordinates":[42.294444,27.165278]}">
|
||
<geoCoordinate id="1335BB66D01A933AC61B1A9A806624CA" degrees="27.165278" direction="north" orientation="latitude" precision="1" value="27.165277">27.165278 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2988FDD6EEBD2BFC2BD788CCF3BB61F5" degrees="42.294444" direction="east" orientation="longitude" precision="1" value="42.294445">42.294444 ° E</geoCoordinate>
|
||
</named-content>
|
||
), elevation
|
||
<quantity id="9446B3243AA7C6B4968EF26220F47510" metricMagnitude="3" metricUnit="m" metricValue="1.052" unit="m" value="1052.0">
|
||
<elevation id="09460710584F5BCCA9E16A0B7DC8E6C2" metricMagnitude="3" metricUnit="m" metricValue="1.052" unit="m" value="1052.0">1052 m</elevation>
|
||
</quantity>
|
||
asl, MorphoBank accessions:
|
||
<morphoBankNumber id="FE13E4A5A85B9EBF2C7B68BD052C6170">
|
||
M
|
||
<ext-link id="5CEF9C3A06785194D1FAB0646063E214" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902290" xlink_type="simple">902290</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="BBB7BEA16A7F3447796D7526DF278258">
|
||
M
|
||
<ext-link id="4881149932E862C6BED9AC30484774D2" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902292" xlink_type="simple">902292</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="21248BD6F50120CB1A46D728B02A1EED" collectingDate="2017-05-11" collectionCode="MNHN" collectorName="Adel A. Ibrahim & Ahmed Mohajja AlShammari" country="Saudi Arabia" county="Jabal Salma" elevation="1099" latitude="26.948889" location="Wadi Al - Azraq" longLatPrecision="1" longitude="41.97389" municipality="Zekheen village" specimenCode="MNHN - RA - 2023.0013" specimenCount="1" stateProvince="Hail Province">
|
||
<specimenCode id="4746C0D05DD84EB2778CB39848F0C6DE">
|
||
<named-content id="A49FAE0382487DAA15A83400D088F8BB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-historia-natural-0" xlink_title="Museo Nacional de Historia Natural">MNHN</named-content>
|
||
– RA – 2023.0013
|
||
</specimenCode>
|
||
(sample code JIR 544; Fig.
|
||
<figureCitation id="6DD164CBF47466838CF01033D84718A2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Colour variation within R. hejazicus sp. nov. (or its lack, thereof). Top row: two unvouchered specimens from NEOM, Tabuk Province, KSA (photo credit: Euan Ferguson and Neil Rowntree); bottom left: paratype NMP 76815 (sample code LP 760, photo credit: DME); bottom right: paratype MNHN – RA – 2023.0013 (sample code JIR 544, photo credit: AAI)." figureDoi="10.3897/zse.100.123441.figure4" httpUri="https://binary.pensoft.net/fig/1059095">4</figureCitation>
|
||
; Table
|
||
<tableCitation id="85E431C6567B9388C6E663DAC7E3F649" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Measurements (in mm) of the type series of Rhynchocalamus hejazicus sp. nov. For abbreviations see the Methods section.">1</tableCitation>
|
||
)
|
||
<specimenCount id="516F92F8FA47F6CD16DBC07AEC0124C5" type="adult">adult female</specimenCount>
|
||
collected by
|
||
<collectorName id="40ADD5555B92C193CF1720CF0710BC94">Adel A. Ibrahim</collectorName>
|
||
and
|
||
<collectorName id="31BF2B38FCF8547B6E5FCC9306F34B03">Ahmed Mohajja AlShammari</collectorName>
|
||
on
|
||
<date id="EAE9F39595FC91739FF88473C0ED72B0" value="2017-05-11">
|
||
<collectingDate id="25C5528EAAE44FF78D2126DF398410E7" value="2017-05-11">May 11, 2017</collectingDate>
|
||
</date>
|
||
,
|
||
<abbrev id="ABBRID0EN1AE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="E60757D926CBBD5655CE2886DD9EA1E4" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
,
|
||
<collectingRegion id="C6A5194FD30738496B5CE1AAFE0FB3B8" country="Saudi Arabia" name="Ha'il">Hail Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="F074CB4D94E9419D185A0D3C55B3BAD6">Jabal Salma</collectingCounty>
|
||
,
|
||
<location id="C4CB30FF9CB93D697DDE491FBB30B713" LSID="urn:lsid:plazi:treatment:7B8E986F6C7D5857A65AE8D79456F576:C4CB30FF9CB93D697DDE491FBB30B713" country="Saudi Arabia" county="Jabal Salma" latitude="26.948889" longLatPrecision="1" longitude="41.97389" municipality="Zekheen village" name="Wadi Al - Azraq" stateProvince="Hail Province">Wadi Al – Azraq</location>
|
||
, east of
|
||
<collectingMunicipality id="7E208107B90165828C4F445E0E58E074">Zekheen village</collectingMunicipality>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EU1AE" specific-use="{"type":"Point","coordinates":[41.973889,26.948889]}">
|
||
<geoCoordinate id="E2D3EA5FC086D6C91D56BE80209296FD" degrees="26.948889" direction="north" orientation="latitude" precision="1" value="26.948889">26.948889 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5D21AFFAFDC82630D14172632CEB2F14" degrees="41.973889" direction="east" orientation="longitude" precision="1" value="41.97389">41.973889 ° E</geoCoordinate>
|
||
</named-content>
|
||
), elevation
|
||
<quantity id="B889B95F3CE71C8C93EC3158A0DD2718" metricMagnitude="3" metricUnit="m" metricValue="1.099" unit="m" value="1099.0">
|
||
<elevation id="7D00EF7819797BE955593AE4810A52B9" metricMagnitude="3" metricUnit="m" metricValue="1.099" unit="m" value="1099.0">1099 m</elevation>
|
||
</quantity>
|
||
asl, MorphoBank accessions:
|
||
<morphoBankNumber id="B31A440F48259A01473D1AD267E965BA">
|
||
M
|
||
<ext-link id="9EFDE1B9E43A09201ED2D840B96CAF0D" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902262" xlink_type="simple">902262</ext-link>
|
||
</morphoBankNumber>
|
||
–
|
||
<morphoBankNumber id="105D17C6718CED6D8675DC0E09AA2A4A">
|
||
M
|
||
<ext-link id="A890483369851EF8A928DD67D997596C" ext-link-type="uri" xlink_href="http://www.morphbank.net/?id=902289" xlink_type="simple">902289</ext-link>
|
||
</morphoBankNumber>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ED2AE" type="material">
|
||
<paragraph id="726CE8F235FB21FD921BB41666C2DC44">
|
||
<heading id="FAF23EA3F4B8454EE7A48CE31F5185E0" reason="title">Other material.</heading>
|
||
</paragraph>
|
||
<paragraph id="95D383E31A67E3A12F56F411AA56EDDD">
|
||
Additional unvouchered individuals, assigned to this species based on phenotypical resemblance and unique head pattern, were encountered in May, June, and
|
||
<date id="79E788A475456AFF7869F21D094480EC" value="2023-11">November 2023</date>
|
||
by Vidak Lakušić, Gholam Hosein Yusefi, and Fulvio Licata in Harrat Uwayrid, Al-Gharameel, Wadi Nakhlah, and Sharaan (conservation areas located in the AlUla County,
|
||
<collectingRegion id="509EE6675A39D95DB4EB7A230A412793" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
). Other individuals were observed in
|
||
<date id="DAC2A277F8196B8973F9BD853A229E87" value="2023-07" valueMax="2023-08-31" valueMin="2023-07-00">July and August 2023</date>
|
||
by Neil Rowntree and Euan Ferguson in
|
||
<collectionCode id="3C01B0006CD3D2038B0391B826106F7D">NEOM</collectionCode>
|
||
, the northwestern region of
|
||
<collectingRegion id="0DF75BD84296D60049A585317FE9A03A" country="Saudi Arabia" name="Tabuk">Tabuk Province</collectingRegion>
|
||
,
|
||
<abbrev id="ABBRID0EJ2AE" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="81020F8DF3BF4DA08C5CF6F13118BAE1" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
. One record was found in the public database iNaturalist (
|
||
<ext-link id="36A6809FCBB58799268FCAD1B340BF71" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/20014261" xlink_type="simple">https://www.inaturalist.org/observations/20014261</ext-link>
|
||
). Based on the provided information, the snake was recorded on
|
||
<date id="2EEE8DE89E95ED6878ED05D7C54D1E5F" value="2019-02-01">February 1, 2019</date>
|
||
at As Salam,
|
||
<collectingRegion id="4B70CDF9649C0E9B7FFA6FFC572EE90B" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
(
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EV2AE" specific-use="{"type":"Point","coordinates":[39.537669,24.464058]}">
|
||
<geoCoordinate id="8996243FB7F73FC5DD41E1C7F5D12F14" degrees="24.464058" direction="north" orientation="latitude" precision="1" value="24.464058">24.464058 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7854F4B720A49291325913311E098734" degrees="39.537669" direction="east" orientation="longitude" precision="1" value="39.53767">39.537669 ° E</geoCoordinate>
|
||
</named-content>
|
||
). An additional individual was reported to us by Muteb Masad Al – Malki from the vicinity of Adham city (
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E42AE" specific-use="{"type":"Point","coordinates":[40.879264,20.448653]}">
|
||
<geoCoordinate id="58819B81385F77087CA4206055BEC5E3" degrees="20.4486525" direction="north" orientation="latitude" precision="1" value="20.448652">20.4486525 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BCBD7025E19EFB48859112599055C5A4" degrees="40.8792636" direction="east" orientation="longitude" precision="1" value="40.879265">40.8792636 ° E</geoCoordinate>
|
||
</named-content>
|
||
) from
|
||
<collectingRegion id="B6A5D1B301507DAB823FB70792D23386" country="Saudi Arabia" name="Makkah">Mecca Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EC3AE" type="etymology">
|
||
<paragraph id="795A91962D353ECFDED8C1A710AFF191">
|
||
<heading id="B3EE3624B772250CF20360568FCE62A1" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="107F230DC3FF55613C8F4E0008DD14AE">
|
||
The species name is a latinized noun in masculine gender derived from the word "
|
||
<emphasis id="FC7070D8DFAE027156EFF893F62C6986" italics="true">Hejaz</emphasis>
|
||
– " = Hejaz Mountains, a mountain range located in the Hejaz region (an important region located in western
|
||
<collectingCountry id="31395960154C8771EE0B569DA6DBB1BA" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
, where the two holy cities of Islam,
|
||
<collectingRegion id="D46116ED8CD85A3F3E94272B13B49862" country="Saudi Arabia" name="Makkah">Mecca</collectingRegion>
|
||
, and
|
||
<collectingRegion id="3939BBD7202CC6B4CBD1A5F6ABDEFF16" country="Saudi Arabia" name="Al Madinah">Medina</collectingRegion>
|
||
are located) where most individuals were observed, and the Latin suffix " –
|
||
<emphasis id="38B51EE6D5000B35298BD7CCE6C8E964" italics="true">icus</emphasis>
|
||
" = “ belonging to ”. We suggest the common name “ Hejaz black – collared snake ” in English and أبو حناء [Abu Henna] in Arabic for the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ED4AE" type="diagnosis">
|
||
<paragraph id="6CD1781F542A0617D864E3240FB0BB7F">
|
||
<heading id="B773B9230706C9F854070E43B4D64108" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="27EFCFE0432FF6568B9F1C943C78557D">
|
||
The new species of
|
||
<taxonomicName id="6049CE36EDF61FAE2E7B387E327E9DD8" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="genus">
|
||
<emphasis id="6AC82DE0030C208935DA091D10843D7B" italics="true">Rhynchocalamus</emphasis>
|
||
</taxonomicName>
|
||
from the Hejaz Mountain range in western
|
||
<collectingCountry id="D53011B7959511923AAD420AE8577350" name="Saudi Arabia">Saudi Arabia</collectingCountry>
|
||
is characterised by the following morphological characters: (1)
|
||
<abbrev id="ABBRID0EW4AE" xlink_title="snout-vent length">SVL</abbrev>
|
||
<quantity id="5A29434E9CB376D99ED215513476051B" metricMagnitude="-1" metricUnit="m" metricValue="2.7435" metricValueMax="3.395" metricValueMin="2.0919999999999996" unit="mm" value="274.35" valueMax="339.5" valueMin="209.2">209.2–339.5 mm</quantity>
|
||
in adults; (2) tail length
|
||
<quantity id="5D060A59BB4BFC2DE40C48BE4BB97B3F" metricMagnitude="-2" metricUnit="m" metricValue="5.125" metricValueMax="6.42" metricValueMin="3.8299999999999996" unit="mm" value="51.25" valueMax="64.2" valueMin="38.3">38.3–64.2 mm</quantity>
|
||
in adults; (3) loreal scale present; (4) large 3
|
||
<superScript id="663136FBFAB3C6BE1892185D3E8B1689">rd</superScript>
|
||
and 4
|
||
<superScript id="C65BFD80FB847D5135853A14C2E8F591">th</superScript>
|
||
upper labial scales in contact with the eye; (5) one preocular scale; (6) 1–2 postocular scales; (7) one temporal scale; (8) 0–2 post – temporal scales; (9) six upper labial scales; (10) eight lower labial scales; (11) usually four lower labial scales in contact with the anterior inframaxillars; (12) usually one gular scale in contact with anterior inframaxillars, situated between the posterior inframaxillars; (13) 15 smooth dorsal scales at mid-body; (14) 11–12 dorsal and temporal scales surrounding the margin of parietals; (15) 227–250 ventrals; (16) anal and subcaudal scales divided; (17) 67–70 subcaudal scales; (18) dorsal colouration in life deep reddish with a distinctive black collar extending behind the parietal scales and abruptly stopping or tapering backward in the middle, and a pale reddish band passing behind the eyes, through the middle of the supraoculars and the frontal scale, encompassing the temporal and parietal scales.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E54AE" type="Colouration in life (adults)">
|
||
<paragraph id="09458236F848ECF6A79015EC01F49B60">
|
||
<heading id="BBCDF3C67094CDE0292CCE5C77DACD4F" reason="title">Colouration in life (adults).</heading>
|
||
</paragraph>
|
||
<paragraph id="726D3A505BE3BAA155D0439B0D601785">The upper surface of the head is shiny black from the middle of the supraoculars and the frontal scale to the tip of the snout, which is whitish; a wide band, pale reddish dorsally and fading to whitish ventrally, passes behind the eyes, through the middle of the supraoculars and the frontal scale, encompassing the temporal and parietal scales; a black collar around the neck reaches the ventrals, and abruptly stops or tapers dorsally towards the centre; dorsal surface of the body and tail uniformly deep reddish from the end of the collar to the tail tip; ventral surface of the body deep reddish fading to whitish in the upper part of the body, narrowing in correspondence of the black collar; colour pattern paler in alcohol preserved specimen. A melanistic morph, uniformly black, also occurs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ED5AE" type="comparisons">
|
||
<paragraph id="C4DF1C2B5BFB7C9F9BF16F55687CED7E">
|
||
<heading id="68A8E53213A99FB6F9A283058A8305E9" reason="title">Comparison.</heading>
|
||
</paragraph>
|
||
<paragraph id="41B359CD98D7F91164DC330DF0A2508B">
|
||
<taxonomicName id="8DE2986ABE1532EA4949EEB158E7E042" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="12E86B0175E4D638899E71149C130087" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="17C34D1AFBD219A3EDB99693817BE629" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is morphologically similar to the other
|
||
<taxonomicName id="B5EAEE8F68A2E393644DF93EA89AD73A">
|
||
<emphasis id="704BA8D4E029968C4D765FD4621CE768" italics="true">Rhynchocalamus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, and it can be distinguished by slight differences in size, colouration, and head and body scalation.
|
||
</paragraph>
|
||
<paragraph id="02927B1523D65C463D015706F757353D">
|
||
In comparison with
|
||
<taxonomicName id="A6D78E24FD9D80095C16801BFA23E903" authorityName="Schmidt" authorityYear="1933" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="86A17B21CFE7F81F25FCFE657C8D1D94" italics="true">R. arabicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="86FB06DE7E834BD755A363CBD98B4A54" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="858CD0361309A00ED815652F90E9267B" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="5C26CB9742B0EFAF87CD040190C8B597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a lower number of subcaudal scales (67–70 vs.
|
||
<quantity id="550A3DFD381D4B4377B2D99A7D97A457" metricMagnitude="0" metricUnit="m" metricValue="1.9304" metricValueMax="2.0574" metricValueMin="1.8034" unit="in" value="76.0" valueMax="81.0" valueMin="71.0">71–81 in</quantity>
|
||
<taxonomicName id="3F9FB3D098A3BDCDA8F0FBAA3A35F6B2" authorityName="Schmidt" authorityYear="1933" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="D20568CE55BD4F9DAA5087B5FAF041DD" italics="true">R. arabicus</emphasis>
|
||
</taxonomicName>
|
||
). We show that a melanistic morph of
|
||
<taxonomicName id="C8E483FDCC7118C6F96CF41EAD2E6ABF" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="B893060EE064E2065C163B04A5B4271A" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DA837C3555ACEB62EAF3391688EEFA97" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also occurs, therefore the new species can be easily misidentified as
|
||
<taxonomicName id="1DF691EAD548C9BD6F9EAEC24E0FFEE1" authorityName="Schmidt" authorityYear="1933" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="arabicus">
|
||
<emphasis id="21D76D5E031C2B4F4184FE569673269C" italics="true">R. arabicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B8F788962567742A60A95B631F7A4295">
|
||
<taxonomicName id="A95B44798A28F4BC1A551552976292BA" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="4CAED8C50B5BE718CEB2055B31D3E086" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CFEBA00D2C897937AF195F88BC84D27C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="A0420AE88A94D15D04544ED035FC0B57" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="13C41B0D0B06C646482D84998B681ADB" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
by smaller maximum body size (339.5 vs.
|
||
<quantity id="264AC2E0051B529AD368F916507B8B5C" metricMagnitude="1" metricUnit="m" metricValue="1.097534" unit="in" value="432.1">432.1 in</quantity>
|
||
<taxonomicName id="77EA83F737C7B42484F491BF34B319BB" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="502443FCCDCF4C37436101D4F9B6ADE5" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
), a shorter tail (38.3–64.2 vs.
|
||
<quantity id="D07EA3620B6A8E4108F1890C0AA340FA" metricMagnitude="0" metricUnit="m" metricValue="1.9469100000000004" metricValueMax="2.3901399999999997" metricValueMin="1.5036800000000003" unit="in" value="76.65" valueMax="94.1" valueMin="59.2">59.2–94.1 in</quantity>
|
||
<taxonomicName id="615645596A0279B905D55095E8C5779A" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="AB54798C8D3B86154BE369ACB3FB082C" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
), by a higher number of ventrals (227–250 vs.
|
||
<quantity id="8506BB2B83666D92418DEEF3049AEB1B" metricMagnitude="0" metricUnit="m" metricValue="5.4229" metricValueMax="5.816599999999999" metricValueMin="5.0292" unit="in" value="213.5" valueMax="229.0" valueMin="198.0">198–229 in</quantity>
|
||
<taxonomicName id="36C664D8ADC9ACE0804E99DC6F20D213" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="34165A564E7C593065CFBCB7C28CF03D" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
) and subcaudals (67–70 vs.
|
||
<quantity id="01F61D4D4310C54696E517E9A65C5705" metricMagnitude="0" metricUnit="m" metricValue="1.4731999999999998" metricValueMax="1.5748" metricValueMin="1.3716" unit="in" value="58.0" valueMax="62.0" valueMin="54.0">54–62 in</quantity>
|
||
<taxonomicName id="67E6CA173878CFF96D701846B9AAE908" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="F2FEF7A40A73855437628437137D3C98" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
), Lastly,
|
||
<taxonomicName id="ECAB518B2556CF6C63BE036D4A8AE6CA" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="7F9F655B525A2E7E45FB8BB77FE44DB8" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0CCC55F880DAF6BA287104F3B9E98125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be further differentiated from
|
||
<taxonomicName id="D5655E5CA672059DC75E27588BFE99BF" authorityName="Tamar, Smid, Gocmen, Meiri & Carranza" authorityYear="2016" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="dayanae">
|
||
<emphasis id="6EA50D093FEF1AB0F93C7ABCC865CC89" italics="true">R. dayanae</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a pale reddish band passing between the eyes and the neck.
|
||
</paragraph>
|
||
<paragraph id="81378D59B81C9CB57E2C0B58728CBF03">
|
||
<taxonomicName id="DAF0F69AE44018D0AE70AFC02300C4B6" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="B4962913B87769638A063C65422ED821" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="46E37918B334DB770CD2BE5D52FA272A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="8108F530AC714D433487EDD05CADD187" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="79704A193EC0D413DF083734BC48FB8E" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
(both Southern population from the Negev region in
|
||
<collectingCountry id="7AFFEF0D72CC2AF917C4AB7872E7C223" name="Israel">Israel</collectingCountry>
|
||
and Northern population from the Mediterranean ecoregion;
|
||
<emphasis id="EEB414CA569B567B39C372C7FB378696" italics="true">sensu</emphasis>
|
||
<bibRefCitation id="8B54502815EDD4341F931DA9AE9A3F7B" DOI="10.1080/09397140.2020.1757914" author="Tamar" etAl="et al." firstAuthor="Tamar" issue="2" journalOrPublisher="Zoology in the Middle East" pagination="118-123" refId="B56" refString="Tamar K, Wiedl HJ, Maza E, Jablonski D, Meiri S (2020) Discovery of the Black-headed Ground Snake Rhynchocalamus melanocephalus (Jan, 1862) in Cyprus (Reptilia: Colubridae). Zoology in the Middle East 66 (2): 118–123. https://doi.org/10.1080/09397140.2020.1757914" title="Discovery of the Black-headed Ground Snake Rhynchocalamus melanocephalus (Jan, 1862) in Cyprus (Reptilia: Colubridae)." volume="66" year="2020">Tamar et al. 2020</bibRefCitation>
|
||
) by a smaller maximum body size (339.5 vs.
|
||
<quantity id="8003D57A51AD2C6E7C85EADF47D4E6C2" metricMagnitude="1" metricUnit="m" metricValue="1.267968" unit="in" value="499.2">499.2 in</quantity>
|
||
<taxonomicName id="22DCB596BD90AA6680AF287F5F048B10" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="A62056B29955CFF4E560F72560CECBD8" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
), a lower number of post-temporal scales (1 vs
|
||
<quantity id="6F20769EB13C25AAC6E57BB2E3DF701B" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="06F0BAF9655A43F891409A33DFD260D8" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="9CA0905194D47AC8CF4BD4B6B5D6D856" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
), a higher number of lower labial scales (8 vs.
|
||
<quantity id="D367501DA438DEAAF9C671857F51F045" metricMagnitude="-1" metricUnit="m" metricValue="1.778" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="6F1656FC2658554ED6E4979D1B4D38AF" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="D0AD11E9A1092BDF56AA72B5128EFD6D" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
), a higher number of ventrals (227–250 vs.
|
||
<quantity id="E29D16419E00A81E1F45FB34D9EF5250" metricMagnitude="0" metricUnit="m" metricValue="5.0673" metricValueMax="5.968999999999999" metricValueMin="4.1655999999999995" unit="in" value="199.5" valueMax="235.0" valueMin="164.0">164–235 in</quantity>
|
||
<taxonomicName id="346B34FBE97F1EB2AEF14C461FF96FEF" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="C1C19A1441BE920185061D85D7854EC8" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
), and subcaudals (67–70 vs.
|
||
<quantity id="3B9B7C0BDAB5FD6CA5DBD9BF804C923C" metricMagnitude="0" metricUnit="m" metricValue="1.2446" metricValueMax="1.7526" metricValueMin="0.7365999999999999" unit="in" value="49.0" valueMax="69.0" valueMin="29.0">29–69 in</quantity>
|
||
<taxonomicName id="07DD8AFE847A43CCB4931A836DF7BAAD" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="8FDE2E6BF3B9F9D84AC0AF067C91E962" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
). Lastly,
|
||
<taxonomicName id="283D088D2D42E0630778B1D6E7CCF86E" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="6F47A3FD4CD3FCC7896665E6D468A327" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7553FCCE431B2DE233D40A326AA877E3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be further differentiated from
|
||
<taxonomicName id="643F939FA9B20F6FE85942ADE3574079" baseAuthorityName="Jan" baseAuthorityYear="1862" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="melanocephalus">
|
||
<emphasis id="DDD200813FD35ED739FCE4736A2E75D9" italics="true">R. melanocephalus</emphasis>
|
||
</taxonomicName>
|
||
by a pale reddish band between the eyes and the neck.
|
||
</paragraph>
|
||
<paragraph id="884D5C3F64FE6E8FF5BE873FEE6670EA">
|
||
<taxonomicName id="1802057FC55588EFAF1E31AF6666DA74" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="E49E05816AA35B3E453E7AE6230689EE" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C3C6A5F5B1AFC72EC18301182F7E2C19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="9EA06BD8B04D414C97A97DA631DACE80" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="BC422284D9A798A67376365BC708A232" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
in having a longer tail (64.2 vs.
|
||
<quantity id="A564BD437851AD17458397E4AD7CB2BB" metricMagnitude="-2" metricUnit="m" metricValue="5.4" unit="mm" value="54.0">54 mm</quantity>
|
||
in
|
||
<taxonomicName id="8DCDB412049CC0237F3E519612C4EE41" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="1D24A128C322F1E26051B04EB1B6E412" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
), a higher number of ventrals (227–250 vs.
|
||
<quantity id="AACDFB17C652A9495A06DA5FDD199880" metricMagnitude="0" metricUnit="m" metricValue="5.4229" metricValueMax="5.740399999999999" metricValueMin="5.1054" unit="in" value="213.5" valueMax="226.0" valueMin="201.0">201–226 in</quantity>
|
||
<taxonomicName id="FEF5439DA30C4838B2D5695EC04FC232" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="3F3042AAE59EAD6F6A02FBDD8B817BC2" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
) and subcaudals (67–70 vs.
|
||
<quantity id="018C479EB4D8C9FE095E618C99B2EF96" metricMagnitude="0" metricUnit="m" metricValue="1.4859" metricValueMax="1.6256" metricValueMin="1.3462" unit="in" value="58.5" valueMax="64.0" valueMin="53.0">53–64 in</quantity>
|
||
<taxonomicName id="173E154AB8976244C4A1E92FD45BAC70" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="6F12E05A89766DCDEDFD715F340B439B" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
), and a lower number of upper labials (6 vs.
|
||
<quantity id="B1A3414DDF1EC7CA674B1FF1DA255563" metricMagnitude="-1" metricUnit="m" metricValue="1.778" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="0C878E889B8F8F0D3EDACA8543016C82" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="E3616902B64471EC2B663DEB2B5480D7" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
). Lastly,
|
||
<taxonomicName id="05361B2D5CE3CB5D99FE591877AD5BBE" baseAuthorityName="Nikolsky" baseAuthorityYear="1899" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="satunini">
|
||
<emphasis id="6A9AA36653455117270F15DDF80326F6" italics="true">R. satunini</emphasis>
|
||
</taxonomicName>
|
||
is characterised by two black patches on a pale reddish / whitish background on the prefrontals and the parietals, and a black band around the neck that does not reach the ventrals.
|
||
</paragraph>
|
||
<paragraph id="3ADC7834AA88448E7760060BD7DC4936">
|
||
In comparison with
|
||
<taxonomicName id="C489B95CF1C87CBF56B8B651C4731A2C" baseAuthorityName="Torki" baseAuthorityYear="2017" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="levitoni">
|
||
<emphasis id="A4EB959C9926CDA2CD878DCE647C3824" italics="true">R. levitoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A0E33F6262A31D58FB2EF5BD12B7CB9D" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="12DD346D3E6D1EEE61CBB94099AF973E" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DF9E3D20F74093C422C5862BD1B3E459" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a lower number of upper labials (6 vs.
|
||
<quantity id="8DAB15736554866D24BA3260F8AF9FCD" metricMagnitude="-1" metricUnit="m" metricValue="1.778" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="882DA418695E8DC95A1E6A82AB9AF3CB" baseAuthorityName="Torki" baseAuthorityYear="2017" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="levitoni">
|
||
<emphasis id="674B30449E18B7A5F19A27792B14800C" italics="true">R. levitoni</emphasis>
|
||
</taxonomicName>
|
||
), and it can be distinguished by the overall different colouration (deep reddish vs. lemon yellow in
|
||
<taxonomicName id="81274E7CFAFC736CABA2DBD0D5210D7F" baseAuthorityName="Torki" baseAuthorityYear="2017" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="levitoni">
|
||
<emphasis id="4F693F1396F8633D880B05A1E1CF2002" italics="true">R. levitoni</emphasis>
|
||
</taxonomicName>
|
||
), and the absence of a V-shaped band on the neck, and dark markings on the parietals.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDLAG" type="description">
|
||
<paragraph id="FD04F299A21DBC71C3E8993CD8B956A8">
|
||
<heading id="FE478CA00359F5CF0EA2E4250F9A2F22" reason="title">
|
||
Description of the
|
||
<typeStatus id="D7CEEE27DDDD328BAF2D23E5DE16149E" type="holotype">holotype</typeStatus>
|
||
.
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="1D3D43D2636D53B3A04855FEA3E4F0F0">
|
||
Adult male (voucher code
|
||
<specimenCode id="C45F8759B9A989D93E63C9FB6E82D2F5">
|
||
<abbrev content-type="institution" id="ABBRID0EJLAG" xlink_title="AlUla Museum">
|
||
<collectionCode id="3642080AEB89117605E9DFCA5384410D">RCU-URN</collectionCode>
|
||
</abbrev>
|
||
- 93850
|
||
</specimenCode>
|
||
) (Fig.
|
||
<figureCitation id="4DA44590FC82405F6169A20BCA405BA2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype (RCU-URN - 93850, sample code FLI 447, bottom left) and paratype (RCU-URN - 94064, sample code FLI 330, bottom right) specimens in life. Lateral and ventral views of their heads (above each life picture). Photo credit: FL." figureDoi="10.3897/zse.100.123441.figure3" httpUri="https://binary.pensoft.net/fig/1059094">3</figureCitation>
|
||
). Snout-vent length
|
||
<quantity id="CFD5BCA0D5C4F40146D821428A2CDA73" metricMagnitude="-1" metricUnit="m" metricValue="3.211" unit="mm" value="321.1">321.1 mm</quantity>
|
||
, tail length
|
||
<quantity id="078CB9BAF9768D154709030C9476011D" metricMagnitude="-2" metricUnit="m" metricValue="6.36" unit="mm" value="63.6">63.6 mm</quantity>
|
||
. Body cylindrical and slender. The head is small and relatively narrow, slightly distinct from the neck. Head scales not keeled. Rostral scale elongated, extending backwards and wedged between the internasals, which are separated, and touching the bottom of the suture between the prefrontals. The rostral is bordered by two upper labials, two nasals, two internasals, and two prefrontals. Nostrils are situated on undivided nasal scales internasals of triangular shape, separated by the rostral scale. A trapezoid loreal scale at either side in contact with the 2
|
||
<superScript id="D252913691FEDA5F91A2019CD8A33877">nd</superScript>
|
||
and 3
|
||
<superScript id="096F6D88D7D16E22FD8CC9BDA06A5003">rd</superScript>
|
||
upper labials. The eyes are small, with circular pupils. One preocular scale on both sides. Two postocular scales on both sides. Bell-shaped frontal scale, located between the supraoculars and wedged between the parietals. Six squarish upper labials, the 3
|
||
<superScript id="99D6684D23E0E243CA29EE8CCD2878C9">rd</superScript>
|
||
and 4
|
||
<superScript id="CF25D99B13FA19395A5A951F839021A4">th</superScript>
|
||
in contact with the eye. two large parietal scales; one temporal and one post – temporal scale on the left side, while on the right side, only one enlarged temporal is present. Eight lower labials. Four lower labials are in contact with anterior inframaxillars on each side. Two gular scales are positioned between the posterior inframaxillars and in connection with the anterior inframaxillars. Anterior inframaxillars are almost double in size than posterior inframaxillars. Eleven dorsal and temporal scales surround the posterior margin of the parietals. Fifteen dorsal scale rows at mid-body. 247 ventrals and 67 subcaudals, including a conical scale at the tail tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1LAG" type="distribution">
|
||
<paragraph id="41D49F0487B2A2B7C2B3EDD42779A965">
|
||
<heading id="8D5664738247FADAFD73CBE515DE32A2" reason="title">Known and potential distribution and habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="FF299EC6E6201BDFDB7968C5E7728321">
|
||
The known extent of occurrence (
|
||
<abbrev id="ABBRID0EAMAG" xlink_title="extent of occurrence">EOO</abbrev>
|
||
) of
|
||
<taxonomicName id="D04B974871195C5441BD29DDDF9A0C65" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="BB04E90E3CFE30B8317059AF8CB7C178" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65A210D34196B8ACBEE37B20384DE010" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is more than
|
||
<quantity id="50A04B81C305452145D339BEFA9A5DCB" metricMagnitude="8" metricUnit="m" metricValue="2.74674" unit="km" value="274674.0">274,674 km</quantity>
|
||
<superScript id="FEBBA6BEF2061386176B72259CC6FB83">2</superScript>
|
||
and the area of occurrence (
|
||
<abbrev id="ABBRID0ERMAG" xlink_title="area of occurrence">AOO</abbrev>
|
||
) is
|
||
<quantity id="75566D6FA8327FFB9E451A162D3FC64D" metricMagnitude="4" metricUnit="m" metricValue="5.6" unit="km" value="56.0">56 km</quantity>
|
||
<superScript id="BAD1CFD1A34370C22B621D36B6EC7544">2</superScript>
|
||
. The individual reported from the vicinity of Adham city (
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1MAG" specific-use="{"type":"Point","coordinates":[40.879264,20.448653]}">
|
||
<geoCoordinate id="9D09D6A3CF7CB1E70C0905891AD06786" degrees="20.4486525" direction="north" orientation="latitude" precision="1" value="20.448652">20.4486525 ° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28F515573F88556A39924B826E34C33F" degrees="40.8792636" direction="east" orientation="longitude" precision="1" value="40.879265">40.8792636 ° E</geoCoordinate>
|
||
</named-content>
|
||
) from
|
||
<collectingRegion id="733DB3211C7E62D5B634D46ED57C03D8" country="Saudi Arabia" name="Makkah">Mecca Province</collectingRegion>
|
||
represents the southernmost record to our knowledge.
|
||
</paragraph>
|
||
<paragraph id="BB4B8FD4225968EB5E9BB012057D5591">
|
||
Available material and photographic records suggest that the species is scattered across
|
||
<collectingRegion id="7585E40E88C234CAF0E6BFDB09BDF0BD" country="Saudi Arabia" name="Tabuk">Tabuk</collectingRegion>
|
||
,
|
||
<collectingRegion id="F9106400728E1B55B4FF2B92B97BD522" country="Saudi Arabia" name="Al Madinah">Medina</collectingRegion>
|
||
,
|
||
<collectingRegion id="D123B0D106C746DE458E39935EA3C1FF" country="Saudi Arabia" name="Ha'il">Hail</collectingRegion>
|
||
and
|
||
<collectingRegion id="5C1281C0E6745377BA3929F479B5F3DD" country="Saudi Arabia" name="Makkah">Mecca</collectingRegion>
|
||
Provinces, and is likely endemic to
|
||
<abbrev id="ABBRID0EBNAG" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="1BF0442617E4D9ADFFFCC183DB0DF33D" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
. Nevertheless, it should be noted that additional range extensions of many overlooked species are now being reported with each herpetofaunal survey carried out in the unexplored regions of northwestern and western
|
||
<abbrev id="ABBRID0EFNAG" xlink_title="Kingdom of Saudi Arabia">
|
||
<collectingCountry id="E10BF9639744ABEAA639BBD33C241DA4" name="Saudi Arabia">KSA</collectingCountry>
|
||
</abbrev>
|
||
(e. g.,
|
||
<bibRefCitation id="A38BEBAD4C1BD7F8ED92C9E4C3083C98" author="Aloufi" etAl="et al." firstAuthor="Aloufi" journalOrPublisher="Herpetology Notes" pagination="973-975" refId="B2" refString="Aloufi A, Baker MAA, Amr ZS (2020) First record of the Variable Racer, Platyceps variabilis (Boulenger, 1905), for Saudi Arabia. Herpetology Notes 13: 973–975." title="First record of the Variable Racer, Platyceps variabilis (Boulenger, 1905), for Saudi Arabia." volume="13" year="2020">Aloufi et al. 2020</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="C4FDCA97A1A3E634548BC131BF170C2B" DOI="10.30906/1026-2296-2021-28-3-123-137" author="Aloufi" firstAuthor="Aloufi" issue="3" journalOrPublisher="Russian Journal of Herpetology" pagination="123-137" refId="B3" refString="Aloufi AA, Amr ZS, Baker MA (2021) Reptiles and Amphibians of Al Madinah Al Munawwarah Province, Saudi Arabia. Russian Journal of Herpetology 28 (3): 123–137. https://doi.org/10.30906/1026-2296-2021-28-3-123-137" title="Reptiles and Amphibians of Al Madinah Al Munawwarah Province, Saudi Arabia." volume="28" year="2021">2021</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="CE6033CB5CE9BFAB98D017407DFF217C" author="Aloufi" firstAuthor="Aloufi" journalOrPublisher="Herpetology Notes" pagination="483-491" refId="B4" refString="Aloufi A, Amr Z, Baker MA (2022) Reptiles from ’ Uruq Bani Ma’arid and Harat al Harrah protected areas in Saudi Arabia: Reptiles from two protected areas in Saudi Arabia. Herpetology Notes 15: 483–491." title="Reptiles from ’ Uruq Bani Ma’arid and Harat al Harrah protected areas in Saudi Arabia: Reptiles from two protected areas in Saudi Arabia." volume="15" year="2022">2022</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D5388D67B5559CB3A09053A4FAD098AB" DOI="10.1111/jbi.14823" author="Pola" etAl="et al." firstAuthor="Pola" journalOrPublisher="Journal of Biogeography" pagination="1-15" refId="B44" refString="Pola L, Crochet PA, Geniez P, Shobrak M, Busais S, Jablonski D, Masroor R, Abduraupov T, Carranza S, Šmíd J (2024) Some like it hot: Past and present phylogeography of a desert dwelling gecko across the Arabian Peninsula. Journal of Biogeography 00: 1–15. https://doi.org/10.1111/jbi.14823" title="Some like it hot: Past and present phylogeography of a desert dwelling gecko across the Arabian Peninsula." year="2024">Pola et al. 2024</bibRefCitation>
|
||
), suggesting that the range of this rather secretive species could be even wider. Currently, the northwesternmost individual is reported from the area west of Jabal Al Lawz, approx.
|
||
<quantity id="4D6B3F8E5B78BD51157F2755BED10CBC" metricMagnitude="4" metricUnit="m" metricValue="7.0" unit="km" value="70.0">70 km</quantity>
|
||
south of the Jordanian border. Given the presence of similar habitats harbouring many shared species, we cannot rule out the species occurrence in that area, especially in the mountains of Aqaba.
|
||
</paragraph>
|
||
<paragraph id="D8344DBD8BAFC45023A54B36A22B4402">
|
||
<taxonomicName id="50F4E79128B02EC802DCDE4B76D07438" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="137865096CD263CBDAEF677758396403" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DB65716D35A0BE311C8DCEBE43152BC2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been observed between 456 and
|
||
<quantity id="3B40D6A396A89C7D5A8D1C8E69F03293" metricMagnitude="3" metricUnit="m" metricValue="1.61" unit="m" value="1610.0">1610 m</quantity>
|
||
a. s. l., in the following habitats: (i) sandy flatland with sparse vegetation (
|
||
<taxonomicName id="5EC47F0DE22DE71626C5DE83A759A63A" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="7BE592B3397107047FF9844500C988D4" italics="true">Acacia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, bushes, and tussock grasses; Fig.
|
||
<figureCitation id="BBC70E59B8865AD9EC21C929B8138BF5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitats of the holotype and two paratype specimens of R. hejazicus sp. nov. Top: locality of the holotype RCU-URN - 93850 at the Shaaran NR, AlUla County, Medina Province, KSA (photo credit: FL); bottom left: habitat of the paratype MNHN – RA – 2023.0013 at Wadi Al-Azraq, Jabal Salma, Hail Province, KSA (photo credit: AAI); bottom right: habitat of the paratype RCU-URN - 94065 the Harrat Khaybar, Hail Province, KSA (photo credit: FL)." figureDoi="10.3897/zse.100.123441.figure5" httpUri="https://binary.pensoft.net/fig/1059096">5</figureCitation>
|
||
); (ii) large, stony wadis in lava fields (= harrat) with dense patches of woody vegetation (
|
||
<taxonomicName id="F55967DAA346337C2EA46342DAAE8B6F" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="5650825024F55FB58A4F77E8D405BB53" italics="true">Acacia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
; Fig.
|
||
<figureCitation id="420C3A872F4DB7F5EAFFDC2A19A847FC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitats of the holotype and two paratype specimens of R. hejazicus sp. nov. Top: locality of the holotype RCU-URN - 93850 at the Shaaran NR, AlUla County, Medina Province, KSA (photo credit: FL); bottom left: habitat of the paratype MNHN – RA – 2023.0013 at Wadi Al-Azraq, Jabal Salma, Hail Province, KSA (photo credit: AAI); bottom right: habitat of the paratype RCU-URN - 94065 the Harrat Khaybar, Hail Province, KSA (photo credit: FL)." figureDoi="10.3897/zse.100.123441.figure5" httpUri="https://binary.pensoft.net/fig/1059096">5</figureCitation>
|
||
); (iii) sparsely vegetated rocky creek with temporary pools.
|
||
</paragraph>
|
||
<paragraph id="524D3AC353E8CB4C0A10B7DEA7400DFB">
|
||
The species distribution models achieved good performance levels (AUC: 0.9–0.94;
|
||
<abbrev id="ABBRID0EBPAG" xlink_title="True Skill Statistics">TSS</abbrev>
|
||
: 0.78–0.87; Boyce Index: 0.42–0.99), resulting in an overall good predictive accuracy of the final ensemble models (Boyce index: 0.71). The potential distribution of
|
||
<taxonomicName id="D9AB4DB52D7DC265F60120FBCEA605F1" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="552A0ED25CC5AB11CDD126D806F6A0A8" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E994A22A7887A2683B577CC0AF1F43B5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
mostly includes the known range of the species, but other extralimital suitable areas are identified to the northwest (i. e., the Sinai Mountains in South Sinai,
|
||
<collectingRegion id="FBB36524ECACC65CFFD6127311469BE9" country="Jordan" name="Al 'Aqabah">Aqaba</collectingRegion>
|
||
mountains, and Wadi Rum in
|
||
<collectingCountry id="EF5EBFEAAB423049E5F11B2B912B0A49" name="Jordan">Jordan</collectingCountry>
|
||
), and to the north (i. e., Jabal Tubaiq hills at the border with
|
||
<collectingCountry id="FE533FD47376776A3DA87D3B281A720A" name="Jordan">Jordan</collectingCountry>
|
||
and Upper Galilee mountains in North
|
||
<collectingCountry id="2D5AD2E70D38BF12AA56681BB365C4F0" name="Israel">Israel</collectingCountry>
|
||
; Fig.
|
||
<figureCitation id="AA3C2AE9B2E7F445A5EE9398BB832C03" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Rhynchocalamus hejazicus sp. nov. showing the location of the material examined in this study. Circles indicate samples used for the genetic analyses, squares indicate examined voucher specimens, and diamonds indicate photographic records. The black arrow denotes the type locality (Shaaran NR in AlUla County, Medina Province). Geographical sampling is overlaid by the predicted species distribution model categorised into binary predictions of the suitable areas. The threshold used for the categorisation was the lowest predicted suitability value among the occurrence points (= 0.05)." figureDoi="10.3897/zse.100.123441.figure6" httpUri="https://binary.pensoft.net/fig/1059097">6</figureCitation>
|
||
). The main drivers of environmental suitability in our models were the altitude (26.7 % of explained variance) and the mean temperature of the wettest quarter (
|
||
<collectionCode id="A7CE1EC278F3697F19A4BCEC04E9DB1C" country="Spain" name="University of the Basque Country" type="Herbarium">BIO</collectionCode>
|
||
8; 15.6 %), suggesting that
|
||
<taxonomicName id="9502275F5F1E09BF31BC708468936F84" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="DBE05835493251555DBD277E49795A35" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1C1453FE38E722C6A94E9D4FF4234DCC" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is associated with altitudes above
|
||
<quantity id="0B2064CCE857D9C988657BCA05C53A78" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1,000 m</quantity>
|
||
a. s. l., in regions characterised by cold wet months (<20 ° C). Furthermore, the precipitation of the warmest quarter (
|
||
<collectionCode id="0F2B7F0370FE05DA19F535FDB32B4E1D" country="Spain" name="University of the Basque Country" type="Herbarium">BIO</collectionCode>
|
||
18; 13.5 %) and the precipitation seasonality (
|
||
<collectionCode id="1CEF93893D260CC88E7645329FC06EC8" country="Spain" name="University of the Basque Country" type="Herbarium">BIO</collectionCode>
|
||
15; 13 %) were also related to the probability of occurrence of the species, indicating that the species is likely to occur in areas with relatively dry hot months (precipitations <
|
||
<quantity id="3A20AD1EE37A41171D692570BFA0983D" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="mm" value="50.0">50 mm</quantity>
|
||
) and stable rainfall regimes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E6PAG" type="natural history">
|
||
<paragraph id="6E77E56897FE2EBCB0158AA0CC7F1DF3">
|
||
<heading id="BF6DD9B7A2EDBB60E86253870D6363BF" reason="title">Natural history.</heading>
|
||
</paragraph>
|
||
<paragraph id="59E41EFB17F5272B71960DAA21875480">
|
||
Little is known about the species’ natural history and behaviour. Although the current number of observations is limited, it appears that
|
||
<taxonomicName id="EC7FEF96C37ED8BC5587673BA4F22C24" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="2413B33ADB652EFE5E04E1770A03AF04" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8875611E1BED7804CD9F893867F38F07" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has mainly nocturnal activity as all individuals were encountered active at night. We assume that in many biological aspects, it will be concordant with its congeners (
|
||
<bibRefCitation id="FB21EFE8B917D96EFB2947F393F2EC1D" DOI="10.3897/vz.61.e31150" author="Amr and Disi" firstAuthor="Amr" issue="2" journalOrPublisher="Vertebrate Zoology" pagination="179-266" refId="B7" refString="Amr Z, Disi AM (2011) Systematics, distribution and ecology of the snakes of Jordan. Vertebrate Zoology 61 (2): 179–266. https://doi.org/10.3897/vz.61.e31150" title="Systematics, distribution and ecology of the snakes of Jordan." volume="61" year="2011">Amr and Disi 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5F031CDAC035CF2F8F1D4DF0788980D8" DOI="10.7717/peerj.2769" author="Tamar" etAl="et al." firstAuthor="Tamar" refId="B55" refString="Tamar K, Šmíd J, Göçmen B, Meiri S, Carranza S (2016) An integrative systematic revision and biogeography of Rhynchocalamus snakes (Reptilia, Colubridae) with a description of a new species from Israel. PeerJ 4: e 2769. https://doi.org/10.7717/peerj.2769" year="2016">Tamar et al. 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="041352E96CE1BAB63673A768F795799A" DOI="10.1080/09397140.2017.1299319" author="Torki" firstAuthor="Torki" issue="2" journalOrPublisher="Zoology in the Middle East" pagination="109-116" refId="B59" refString="Torki F (2017) Description of a new species of Lytorhynchus (Squamata: Colubridae) from Iran. Zoology in the Middle East 63 (2): 109–116. https://doi.org/10.1080/09397140.2017.1299319" title="Description of a new species of Lytorhynchus (Squamata: Colubridae) from Iran." volume="63" year="2017">Torki 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="81D44213763F5D6420D311F2C8C0C50B" DOI="10.11646/zootaxa.4282.3.3" author="Fathinia" etAl="et al." firstAuthor="Fathinia" issue="3" journalOrPublisher="Zootaxa" pagination="473-486" refId="B21" refString="Fathinia B, Rastegar-Pouyani E, Rastegar-Pouyani N, Darvishnia H (2017) A new species of the genus Rhynchocalamus Günther, 1864 (Reptilia: Squamata: Colubridae) from Ilam province in western Iran. Zootaxa 4282 (3): 473–486. https://doi.org/10.11646/zootaxa.4282.3.3" title="A new species of the genus Rhynchocalamus Günther, 1864 (Reptilia: Squamata: Colubridae) from Ilam province in western Iran." volume="4282" year="2017">Fathinia et al. 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="405D642E6891B6978206002C1126631D">
|
||
Other reptile species observed in syntopy of the
|
||
<specimenCode id="0B54CF8A7B96F0865AEDD583DACE5B61">
|
||
<named-content id="6E000BCE0A4D68A3AE9F8FA426547803" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-1" xlink_title="National Museum (Prague)">
|
||
<collectionCode id="9C334CB071F738798057DEF44AB3D58E" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" type="">NMP</collectionCode>
|
||
</named-content>
|
||
76815
|
||
</specimenCode>
|
||
(
|
||
<collectionCode id="CA5085E871D8664B42722D45527D6433" country="Russia" name="Laboratory of Palaeontology">LP</collectionCode>
|
||
760) were
|
||
<taxonomicName id="E8B5035BA3290B9D069F83AD14C158C4" baseAuthorityName="Donndorff" baseAuthorityYear="1798" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" isUncertain="true" kingdom="Animalia" phylum="Chordata" rank="species" species="hasselquistii">
|
||
<emphasis id="47845AB81D9813959278A4794915290A" italics="true">Ptyodactylus</emphasis>
|
||
cf.
|
||
<emphasis id="44016D87211A349C62D20EBD0858F75A" italics="true">hasselquistii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="55B68EBEB61A69DEAC304F0934E7D6C6" class="Squamata" family="Phyllodactylidae" kingdom="Animalia" phylum="Chordata" rank="family">Phyllodactylidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="0B78F29D6D1DB5DCA4228C134C865122" authorityName="Haas" authorityYear="1943" class="Squamata" family="Sphaerodactylidae" genus="Pristurus" kingdom="Animalia" phylum="Chordata" rank="species" species="guweirensis">
|
||
<emphasis id="1C8B0B871948EAA4356CE36D6B09580D" italics="true">Pristurus guweirensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="0578DE8CDC710EE6CDC1AEC166561609" class="Squamata" family="Sphaerodactylidae" kingdom="Animalia" phylum="Chordata" rank="family">Sphaerodactylidae</taxonomicName>
|
||
), and
|
||
<taxonomicName id="17D1B2B5F0549D4D218DC864C1AFA790" class="Squamata" family="Viperidae" genus="Echis" kingdom="Animalia" phylum="Chordata" rank="species" species="coloratus">
|
||
<emphasis id="C2C6ED8870A780D11A9EFDD6238471E6" italics="true">Echis coloratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="E044F9709627E352289CD74D80B924BF" authorityName="Oppel" authorityYear="1811" class="Squamata" family="Viperidae" kingdom="Animalia" phylum="Chordata" rank="family">Viperidae</taxonomicName>
|
||
). In
|
||
<collectingRegion id="AF250A3A95057475602B4703D003898B" country="Saudi Arabia" name="Ha'il">Hail Province</collectingRegion>
|
||
, at Jabal Salma range,
|
||
<taxonomicName id="40117EA5662BBD5BDE4A94BCB332A743" class="Squamata" family="Viperidae" genus="Echis" kingdom="Animalia" phylum="Chordata" rank="species" species="coloratus">
|
||
<emphasis id="6488787B5D22C857AEF3520347F1F238" italics="true">Echis coloratus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="BAB0329C8BF6691F2A9C12AB2E0ACABF" authorityName="Leviton & Anderson" authorityYear="1967" class="Squamata" family="Viperidae" genus="Cerastes" kingdom="Animalia" phylum="Chordata" rank="species" species="gasperettii">
|
||
<emphasis id="6063B200DCDF989C8E8B7D8C53D802F3" italics="true">Cerastes gasperettii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="D5130BBD8A1EC81CE79474288E394D72" authorityName="Oppel" authorityYear="1811" class="Squamata" family="Viperidae" kingdom="Animalia" phylum="Chordata" rank="family">Viperidae</taxonomicName>
|
||
) and
|
||
<taxonomicName id="353DD3B123BDF1550EE4CE4DA080DDDD" authorityName="Lataste" authorityYear="1887" class="Squamata" family="Elapidae" genus="Walterinnesia" kingdom="Animalia" phylum="Chordata" rank="species" species="aegyptia">
|
||
<emphasis id="EFCC91A359E229285203F2D4A65143C1" italics="true">Walterinnesia aegyptia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="818470107A4BC0123D6D6C92A598580B" authorityName="F.Boie" authorityYear="1827" class="Squamata" family="Elapidae" kingdom="Animalia" phylum="Chordata" rank="family">Elapidae</taxonomicName>
|
||
) were found in sympatry (
|
||
<bibRefCitation id="6E80E8E444E5AB600B6E53CD49AE1C73" author="Alshammari and Ibrahim" firstAuthor="Alshammari" journalOrPublisher="Herpetological Conservation and Biology" pagination="1021-1029" refId="B5" refString="Alshammari AM, Ibrahim AA (2015) Lizards and snakes in the historical Faid protected area (Faid Hema), Ha’il region, Saudi Arabia. Herpetological Conservation and Biology 10: 1021–1029." title="Lizards and snakes in the historical Faid protected area (Faid Hema), Ha’il region, Saudi Arabia." volume="10" year="2015">Alshammari and Ibrahim 2015</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D4A99AE50788F512C0CA76853906E861" author="Alshammari" etAl="et al." firstAuthor="Alshammari" journalOrPublisher="Herpetozoa (Wien)" pagination="59-63" refId="B6" refString="Alshammari AM, Busais SM, Ibrahim AA (2017) Snakes in the Province of Ha’il, Kingdom of Saudi Arabia, including two new records. Herpetozoa (Wien) 30: 59–63." title="Snakes in the Province of Ha’il, Kingdom of Saudi Arabia, including two new records." volume="30" year="2017">Alshammari et al. 2017</bibRefCitation>
|
||
). In AlUla County, it was found in sympatry with
|
||
<taxonomicName id="BAA5E3946D70BF95E94DDC27119C14FE" authorityName="Heyden" authorityYear="1827" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="guttatus">
|
||
<emphasis id="7AF2CBE9770E73E3D2DE9C2730BF183F" italics="true">Ptyodactylus guttatus</emphasis>
|
||
</taxonomicName>
|
||
(in prep.),
|
||
<taxonomicName id="3B43AF7F484DAB76338CFAED725F8EC5" baseAuthorityName="Donndorff" baseAuthorityYear="1798" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" isUncertain="true" kingdom="Animalia" phylum="Chordata" rank="species" species="hasselquistii">
|
||
<emphasis id="416E76B381E2F6F9FE9F0ABBDC281CE1" italics="true">Ptyodactylus</emphasis>
|
||
cf.
|
||
<emphasis id="D4C3825F9B0CA3D982CDC00178E46C48" italics="true">hasselquistii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F9EB3BB14810A8E5C7F6DDDE3C1C3565" authorityName="Blanford" authorityYear="1874" class="Squamata" family="Gekkonidae" genus="Bunopus" kingdom="Animalia" phylum="Chordata" rank="species" species="tuberculatus">
|
||
<emphasis id="48FF112DFC528B20D1752AEF396E6BA3" italics="true">Bunopus tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D09C4431EC942D395AB88FB22A04178E" authorityName="Heyden" authorityYear="1827" class="Squamata" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="granosus">
|
||
<emphasis id="B16977179481C1D3ACDD7C62B7D152A0" italics="true">Hemidactylus granosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="332662A82A2E9B9F2795068F1D65CCEE" baseAuthorityName="Blanford" baseAuthorityYear="1874" class="Squamata" family="Gekkonidae" genus="Stenodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="doriae">
|
||
<emphasis id="08A03FFDA57C3E1B3E17F41FEB154854" italics="true">Stenodactylus doriae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="56BA05CF6D7346F301C4AED6CCB1E648" authorityName="Haas" authorityYear="1957" class="Squamata" family="Gekkonidae" genus="Stenodactylus" kingdom="Animalia" phylum="Chordata" rank="species" species="slevini">
|
||
<emphasis id="38D0F3130027CC475589FF19C4E5F6E3" italics="true">Stenodactylus slevini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5479EA8E6A48AD350F3FAB49AC4904DF" authorityName="Ribeiro-Junior, Tamar, Maza, Flecks, Wagner, Shacham, Calvo, Geniez, Crochet, Koch & Meiri" authorityYear="2022" class="Squamata" family="Gekkonidae" genus="Tropiocolotes" kingdom="Animalia" phylum="Chordata" rank="species" species="yomtovi">
|
||
<emphasis id="D16467014A987A6754A428A094D0E9F5" italics="true">Tropiocolotes yomtovi</emphasis>
|
||
</taxonomicName>
|
||
(the latter five all
|
||
<taxonomicName id="E44026B167A41EA4E255F20D7F779B11" class="Squamata" family="Gekkonidae" kingdom="Animalia" phylum="Chordata" rank="family">Gekkonidae</taxonomicName>
|
||
), and
|
||
<taxonomicName id="F89A7192D575A01868964EC46E190212" class="Squamata" family="Viperidae" genus="Echis" kingdom="Animalia" phylum="Chordata" rank="species" species="coloratus">
|
||
<emphasis id="9A6045BCF8E6A348D01947F14D7A74AC" italics="true">Echis coloratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELXAG" type="conservation status">
|
||
<paragraph id="B0DA1891ADFBE74CDC4E47927BFB202E">
|
||
<heading id="4C659474367B5B7EFF6144F421E200E3" reason="title">Conservation status.</heading>
|
||
</paragraph>
|
||
<paragraph id="2B05DBEF4D34B61C2608D390B3C912A0">
|
||
We propose to classify
|
||
<taxonomicName id="06F5F75263E1DD1EF6B5B4472AF7515B" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="EC2F521288B2BFCC637312DEBF427F25" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="68BA930FDABBF15555FE3002BA646336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
as Least Concern, based on the wide
|
||
<abbrev id="ABBRID0E3XAG" xlink_title="Extent of Occurrence">EOO</abbrev>
|
||
, and the diverse and continuous habitats where the species is encountered, including habitats with heavy anthropogenic disturbance (e. g., overgrazing) and urban habitats (see
|
||
<ext-link id="AA993DAFEC7845674766B6917C7C3384" ext-link-type="uri" xlink_href="https://www.inaturalist.org/observations/20014261" xlink_type="simple">https://www.inaturalist.org/observations/20014261</ext-link>
|
||
). Although
|
||
<taxonomicName id="D02C30A9F899ED673ED0D650EEC63980" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="8FC2945CE75D513ABD893503A344AF90" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C464080F614AF1B0E89D38FE614B647D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is at the moment known from 14 localities and its
|
||
<abbrev id="ABBRID0EQYAG" xlink_title="area of occurrence">AOO</abbrev>
|
||
is only
|
||
<quantity id="36DAC3098C48A4B8C438EC1527996EEF" metricMagnitude="4" metricUnit="m" metricValue="5.6" unit="km" value="56.0">56 km</quantity>
|
||
<superScript id="F503DAD5C95C8C159F792426D0C08A13">2</superScript>
|
||
, we consider it premature to classify it as Near Threatened as the limited number of localities could reflect the generally undersampled distribution and secretive habits of the species. Furthermore, six of the known localities occur within conservation areas located in AlUla County (
|
||
<collectingRegion id="B65F50EBB8F37115008148DAD162F177" country="Saudi Arabia" name="Al Madinah">Medina Province</collectingRegion>
|
||
), where protection measures have been implemented.
|
||
</paragraph>
|
||
<caption id="26CF998ABE8F334C4565F9946A999DA2" ID-DOI="10.3897/zse.100.123441.figure3" ID-arpha="E3E8E600-8802-58EE-8427-64A1CD26E77C" httpUri="https://binary.pensoft.net/fig/1059094" startId="F3">
|
||
<paragraph id="4C7C57A016FCF17E085DD371101CA57F">
|
||
<label id="F023733A83F432BBF184D23DB4BB3DE4">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="1FD3E74C04BABD2D2F4546473ECD4205">
|
||
Holotype (
|
||
<specimenCode id="2A430AF25270AF8C5322FBFF89A33EAB">
|
||
<abbrev content-type="institution" id="ABBRID0ELJDI" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 93850
|
||
</specimenCode>
|
||
, sample code FLI 447, bottom left) and paratype (
|
||
<specimenCode id="5A10BFE87735895977FE90934050753C">
|
||
<abbrev content-type="institution" id="ABBRID0EQJDI" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 94064
|
||
</specimenCode>
|
||
, sample code FLI 330, bottom right) specimens in life. Lateral and ventral views of their heads (above each life picture). Photo credit: FL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="250B8B2ED84125EF39AC22EED491C8E5" ID-DOI="10.3897/zse.100.123441.figure4" ID-arpha="9E62C61C-CB03-5B6D-A52E-71DFE2165C03" httpUri="https://binary.pensoft.net/fig/1059095" startId="F4">
|
||
<paragraph id="60A0B6C91899821006B749EB6D961AED">
|
||
<label id="52EBDCAF17802E81EF851873F06E8896">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="8AA2A1FE1C049A560A56AE1F2A5635FD">
|
||
Colour variation within
|
||
<taxonomicName id="17BC64542082E8890519C716DCCAA6F1" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="7837A782936102A0D6F1E433C3188770" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="97FDE68D9B3B7B189C22CD95612424D6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(or its lack, thereof). Top row: two unvouchered specimens from NEOM, Tabuk Province,
|
||
<abbrev id="ABBRID0ENKDI" xlink_title="Kingdom of Saudi Arabia">KSA</abbrev>
|
||
(photo credit: Euan Ferguson and Neil Rowntree); bottom left: paratype
|
||
<specimenCode id="853A7ADE30EC77FE95E2B85CDA2B2B28">
|
||
<named-content id="B302170B8748687826A916B8130135C9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/national-museum-prague-1" xlink_title="National Museum (Prague)">NMP</named-content>
|
||
76815
|
||
</specimenCode>
|
||
(sample code LP 760, photo credit: DME); bottom right: paratype
|
||
<named-content id="82404253B0ECA022F87318B0E88838DD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-historia-natural-0" xlink_title="Museo Nacional de Historia Natural">MNHN</named-content>
|
||
– RA – 2023.0013 (sample code JIR 544, photo credit: AAI).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0A54BAEBCA814B4C745A7D8668CB8A0F" ID-DOI="10.3897/zse.100.123441.figure5" ID-arpha="AB1E3665-53B2-5868-8FFA-5D48FD4C4834" httpUri="https://binary.pensoft.net/fig/1059096" startId="F5">
|
||
<paragraph id="F228093B3C90E6B8EC65F5929EEEBADF">
|
||
<label id="73758406B016AFA37A9047E62487B4E5">Figure 5.</label>
|
||
</paragraph>
|
||
<paragraph id="ADC293BC0CD158484FF42346FE0CA811">
|
||
Habitats of the holotype and two paratype specimens of
|
||
<taxonomicName id="B5F76D55D508D62FE62804A58934C3A4" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="690AE694564CB6DAC11CA4FF95A9782E" italics="true">R. hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A3D89BC33A7982101E66404BB790DB32" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Top: locality of the holotype
|
||
<specimenCode id="C3C48E1AF2DFF0EB64929399695515AC">
|
||
<abbrev content-type="institution" id="ABBRID0ETLDI" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 93850
|
||
</specimenCode>
|
||
at the Shaaran NR, AlUla County, Medina Province,
|
||
<abbrev id="ABBRID0EYLDI" xlink_title="Kingdom of Saudi Arabia">KSA</abbrev>
|
||
(photo credit: FL); bottom left: habitat of the paratype
|
||
<named-content id="C34C1865B4ECF1D857A0E71FB1CCEC6D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-nacional-de-historia-natural-0" xlink_title="Museo Nacional de Historia Natural">MNHN</named-content>
|
||
– RA – 2023.0013 at Wadi Al-Azraq, Jabal Salma, Hail Province,
|
||
<abbrev id="ABBRID0EBMDI" xlink_title="Kingdom of Saudi Arabia">KSA</abbrev>
|
||
(photo credit: AAI); bottom right: habitat of the paratype
|
||
<specimenCode id="5BC061CBF5ED7DCA740BF590986B9219">
|
||
<abbrev content-type="institution" id="ABBRID0EFMDI" xlink_title="AlUla Museum">RCU-URN</abbrev>
|
||
- 94065
|
||
</specimenCode>
|
||
the Harrat Khaybar, Hail Province,
|
||
<abbrev id="ABBRID0EKMDI" xlink_title="Kingdom of Saudi Arabia">KSA</abbrev>
|
||
(photo credit: FL).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3327FB2D7C26A7E577B984F1AE10DE30" ID-DOI="10.3897/zse.100.123441.figure6" ID-arpha="14C62C57-D03C-5869-9899-625973D88B30" httpUri="https://binary.pensoft.net/fig/1059097" startId="F6">
|
||
<paragraph id="9477F85D703C1B99E8BD8BB08E39E2EC">
|
||
<label id="026B1102DAD347D12619CE47C5252AB0">Figure 6.</label>
|
||
</paragraph>
|
||
<paragraph id="79AFD262710F7514B56C63C14230222D">
|
||
Distribution of
|
||
<taxonomicName id="21F7B36245A447F72EB4D403E25BA625" authority="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito, 2024" authorityName="Licata & Pola & Šmíd & Ibrahim & Liz & Santos & Patkó & Abdulkareem & Gonçalves & AlShammari & Busais & Egan & Ramalho & Smithson & Brito" authorityYear="2024" class="Squamata" family="Colubridae" genus="Rhynchocalamus" kingdom="Animalia" phylum="Chordata" rank="species" species="hejazicus" status="sp. nov.">
|
||
<emphasis id="6909F6C4E10222F85D109536A8A33D4E" italics="true">Rhynchocalamus hejazicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="763FF25FFC61AE6451FB9E4B2183FFF2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
showing the location of the material examined in this study. Circles indicate samples used for the genetic analyses, squares indicate examined voucher specimens, and diamonds indicate photographic records. The black arrow denotes the type locality (Shaaran NR in AlUla County, Medina Province). Geographical sampling is overlaid by the predicted species distribution model categorised into binary predictions of the suitable areas. The threshold used for the categorisation was the lowest predicted suitability value among the occurrence points (= 0.05).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |