treatments-xml/data/77/5C/1A/775C1A1E890053C5A50DD784E840DDB6.xml

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<mods:title id="1811B5EDEB4607EF808E816812F01EBA">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:namePart id="10BF4F39D220A63B14F468AC868CD83A">Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation id="4C2AD238A5F32DF677E5D5E5B1D09D2C">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
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<mods:namePart id="707A4835371B11DE90ACC1C8E81AA507">Pinto-da-Rocha, Ricardo</mods:namePart>
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<mods:affiliation id="4F4A5306B4D44AD0F47D61A1BD9E0633">Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<treatment id="775C1A1E890053C5A50DD784E840DDB6" ID-GBIF-Taxon="199468675" LSID="urn:lsid:zoobank.org:act:11B40D2F-108B-49D2-AF56-79AC64978832" httpUri="http://treatment.plazi.org/id/775C1A1E890053C5A50DD784E840DDB6" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="DCAEAD405FD3ABD82B79D6BC0E833DBC" pageId="0" pageNumber="309" type="nomenclature">
<paragraph id="F218FBCA78AE8AD2E9D20E67285746EE" pageId="0" pageNumber="309">
3.41. 
<taxonomicName id="63E9D9062B4312F2D9DC591C7970D6C8" ID-CoL="9X65Y" LSID="http://zoobank.org/11B40D2F-108B-49D2-AF56-79AC64978832" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus bergmani" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="bergmani" status="sp. nov.">Metasarcus bergmani</taxonomicName>
<taxonomicNameLabel id="B7E8D42FE016D7C59997A09346C8B442" pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="297212EB5046921987DC6E9FAA15DE27" pageId="0" pageNumber="309" type="description">
<paragraph id="1C0E4021A8FECC910272A6362441685B" pageId="0" pageNumber="309">
<figureCitation id="68CB8F742A63C147A041ECBAE99F38DA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5B</figureCitation>
<figureCitation id="B042AA41053B7D432A6F447DF812C085" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10C, D</figureCitation>
<figureCitation id="3243E252B3FD437A594BC325DF577F3F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">, 19A-C</figureCitation>
<figureCitation id="3677CBA1D37CF4EB9966D31AB64ACD52" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">, 25A, B</figureCitation>
<figureCitation id="C433FF097AE11FEF2E836DD3E5B3B179" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95136E83374A99E26EAD0C0BB08D4573" pageId="0" pageNumber="309" type="description">
<paragraph id="CBD750CA3CF20F197E942684AAED8954" pageId="0" pageNumber="309">Description.</paragraph>
<paragraph id="94E667AAA3D7D9AA7039ADB12A0A1E6B" pageId="0" pageNumber="309">
<emphasis id="6707BC9F98F922090A087B17BF37F8B6" bold="true" pageId="0" pageNumber="309">
MALE: 
<emphasis id="14270F87A8A9068449FBA33C7ECF3424" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="3B581B86A993490BED62BB672B70E127" italics="true" pageId="0" pageNumber="309">n</emphasis>
=5) DSW: 5.2-6.0 (5.2); DSL: 5.5-6.8 (5.5); CL: 2.1-2.5 (2.1). FIVL: 12.5-13.0 (12.9). ChL: 2.0-2.1 (2.1). 
<emphasis id="F652865E5B5DCCE5E3639AA25CF4CA64" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
(Fig. 
<figureCitation id="1B6B8F340AE927475D7659ACEA848EE2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">25A</figureCitation>
): Chelicerae, pedipalpus and trochanters I-III yellowish green. Ocularium, anterior and lateral margin of DS, anterior part of the carapace, coxa and trochanter IV reddish. Posterior part of the carapace and areas I-IV orange, in lighter tone. Spines of area III, apophysis of coxa IV and anterior angle on area I most blackened. Granules of posterior margin of DS and free tergites I-III orange. Grooves of dorsal scutum white. 
<emphasis id="F9DB501C1D7A62A059EA664D7F46D05B" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig. 
<figureCitation id="9F7EBCA8E3042607AE04907D0D4BA3B6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5B</figureCitation>
) Gamma-P-type DSS, with concave posterior margin of DS. Anterior margin of carapace with median elevation, covered with granules. Ocularium with median depression granular, and 4-5 small tubercles near the eyes. Carapace with sparse granules. Areas I-IV covered by few granules; Area I divided by a longitudinal groove and invaded by area II; with a pair of median small tubercles. Area II and IV unarmed. Area III with a pair of median spines, parallel, directed posteriorly. Lateral margins of DS granular. Posterior margin of DS and free tergites with a row of small granules. Free tergite III with a spiniform apophysis slightly longer than the length of this tergite. 
<emphasis id="C3B7D98258CC7BD0D0477BFC480BC42B" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig. 
<figureCitation id="9131F3B42961C10015EEC52F66AB51D9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5B</figureCitation>
) Not swollen, similar to females. Segment I covered with small granules. Segment II covered with granules larger than those of segment I; finger with four teeth. Segment III with three teeth. 
<taxonomicName id="CD2B7E7B76ED3922BD2B6EAD7957D3D6" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="E0D2A6D8D0322D7B26613D68EA26C424" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Trochanter with a ventroapical setiferous tubercle. Femur with a ventral row of four spines and a proapical spine. Patella with a proventral small setiferous tubercle. Tibia: prolateral IiII/iIiII, retrolateral IiIIi. Tarsus: prolateral iIiIi, retrolateral iIiIi/iIiIii. 
<emphasis id="21A61328BF4512C483BECAFBF01B8C09" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a median row of 5-6 setiferous tubercles and small sparse granules. Coxae II-III with setiferous tubercles and sparse granules. Coxa IV with granules smaller than those on other coxae, sparsely distributed. Smooth genital area. Free sternites with few granules. 
<emphasis id="AF271CA848D3A7CFC571750690C89C13" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs 
<figureCitation id="1B92FFF9AA6A21988202077B95EC34BA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5B</figureCitation>
, 
<figureCitation id="B729593FB7DB81569EEE9BC59CB999E4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10C, D</figureCitation>
) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with a large retrolateral apophysis, curved posteriorly and laterally. Trochanters I-IV with small granules. Trochanter IV with prolateral base expanded and a retrolateral tubercle. Femora I-III straight and unarmed, with few small granules. Femur IV straight, with few sparsely scattered small granules and a small retrobasal blunt tubercle. Tarsal segmentation: (
<emphasis id="5A36651DC38853E2957A310560103E21" italics="true" pageId="0" pageNumber="309">n</emphasis>
=5) 9, 17-19 (18), 11-12 (12), 12-14 (14). 
<emphasis id="89B492BD677C1E01FA3EFD676BB31C5E" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig. 
<figureCitation id="D7F5AC690728B933FBA22EFCD040E011" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Penis of Metasarcus, dorsal, lateral and ventral views, respectively. A - C M. bergmani sp. nov.; D - F M. clavifemur (Roewer, 1929); G - I M. fellinii sp. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure19" httpUri="https://binary.pensoft.net/fig/729893" pageId="0" pageNumber="309">19A-C</figureCitation>
) VP rectangular, distal margin straight and with small lateral projections; with distal laterodorsal depressions; slightly concave in lateral view. MS C1-C3 subapical long and straight; MS A1 sub basal long and straight; MS B1 basal short and straight. Lateral sacs long and robust, with large base and blunt apex; with long tapered T3-like microsetae near the base and short and flattened near the apex. Stylus long and cylindrical (apex cannot be viewed in SEM). Dorsal process absent. Promontory conspicuous and convex. - 
<emphasis id="C190D5930A466C977F7A50B7403BB3DF" bold="true" pageId="0" pageNumber="309">
FEMALE: 
<emphasis id="8DA4A7EDEF968153AA12BE886ABDB9FA" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis id="E50DC5DC4AEB15EBD983E995D9BB1E5E" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 5.0-5.2; DSL: 5.2-5.3; CL: 2.0. FIVL: 12.1-11.3. ChL: 2.0-2.2. (Fig. 
<figureCitation id="6157A868D7260F4148558D4455022007" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Live specimens of Metasarcus. A M. bergmani sp. nov., male; B female; C M. clavifemur (Roewer, 1929), male; D female; E M. fellinii sp. nov., male; F female; G M. kurosawai sp. nov., male; H female." figureDoi="10.3897/asp.80.e73829.figure25" httpUri="https://binary.pensoft.net/fig/729899" pageId="0" pageNumber="309">25B</figureCitation>
) Granulation denser than in male, mainly in the areas of the dorsal scutum. Spine of area III with multiple granules at base. Retrolateral apophysis in the coxa IV short, much shorter than in male. Trochanter IV without expansion of the prolateral base. Apophysis of free tergite III longer than in male. Tarsal segmentation: (
<emphasis id="39E7E4598FC072DB9AC45F9B2D14C1B5" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 8-9, 16-18, 10, 11-12.
</paragraph>
</subSubSection>
<subSubSection id="9A03B84FC703404341974AD0446CDE84" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="FE58875F0A27946D20A63AC950F1F94E" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="71CC040CDFBC9C4EB4A06D6F7E9C3C24" pageId="0" pageNumber="309">
It differs from other species of the genus by having a large retrolateral apophysis on coxa IV, curved toward externally, and a short spiniform apophysis in free tergite III (Fig. 
<figureCitation id="E2C4B7A5D00B9DF028C76113F3AEC351" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="132D4267B25129BA6A90F04AB5E37DB9" pageId="0" pageNumber="309" type="etymology">
<paragraph id="D6E828EAED0A57C54E6EE7021787A289" pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph id="6D7243B032ADCF59463D60AF2268FA85" pageId="0" pageNumber="309">The specific epithet of masculine gender, in the genitive form, dedicated to the Swedish filmmaker, director, producer and writer Ernst Ingmar Bergman (1918-2007).</paragraph>
</subSubSection>
<subSubSection id="C4DFEDE647883906FF068BD86DF2016C" pageId="0" pageNumber="309" type="distribution">
<paragraph id="5D88E0AE889FA5E9791A035D6D013239" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="56FE40783EC2690DD2B8CA3F8732A745" pageId="0" pageNumber="309">
(Fig. 
<figureCitation id="E821C250443DD9F90ABC1ACC40BE958C" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
) BOLIVIA. Cochabamba. Near Corani.
</paragraph>
</subSubSection>
<subSubSection id="257BF71B26CC79E67613A66D4B221089" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="8226BB27BAC2FF247FE8977D982BF286" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="E8F2809C0787DCA491CAA2E07028BB46" pageId="0" pageNumber="309">
<emphasis id="A488AB9A90DDF5BBAE40BB511B44B55E" bold="true" pageId="0" pageNumber="309">
<emphasis id="8EBB7F1C438D50915D88848DE569B8BB" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
: 
<materialsCitation id="B1DCDDC3F5108800A804F4CE66C0B5C6" ID-GBIF-Occurrence="3889641341" collectingDate="2010-12-01" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, A. Saravia" country="BOLIVIA" latitude="-17.188583" location="Corani" longLatPrecision="1" longitude="-65.896996" specimenCount="♂" stateProvince="Cochabamba" typeStatus="Holotype">
<typeStatus id="AF86960A65D5AAB6CB137E655DA45DC4">Holotype</typeStatus>
<specimenCount id="EEDAC8F6BF87BE46AB7E56942205C0D7">♂</specimenCount>
, ' 
<collectingCountry id="859B8AF364F0987D3003A05C4EFE13A2" name="Bolivia">BOLIVIA</collectingCountry>
, 
<collectingRegion id="2BCA53239F819D772C3534528989588D" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
, road next to 
<location id="7EDFF29CB64716EAA28C8F3FBACB5FF7" LSID="urn:lsid:plazi:treatment:775C1A1E890053C5A50DD784E840DDB6:7EDFF29CB64716EAA28C8F3FBACB5FF7" country="BOLIVIA" latitude="-17.188583" longLatPrecision="1" longitude="-65.896996" name="Corani" stateProvince="Cochabamba">Corani</location>
,| 
<geoCoordinate id="713A397E753574771E9B01A2A7C7FA70" degrees="17" direction="south" minutes="11" orientation="latitude" precision="1" seconds="18.9" value="-17.188583">17°11′18.9″S</geoCoordinate>
<geoCoordinate id="9D72D511CA5597A09DCBA11B8BC3BB7D" degrees="65" direction="west" minutes="53" orientation="longitude" precision="1" seconds="49.2" value="-65.896996">65°53′49.2″W</geoCoordinate>
, 
<collectingDate id="4F5A4FE1D5299A9B088F589A82AF60CB" value="2010-12-01">01/XII/2010</collectingDate>
, 
<collectorName id="18E4C390075C9866BBFC18D8A07B25E4">R. Pinto-da-Rocha</collectorName>
, 
<collectorName id="4B17C2402E7551ED9245CCE7F46BF109">A. Benedetti</collectorName>
, 
<collectorName id="D576B8D01BA4491394D762D2DF3CB549">J. Ochoa</collectorName>
&amp; 
<collectorName id="CAEAAE140C081EA2883E58E2A1843C1A">A. Saravia</collectorName>
leg. (CBF)
</materialsCitation>
</emphasis>
- 
<emphasis id="4C0E5B4D0C08CE53CAD2CF9A652A151B" bold="true" pageId="0" pageNumber="309">
<typeStatus id="B78B6A79742A33C1064D4236236FB024">Paratypes</typeStatus>
</emphasis>
<specimenCount id="6AC8A161387D9D9E7F86646666CA9B81" type="male">1 ♂</specimenCount>
, 
<specimenCount id="754FD64974767D74F3D653524C89A5C1" type="female">1 ♀</specimenCount>
, 
<normalizedToken id="FC4F66D75038B1A16B7CC821BFB5D91A" originalValue="‘ditto’">'ditto'</normalizedToken>
(CBF); 
<emphasis id="284366FF5FD70285F73A97C33838526B" bold="true" pageId="0" pageNumber="309">
<typeStatus id="348DE9ABF0AC9E5EFC99C0BEC8167793">Paratypes</typeStatus>
</emphasis>
<specimenCount id="35C8E1E1434CABE9B0351C8E7A46E105" type="male">3 ♂</specimenCount>
, 
<specimenCount id="C378828AFF296D5F79BD37DD5ECD25A1" type="female">3 ♀</specimenCount>
, 
<normalizedToken id="6AD584FD517DD18D3822ACF77A868866" originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36983).
</paragraph>
</subSubSection>
</treatment>
</document>