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<mods:title id="BA7EC7000B3EA34D48296DE2D8230C4D">A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species.</mods:title>
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<mods:namePart id="EAB25E05A6B7D42278308FF7F6423CD6">Juan J. Schmitter-Soto</mods:namePart>
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<mods:date id="29B101020E3351F57AB2FC0676D86017">2007</mods:date>
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<treatment id="616F60769F866FBAA81A072270CAA8D8" ID-DOI="http://doi.org/10.5281/zenodo.6248732" ID-GBIF-Taxon="100117427" ID-Zenodo-Dep="6248732" LSID="urn:lsid:plazi:treatment:616F60769F866FBAA81A072270CAA8D8" httpUri="http://treatment.plazi.org/id/616F60769F866FBAA81A072270CAA8D8" lastPageNumber="6" pageNumber="4">
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[[ Genus
<taxonomicName id="4066ABE2129365CA4743304C7F848AD4" ID-CoL="328M" ID-ENA="61816" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus Gill</taxonomicName>
]]
</paragraph>
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<subSubSection id="5A74C696163EEF1BB92CAFB99856648E" pageNumber="3" type="discussion">
<paragraph id="C12D073042F419D0DBADCFF39CDF87DF" lastPageNumber="5" pageNumber="4">
The cichlid fish genus
<taxonomicName id="A6D4D449BFDE4FFFE81DC6BDE45CC3FF" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
was originally proposed as a subgenus of
<taxonomicName id="5324C3D5297157FA57A7ED61EEC62A20" LSID-ZBK="urn:lsid:zoobank.org:act:3A1BBC79-44D3-4A63-8873-730518BF6096" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros Heckel 1840:362" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Heros Heckel</taxonomicName>
by Gill (in
<bibRefCitation id="2CEEF6EBFA6197017A5AC3AD57BB9C4A" author="Gill, T. &amp; Bransford, J. F." pageNumber="3" pagination="175 - 218" refId="ref35271" refString="Gill, T. &amp; Bransford, J. F. (1877) Synopsis of the fishes of Lake Nicaragua. Proceedings of the Academy of Natural Sciences of Philadelphia, 29, 175 - 218." type="journal article" year="1877">Gill &amp; Bransford 1877</bibRefCitation>
), and later treated as an informal “section” of
<taxonomicName id="F8D2598898EB22352708F8D94424E7C1" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma Swainson</taxonomicName>
in Regan’s (1905) revision of the genus. The generic status of
<taxonomicName id="34CA680FBCBC133AC67EDB530F578B87" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
sensu
<bibRefCitation id="CF694AA6BD7B9CD97949A1B72516FF69" author="Regan, C. T." pageNumber="3" pagination="433 - 445" refId="ref36996" refString="Regan, C. T. (1905) A revision of the fishes of the American cichlid genus Cichlosoma and of the allied genera. Annual Magazine of Natural History, 16, 60 - 77, 225 - 243, 316 - 340, 433 - 445." type="journal article" year="1905">Regan (1905)</bibRefCitation>
, encompassing more than 100 species, has been in a state of uncertainty since Kullander’s (1983) restriction of
<taxonomicName id="0C16721D26EBF6B32AF34C0B394C81B3" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
sensu stricto to 12 South American species. Of the many remaining groups of species formerly referred to
<taxonomicName id="7D80A84B366A0887B555327109B6D52B" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
, those
<pageBreakToken id="FBEA60CE405DC10FE52571AC922A8216" pageNumber="3">referred</pageBreakToken>
to
<taxonomicName id="665025FFDE5B51570B38D16A68B8E70B" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
have been among the strongest candidates for recognition as a monophyletic group (e.g.
<bibRefCitation id="2F0315DE6C1C7151B9FE5B48F0361F89" author="Miller, R. R." pageNumber="3" pagination="18 - 19" refId="ref36588" refString="Miller, R. R. (1993) [Review of] North and Central American Cichlids, by Donald Conkel. Buntbarsche Bulletin, 157, 18 - 19." type="journal article" year="1993">Miller 1993</bibRefCitation>
,
<bibRefCitation id="6EE8BA28E478C48D0FA22431DDD4A3ED" author="Miller, R. R. &amp; Minckley, W. L. &amp; Norris, S. M." pageNumber="3" refId="ref36620" refString="Miller, R. R., Minckley, W. L. &amp; Norris, S. M. (2005) Freshwater Fishes of Mexico. Museum of Zoology, University of Michigan, &amp; University of Chicago Press, Chicago, 652 pp." type="book" year="2005">Miller et al. 2005</bibRefCitation>
). Species assigned to
<taxonomicName id="5289B8B7869FFC7B71AD010A76F721D2" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
are major components of the neotropical Central American ichthyofauna from southern Mexico to Panama, occurring across all the ichthyolimnological provinces of
<bibRefCitation id="F1F97AE5948878BD4CB5477BFB55496A" author="Bussing, W. A." pageNumber="3" pagination="157 - 175" refId="ref34665" refString="Bussing, W. A. (1976) Geographic distribution of the San Juan ichthyofauna of Central America with remarks on its origin and ecology. In: Thorson, T. B. (Ed.), Investigations of the Ichthyofauna of Nicaraguan Lakes. University of Nebraska, Lincoln, pp. 157 - 175." type="book chapter" year="1976">Bussing (1976)</bibRefCitation>
, but the monophyly of the genus has not been explicitly demonstrated and its species composition has been far from clear.
</paragraph>
<paragraph id="BFB6A382D329BB15AE1CF211E28764C1" pageNumber="5">
Since a comprehensive analysis of
<taxonomicName id="85653BA31CDDD5BD12ED06FF53FE91A5" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
sensu lato appeared unlikely,
<bibRefCitation id="96CD6FA63F8AD88D19D5D827369F12A3" author="Kullander, S. O." pageNumber="3" pagination="149 - 172" refId="ref35926" refString="Kullander, S. O. (1996) Heroina isonycterina, a new genus and species of cichlid fish from Western Amazonia, with comments on cichlasomine systematics. Ichthyological Exploration of Freshwaters, 7, 149 - 172." type="journal article" year="1996">Kullander (1996)</bibRefCitation>
advocated the provisional use of available generic names, a procedure that some workers had already adopted. Kullander himself (1983),
<bibRefCitation id="8307840423B9E3950C0946B490EF7F36" author="Stiassny, M. L. J." pageNumber="3" pagination="1 - 31" refId="ref37342" refString="Stiassny, M. L. J. (1991) Phylogenetic intrarelationships of the family Cichlidae. In: Keenleyside, M. H. A. (Ed.), Cichlid Fishes - Behavior, Ecology and Evolution. Chapman &amp; Hall, London, 1 - 31." type="book chapter" year="1991">Stiassny (1991)</bibRefCitation>
, and
<bibRefCitation id="17F715F671DB858D108C0657A787E15B" author="Miller, R. R." pageNumber="3" pagination="18 - 19" refId="ref36588" refString="Miller, R. R. (1993) [Review of] North and Central American Cichlids, by Donald Conkel. Buntbarsche Bulletin, 157, 18 - 19." type="journal article" year="1993">Miller (1993)</bibRefCitation>
had earlier recommended that the use of ‘
<taxonomicName id="146C2CEA4F8C63A074F1D36F2154A741" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
’- with the quotation marks indicating its informality - would prevent further confusion. Because the systematics of all species previously referred to
<taxonomicName id="5E72ED661BF7DA29BA246A8F20BECE65" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
had not yet been completely resolved,
<bibRefCitation id="4487F14CCCA49C53426B2DF77534AC86" author="Nelson, J. S. &amp; Crossman, E. J. &amp; Espinosa-Perez, H. &amp; Findley, L. T. &amp; Gilbert, C. R. &amp; Lea, R. N. &amp; Williams, J. D." pageNumber="3" refId="ref36696" refString="Nelson, J. S., Crossman, E. J., Espinosa-Perez, H., Findley, L. T., Gilbert, C. R., Lea, R. N. &amp; Williams, J. D. (2004) Common and Scientific Names of Fishes from the United States, Canada, and Mexico. 6 th ed. American Fisheries Society, Bethesda. 386 pp." type="book" year="2004">Nelson et al. (2004: 151-153)</bibRefCitation>
chose the conservative approach and tentatively referred all species to
<taxonomicName id="F934B5E2ECC71AC2E6A878E728FC1392" LSID-ZBK="urn:lsid:zoobank.org:act:2D5E18F7-A978-4A06-8907-A1F0233A2D39" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma Plesiops Swainson 1839:230" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cichlasoma</taxonomicName>
in the most recent checklist of fishes for the United States, Canada, and Mexico.
</paragraph>
<paragraph id="D272DC394370AB9F9CF7A67F841B1A39" pageNumber="5">
Several species have been assigned to
<taxonomicName id="09A1AC85621408AB463FFE49D9B25687" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
more or less consistently in the literature (Table 1).
<bibRefCitation id="0AFFFBE692206809C467AA49C0118308" author="Allgayer, R." pageNumber="3" pagination="13 - 20" refId="ref34520" refString="Allgayer, R. (2001) Description d'un genre nouveau, Cryptoheros, d'Amerique Centrale et d'une espece nouvelle du Panama (Pisces: Cichlidae). L’An Cichlide, 1, 13 - 20." type="journal article" year="2001">Allgayer (2001)</bibRefCitation>
restricted
<taxonomicName id="C43D1E136751397E57FE57F055859ACE" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
to include the two largest species (
<taxonomicName id="11E2013A4108196C6BC800EE0FAC0578" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="centrarchus">Ar. centrarchus</taxonomicName>
and
<taxonomicName id="350B83001B17FEA41E4A21FBFB219C96" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spinosissimus">Ar. spinosissimus</taxonomicName>
), and referred six others to a new genus,
<taxonomicName id="E58B238D2F05AD6DEA779DD2DFCB23FE" LSID-ZBK="urn:lsid:zoobank.org:act:273F29E5-91F4-45BE-8447-659BDE8939E2" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" lsidName-ZBK="Cryptoheros Allgayer 2001" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cryptoheros</taxonomicName>
(type species,
<taxonomicName id="61BD7C7F19A0C46675F2664624F5FB00" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cr. spilurus</taxonomicName>
), which also included the new species
<taxonomicName id="636074061B9B0EEC03A784AE71430EAD" LSID-ZBK="urn:lsid:zoobank.org:act:B8A94766-C505-4E59-98D6-19AEF7E7B80A" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" lsidName-ZBK="Cryptoheros altoflavus Allgayer 2001:16" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="altoflavus">Cr. altoflavus</taxonomicName>
. Other species that have sometimes been assigned to
<taxonomicName id="4D2CB9DBEA0F6A88019B52A265112DB3" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
(Table 1) include:
<taxonomicName id="261DAB08F3FB2C75E6EBA012F44B4ACD" LSID-ZBK="urn:lsid:zoobank.org:act:D76352FE-1F2C-4AFD-9081-678A86D6C797" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros octofasciatus Regan 1903:417" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatus">Heros octofasciatus</taxonomicName>
, though more frequently placed in incertae sedis;
<taxonomicName id="376CF98541B8343BD705C3A724B97AC6" LSID-ZBK="urn:lsid:zoobank.org:act:9DC9350D-12F3-4E33-9712-2080915675C8" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros citrinellus Günther 1864:153" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="citrinellus">H. citrinellus</taxonomicName>
, albeit demonstrated by
<bibRefCitation id="BB1AE3B6058F5FDD47E027A4FF0EDBCB" author="Roe, K. J. &amp; Conkel, D. &amp; Lydeard, C." pageNumber="3" pagination="366 - 376" refId="ref37128" refString="Roe, K. J., Conkel, D. &amp; Lydeard, C. (1997) Molecular systematics of Middle American cichlid fishes and the evolution of trophic-types in ‘ Cichlasoma (Amphilophus) ’ and ‘ C. (Thorichthys) ’. Molecular Phylogenetics and Evolution, 7, 366 - 376." type="journal article" year="1997">Roe et al. (1997)</bibRefCitation>
to be an
<taxonomicName id="6243B4A8535A185BBF11D28135BFBB13" LSID-ZBK="urn:lsid:zoobank.org:act:60986D0C-086A-4E68-8914-90992063465B" family="Cichlidae" genus="Amphilophus" kingdom="Animalia" lsidName-ZBK="Amphilophus Agassiz 1859:408" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Amphilophus</taxonomicName>
;
<taxonomicName id="821C8F6CA30D1A13E2917C477690CDFD" LSID-ZBK="urn:lsid:zoobank.org:act:3CC6C0EE-F67A-4C3C-BC4D-7F797A1E0913" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros multispinosus Günther 1867:601" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="multispinosus">H. multispinosus</taxonomicName>
, generally considered in the monotypic genus
<taxonomicName id="70DF8187B05FCD330C3048874FD2B960" LSID-ZBK="urn:lsid:zoobank.org:act:4B9BCDD6-7318-4D82-B13F-E89502F50F24" family="Cichlidae" genus="Herotilapia" kingdom="Animalia" lsidName-ZBK="Herotilapia Pellegrin 1904:(165) 247" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Herotilapia</taxonomicName>
; and
<taxonomicName id="81AC2059B984A105D7CD326F45BFB500" LSID-ZBK="urn:lsid:zoobank.org:act:2411EEF8-6445-445C-AFA1-B6C5A08A5161" family="Cichlidae" genus="Neetroplus" kingdom="Animalia" lsidName-ZBK="Neetroplus panamensis Meek &amp; Hildebrand 1913:90" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="panamensis">Neetroplus panamensis</taxonomicName>
, variously included in
<taxonomicName id="87584CAD878AB74DCAB1DFD5F7B19572" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
(in
<bibRefCitation id="137084C20C6F6D3D7FDA5368A311549C" author="Kullander, S. O." pageNumber="3" pagination="605 - 654" refId="ref35967" refString="Kullander, S. O. (2003) Family Cichlidae (Cichlids). In: Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J., Jr. (eds.). Checklist of the Freshwater Fishes of South and Central America. Edipucrs, Porto Alegre, pp. 605 - 654." type="book chapter" year="2003">Kullander 2003</bibRefCitation>
),
<taxonomicName id="BBA8124AF283B65B2F00F09B62250097" LSID-ZBK="urn:lsid:zoobank.org:act:0D9C9FBB-6F98-49B8-B90F-61D45F3FDF69" family="Cichlidae" genus="Theraps" kingdom="Animalia" lsidName-ZBK="Theraps Günther 1862:284" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Theraps</taxonomicName>
(in
<bibRefCitation id="37398A9615943E84DD48761597C428B4" author="Eschmeyer, W. N." pageNumber="3" refId="ref34964" refString="Eschmeyer, W. N. (Ed.) (2001) Catalog of Fishes. California Academy of Sciences, San Francisco. Available from http: // www. calacademy. org / research / ichthyology / Catalog / (accessed 15 December 2001)" type="url" year="2001">Eschmeyer 2001</bibRefCitation>
), or
<taxonomicName id="1C34D66202A28330E758235CE833F8A8" LSID-ZBK="urn:lsid:zoobank.org:act:7A9BA3FB-1462-4C73-A40A-F9A660B3AA90" class="Malacostraca" family="Homolidae" genus="Hypsophrys" kingdom="Animalia" lsidName-ZBK="Hypsophrys Agassiz 1859:408" order="Decapoda" pageNumber="3" phylum="Arthropoda" rank="genus">Hypsophrys</taxonomicName>
(in
<bibRefCitation id="27E705F1ED9FB966982EB067805ACE50" author="Bussing, W. A." pageNumber="3" pagination="1 - 468" refId="ref34754" refString="Bussing, W. A. (1998) Peces de las aguas continentales de Costa Rica. Revista de Biologia Tropical, 46, 1 - 468." type="journal article" year="1998">Bussing 1998</bibRefCitation>
).
</paragraph>
<caption id="120531CAD34C10B7AFAE4C77E9798AE2" pageNumber="3">
<paragraph id="78E5140A9D1CB11480515C67E97EAEB5" pageNumber="5">
TABLE 1. Valid and nominal species once or currently included in
<taxonomicName id="5B650C498BE19604082AF2728D7CA297" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
. Authorship, year of description, orthography, and present status or name, after
<bibRefCitation id="3C234C9AB81E94B337575711ACE9719E" author="Eschmeyer, W. N." pageNumber="3" refId="ref35012" refString="Eschmeyer, W. N. (Ed.) (2005) Catalog of Fishes. California Academy of Sciences, San Francisco. Available from http: // www. calacademy. org / research / ichthyology / Catalog / (accessed 15 January 2005)" type="url" year="2005">Eschmeyer 2005</bibRefCitation>
, except for changes proposed or supported in this work.
</paragraph>
</caption>
<paragraph id="F893D4FD45856D57733438B1F5B4535D" pageNumber="3">
<table id="E8677F7C46F34675F21B065D371D9B36" border="0" frame="box" pageNumber="3" rules="all">
<tr id="73E5DC5F7BEC2DFEB8198584FA9C652E" pageNumber="3">
<td id="75EEFDB3B6CD2C7D6405D90286A5024A" pageNumber="3">Basonym</td>
<td id="59CCF001ED4F952C09967FD3B84E3D3A" pageNumber="3">Author and year</td>
<td id="0C6A07315F39D1CC4FD772BEB0E2F99C" pageNumber="3">
Included in
<taxonomicName id="F54B5C340C01189959D2394E3CD86BD3" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
by:
</td>
<td id="4C26588C1C6DB9456E71731F39503673" pageNumber="3">Present status or name</td>
</tr>
<tr id="60456CA52517EE12948C92837CEFFA41" pageNumber="3">
<taxonomicName id="F3C8114D98C23E5F2DED1CB64E57C99A" LSID-ZBK="urn:lsid:zoobank.org:act:CEB65659-51E8-484D-8200-F9850A626462" authority="Guenther, 1862" authorityName="Guenther" authorityYear="1862" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros spilurus Günther 1862:289" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">
<td id="41081A7F2F9D0F83F2497C0412381A43" pageNumber="3">Heros spilurus</td>
<td id="0477FBD3D49F37272A81AA826412B588" pageNumber="3">
<bibRefCitation id="E0187E9E012BC8257EA101FB0521DE62" author="Guenther, A." pageNumber="3" refId="ref35343" refString="Guenther, A. (1862) Catalogue of the Fishes in the British Museum. Vol. IV. British Museum, London, 534 pp." type="book" year="1862">
<normalizedToken id="823C2DE6ABFB948BFFA8A034E2B585F8" originalValue="Günther" pageNumber="3">Guenther</normalizedToken>
, 1862
</bibRefCitation>
</td>
</taxonomicName>
<td id="59747B9F565E4128FA5196EE384042C4" pageNumber="3">
<bibRefCitation id="C5337967DF69D056EA26E50C2D7E6CE7" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="766A188B9070A7C4EFD810FB66F6C836" pageNumber="3">
<taxonomicName id="A413EEBEB33D8B7A53B567044DA62CF5" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cryptoheros spilurus</taxonomicName>
</td>
</tr>
<tr id="5507BFCCF2674D72E936A1486035BAAA" pageNumber="3">
<taxonomicName id="81BD6D2F758E8ADBA28CA07CC93DA5AD" LSID-ZBK="urn:lsid:zoobank.org:act:9DC9350D-12F3-4E33-9712-2080915675C8" authority="Guenther, 1864" authorityName="Guenther" authorityYear="1864" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros citrinellus Günther 1864:153" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="citrinellus">
<td id="FC0AF19465613DB98E914C3496AC86D1" pageNumber="3">Heros citrinellus</td>
<td id="48325BFED55EF5657F3DB1DDBA25985D" pageNumber="3">
<bibRefCitation id="89B0D67B6A92BC20A2B4F1EBFC30B24D" author="Guenther, A." pageNumber="3" pagination="114 - 154" refId="ref35371" refString="Guenther, A. (1864) Report on a collection of fishes made by Messrs. Dow, Godman and Salvin in Guatemala. Proceedings of the Zoological Society of London, 1864, 114 - 154." type="journal article" year="1864">
<normalizedToken id="9652A2A8775345449386CA5408B5B8A9" originalValue="Günther" pageNumber="3">Guenther</normalizedToken>
, 1864
</bibRefCitation>
</td>
</taxonomicName>
<td id="CE6DBA8DFF50D74DEEE4618B9519177D" pageNumber="3">
<bibRefCitation id="CA9BD0C15F1097EF26711344EB6B073A" author="Farias, L. P. &amp; Orti, G. &amp; Meyer, A." pageNumber="3" pagination="76 - 92" refId="ref35110" refString="Farias, L. P., Orti, G. &amp; Meyer, A. (2000) Total evidence: molecules, morphology, and the phylogenetics of cichlid fishes. Journal of Experimental Zoology, 288, 76 - 92." type="journal article" year="2000">Farias et al. 2000</bibRefCitation>
</td>
<td id="C2AC8B0BC581C341089500476C2FCBFC" pageNumber="3">
<taxonomicName id="9F54CCFC6E93BD197D3918EB204E98D6" family="Cichlidae" genus="Amphilophus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="citrinellus">Amphilophus citrinellus</taxonomicName>
</td>
</tr>
<tr id="7561D652657E0F5916243F5C916E8D8A" pageNumber="3">
<taxonomicName id="99621FCDFC68EAC0F024766D2184C764" LSID-ZBK="urn:lsid:zoobank.org:act:F07C767D-B0AE-4BED-9A60-42ED373CF5AB" authority="Gill, 1877" authorityName="Gill" authorityYear="1877" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros basilaris Gill 1877:182" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="basilaris">
<td id="CB715DCD492139DBF38B9740EFCE6A53" pageNumber="3">Heros basilaris</td>
<td id="A281A51CD6282318F9F751E81EC50C2D" pageNumber="3">
<bibRefCitation id="8527909BA34712D5B6928B7296D5736D" author="Gill, T. &amp; Bransford, J. F." pageNumber="3" pagination="175 - 218" refId="ref35271" refString="Gill, T. &amp; Bransford, J. F. (1877) Synopsis of the fishes of Lake Nicaragua. Proceedings of the Academy of Natural Sciences of Philadelphia, 29, 175 - 218." type="journal article" year="1877">Gill, 1877</bibRefCitation>
</td>
</taxonomicName>
<td id="1C4EA56F70B7E525CF1D071896CE1B6A" pageNumber="3">
Synonym of
<taxonomicName id="0B1581061E56A391924B2B2FDD22463D" family="Cichlidae" genus="Amphilophus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="citrinellus">A. citrinellus</taxonomicName>
</td>
</tr>
<tr id="EFEE14429905C391B435011CC6119997" pageNumber="3">
<taxonomicName id="8BEB97E655290E39C87BBCB7E35D92DF" LSID-ZBK="urn:lsid:zoobank.org:act:3CC6C0EE-F67A-4C3C-BC4D-7F797A1E0913" authority="Guenther, 1867" authorityName="Guenther" authorityYear="1867" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros multispinosus Günther 1867:601" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="multispinosus">
<td id="4A82B1DF3357572B47D9C17164E28E21" pageNumber="3">Heros multispinosus</td>
<td id="5C69B291B22E501FA8FA051295891503" pageNumber="3">
<bibRefCitation id="A67F6C4B87942864F97421AA7A322FD6" author="Guenther, A." pageNumber="3" pagination="600 - 604" refId="ref35410" refString="Guenther, A. (1867) Memoir on the Fishes of the States of Central America, founded upon specimens collected in the fresh and marine waters of various parts of that country by Messrs. Salvin and Godman and Capt. J. M. Dow. Proceedings of the Zoological Society of London, 1866, 600 - 604." type="journal article" year="1867">
<normalizedToken id="1C572684AF77B3F35C36DA4024229D22" originalValue="Günther" pageNumber="3">Guenther</normalizedToken>
, 1867
</bibRefCitation>
</td>
</taxonomicName>
<td id="1D39FDA72FC900F5F55F6B8AA6113DAD" pageNumber="3">
<bibRefCitation id="AB6A1F55061A319F26A051F7179EBC1B" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="47E539F94A3F13CA244C8A7D6CB53CAC" pageNumber="3">
<taxonomicName id="A19D85AA9D6B341D96326489CE48A2D1" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="multispinosus">Archocentrus multispinosus</taxonomicName>
</td>
</tr>
<tr id="8B839B54F37CDD0774D2527B0B5E55CC" pageNumber="3">
<taxonomicName id="303A4DA43B40880108B04015FD57CAC3" LSID-ZBK="urn:lsid:zoobank.org:act:A1189A04-DCEB-4EC5-85DE-5BE629152BFE" authority="Guenther, 1867" authorityName="Guenther" authorityYear="1867" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros nigrofasciatus Günther 1867:601" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="nigrofasciatus">
<td id="6BDAC3EE524B8762AEEBEF25D8ED147A" pageNumber="3">Heros nigrofasciatus</td>
<td id="976A8F3688C2FD314AC0FC7DB8F49506" pageNumber="3">
<bibRefCitation id="3B75D8DFFBEDFE80806A25A963E577BF" author="Guenther, A." pageNumber="3" pagination="600 - 604" refId="ref35410" refString="Guenther, A. (1867) Memoir on the Fishes of the States of Central America, founded upon specimens collected in the fresh and marine waters of various parts of that country by Messrs. Salvin and Godman and Capt. J. M. Dow. Proceedings of the Zoological Society of London, 1866, 600 - 604." type="journal article" year="1867">
<normalizedToken id="FA57CFAD12E2FBBF3AAA918ACEBEF699" originalValue="Günther" pageNumber="3">Guenther</normalizedToken>
, 1867
</bibRefCitation>
</td>
</taxonomicName>
<td id="92A593B01EC53C86B71CC5BF8A5C0CE7" pageNumber="3">
<bibRefCitation id="BD6E4648512D80EFD537B314D7CECA3E" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="223F7899EEB2BAA994E71B908551DB34" pageNumber="3">
<taxonomicName id="3EAFE0AF34417DEF213EFE5876A8EE56" family="Cichlidae" genus="Amatitlania" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="nigrofasciata">Amatitlania nigrofasciata</taxonomicName>
</td>
</tr>
<tr id="C22FF5F6601E07CC667FE078980C0FB4" pageNumber="3">
<taxonomicName id="2E0FCF7A8186EF05967C40DF1B255094" authority="Gill, 1877" authorityName="Gill" authorityYear="1877" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="centrarchum" subGenus="Archocentrus">
<td id="D01938030936A1FDF0870C2E06699D46" pageNumber="3">Cichlasoma (Archocentrus) centrarchum</td>
<td id="75B3636C64917D185F569D476D3E860A" pageNumber="3">
<bibRefCitation id="FD6C6C714A1F212440B684B2CD6428E9" author="Gill, T. &amp; Bransford, J. F." pageNumber="3" pagination="175 - 218" refId="ref35271" refString="Gill, T. &amp; Bransford, J. F. (1877) Synopsis of the fishes of Lake Nicaragua. Proceedings of the Academy of Natural Sciences of Philadelphia, 29, 175 - 218." type="journal article" year="1877">Gill, 1877</bibRefCitation>
</td>
</taxonomicName>
<td id="421EC13B8F9552DC7E717CE38D2855AF" pageNumber="3">Original author</td>
<td id="D58404D96B53D82697C7759AD42BF53A" pageNumber="3">
<taxonomicName id="B1FAF633298FF509DAB67C769EC1CE19" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="centrarchus">Archocentrus centrarchus</taxonomicName>
</td>
</tr>
<tr id="D16573DB1039C4DAF703D174DC96BF40" pageNumber="3">
<td id="882D5074B4D5E7DC6740AC3B3544A8BE" pageNumber="3">
<taxonomicName id="B4A1ADF61BCDDC036A6CCD2B1B884CEB" LSID-ZBK="urn:lsid:zoobank.org:act:E115CBE3-0AEE-42FA-8B3B-31DC278DE831" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros Cichlasoma spinosissimus Vaillant &amp; Pellegrin 1902:87 [3]" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spinosissimus">Heros spinosissimus</taxonomicName>
</td>
<td id="D012F3CFA0C549C7E1C4AFC253E91426" pageNumber="3">
<bibRefCitation id="1EAC3769C7650AC06B7550FDE09DB44A" author="Vaillant, L. &amp; Pellegrin, J." pageNumber="3" pagination="84 - 88" refId="ref37436" refString="Vaillant, L. &amp; Pellegrin, J. (1902) Cichlides nouveaux de l’ Amerique centrale. Bulletin du Museum National d’Histoire Naturelle, 8, 84 - 88." type="journal article" year="1902">Vaillant &amp; Pellegrin, 1902</bibRefCitation>
</td>
<td id="B699BCC6CCD9720F87CD5B4EB3D2802C" pageNumber="3">
<bibRefCitation id="3DF8258B0FCDA13515AF7DAEE2DDBA55" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="3A9E59C07FD62A33358A0EFF1E255BFD" pageNumber="3">
<taxonomicName id="D907C659115D13072AEC463D0238F557" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spinosissimus">Archocentrus spinosissimus</taxonomicName>
</td>
</tr>
<tr id="23252C434F05F8038F75D264E254ECCA" pageNumber="3">
<taxonomicName id="AA4744C9A799758926F7189B1F0730E3" LSID-ZBK="urn:lsid:zoobank.org:act:D76352FE-1F2C-4AFD-9081-678A86D6C797" authority="Regan, 1903" authorityName="Regan" authorityYear="1903" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros octofasciatus Regan 1903:417" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatus">
<td id="C0A9FA69C711F6A8BEB62A3DB4A08F96" pageNumber="3">Heros octofasciatus</td>
<td id="AABE45657925C6A9DB80EFB76A1AC4EA" pageNumber="3">
<bibRefCitation id="5B75582ADA68D132D03EE228913D25BD" author="Regan, C. T." pageNumber="3" pagination="413 - 418" refId="ref36927" refString="Regan, C. T. (1903) Descriptions de poissons nouveaux faisant partie de la collection du Musee d'Histoire Naturelle de Geneve. Revue Suisse de Zoologie, 11, 413 - 418." type="journal article" year="1903">Regan, 1903</bibRefCitation>
</td>
</taxonomicName>
<td id="07208585DDC36E7CE48EE590737C990F" pageNumber="3">
<bibRefCitation id="5CF6A162B3DB243265081E33A593C622" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="17E50E693A8CA9283E2A2E8782534CDE" pageNumber="3">
<taxonomicName id="AE406A67049C7A7D6BE86BA1A9ABD96E" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Rocio octofasciata</taxonomicName>
</td>
</tr>
<tr id="D8D58CFBE882512D670E20E2B1727745" pageNumber="3">
<taxonomicName id="E2A81212BB07A933C1662D7025295932" LSID-ZBK="urn:lsid:zoobank.org:act:3A693BD7-FD95-4532-9F14-41B963B40D09" authority="Meek, 1904" authorityName="Meek" authorityYear="1904" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma hedricki Meek 1904:208" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="hedricki">
<td id="7EC67488E728BAE4EA61AE6F65D3CE99" pageNumber="3">Cichlasoma hedricki</td>
<td id="6847D710D3AC22F4C7D6DB16D7FB8991" pageNumber="3">
<bibRefCitation id="3C37DBB4861A52659658A35D0AA06804" author="Meek, S. E." pageNumber="3" pagination="1 - 252" refId="ref36415" refString="Meek, S. E. (1904) The fresh-water fishes of Mexico north of the Isthmus of Tehuantepec. Field Columbian Museum of Natural History Publications, Zoological Series, 93, 1 - 252." type="journal article" year="1904">Meek, 1904</bibRefCitation>
</td>
</taxonomicName>
<td id="D2A3A4B32A4CCCE6026DDCA8E824047D" pageNumber="3">
Synonym of
<taxonomicName id="AF72A5FB8378C763F84B457E13AC2903" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">R. octofasciata</taxonomicName>
</td>
</tr>
<tr id="7DD51911C0500244FD64FAF96A94CEE4" pageNumber="3">
<taxonomicName id="F1CA47AF12AB6E77DEE0320DD48A68BB" authority="Pellegrin, 1904" authorityName="Pellegrin" authorityYear="1904" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="variety" species="spinosissimum" variety="immaculatum">
<td id="B9241BDDB9831A1BDD0BEF5A0D6D8BDE" pageNumber="3">Cichlasoma spinosissimum var. immaculatum</td>
<td id="05558E8A460C1D5EFC77519C95297CE2" pageNumber="3">
<bibRefCitation id="BEA7F573FE078E8B8AE9409CF3E4E693" author="Pellegrin, J." pageNumber="3" pagination="41 - 399" refId="ref36888" refString="Pellegrin, J. (1904) Contribution a l’ etude anatomique, biologique et taxinomique des poissons de la famille des Cichlides. Memoires de la Societe Zoologique de la France, 16, 41 - 399." type="journal article" year="1904">Pellegrin, 1904</bibRefCitation>
</td>
</taxonomicName>
<td id="CF39A7D0B66BD4C956DD3C1011FEBAC9" pageNumber="3">
<bibRefCitation id="669E9F34C69BEE8272B2990C59FDBCE1" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan 1908</bibRefCitation>
</td>
<td id="B81BB4A9A7264918157951E279D2D7D9" pageNumber="3">
Synonym of
<taxonomicName id="3FA6C30BEB6E9CFEB5236D55DAE089A7" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cr. spilurus</taxonomicName>
</td>
</tr>
<tr id="61BAB11E3AD6E12B559AF377C8CC9341" pageNumber="3">
<taxonomicName id="A607E9460EE9512B9DB029EAE139487D" LSID-ZBK="urn:lsid:zoobank.org:act:E49CFF2D-9CD3-43A0-9F1B-A57FF7E4F957" authority="Meek, 1907" authorityName="Meek" authorityYear="1907" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma granadense Meek 1907:121" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="granadense">
<td id="24DC7828A5AD8ECA3C8AF63835E57C26" pageNumber="3">Cichlasoma granadense</td>
<td id="162DAAB9C31E8F8293129F768E26314F" pageNumber="3">Meek, 1907</td>
</taxonomicName>
<td id="F1BFCC9A53825318AFF55CB75711C6C6" pageNumber="3">
Synonym of
<taxonomicName id="389DE4FF96E796D7B67CC6410972E457" family="Cichlidae" genus="Amphilophus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="citrinellus">Am. citrinellus</taxonomicName>
</td>
</tr>
<tr id="52AFC510C5668AC29DCCFDDCB4E1E240" pageNumber="3">
<taxonomicName id="1097E977EDD7F3629DDE0AF38BFF9A45" authority="Regan, 1908" authorityName="Regan" authorityYear="1908" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="septemfasciatum">
<td id="6882A2E47681B267B39A0F019E136F14" pageNumber="3">Cichlasoma septemfasciatum</td>
<td id="98D6503B4F23683D77181C0ACCB26CA5" pageNumber="3">
<bibRefCitation id="A82079C01592229DAE9F6FE6E4941F07" author="Regan, C. T." pageNumber="3" pagination="455 - 464" refId="ref37046" refString="Regan, C. T. (1908) A collection of freshwater fishes made by Mr. C. F. Underwood in Costa Rica. Annual Magazine of Natural History, 2, 455 - 464." type="journal article" year="1908">Regan, 1908</bibRefCitation>
</td>
</taxonomicName>
<td id="CFC6D766C450F5C4AB6A8E9CAD30FAB6" pageNumber="3">
<bibRefCitation id="CD99F53FD1D024F90BCC1CFD9B294F39" author="Loiselle, P. V." pageNumber="3" pagination="1 - 8" refId="ref36179" refString="Loiselle, P. V. (1997) Diagnosis of two new cichlids from the Rio Sixaola drainage, Costa Rica. Buntbarsche Bulletin, 180, 1 - 8." type="journal article" year="1997">Loiselle 1997</bibRefCitation>
</td>
<td id="9D47514FA85637DEC6BB543A74817E1C" pageNumber="3">
<taxonomicName id="0665B6CE135F482318DCF0803866BD6F" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="septemfasciatus">Cryptoheros septemfasciatus</taxonomicName>
</td>
</tr>
<tr id="9FFACAEB7D06FF82DD109715611FBB75" pageNumber="3">
<taxonomicName id="342DEA7AEE5FBABAC2351EFA9BC59996" LSID-ZBK="urn:lsid:zoobank.org:act:E7A510F9-CF8C-4181-A6CE-DD7BA7F860F2" authority="Regan, 1909" authorityName="Regan" authorityYear="1909" family="Cichlidae" genus="Cichlosoma" kingdom="Animalia" lsidName-ZBK="Cichlosoma biocellatum Regan 1909:234" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="biocellatum">
<td id="C2778FA741F3E12F78B1B48FBE182D7F" pageNumber="3">Cichlosoma biocellatum</td>
<td id="663A71FCD1DE4F3113263C8EFF2CF938" pageNumber="3">
<bibRefCitation id="6323D098AFE5A97270FACE70CE63B5E7" author="Regan, C. T." pageNumber="3" pagination="234 - 235" refId="ref37085" refString="Regan, C. T. (1909) Descriptions of three new freshwater fishes from South America, presented to the British Museum by Herr J. Paul Arnold. Annual Magazine of Natural History, 3, 234 - 235." type="journal article" year="1909">Regan, 1909</bibRefCitation>
</td>
</taxonomicName>
<td id="3EA72A146602BB0594467D1C5F3DF48F" pageNumber="3">
Synonym of
<taxonomicName id="D50B1FB862EF35B3A56A88A7E298CA79" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">R. octofasciata</taxonomicName>
</td>
</tr>
<tr id="03CD6BFCD22C03DC7AED04AA9DC9461A" pageNumber="3">
<td id="829A55825D5A4ABF88A9398F527DD1EE" pageNumber="3">
<taxonomicName id="CB227CD52063339583EA7FDAB891DE90" LSID-ZBK="urn:lsid:zoobank.org:act:2411EEF8-6445-445C-AFA1-B6C5A08A5161" family="Cichlidae" genus="Neetroplus" kingdom="Animalia" lsidName-ZBK="Neetroplus panamensis Meek &amp; Hildebrand 1913:90" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="panamensis">Neetroplus panamensis</taxonomicName>
</td>
<td id="E5831998AAA4D8F80F98849E6091C209" pageNumber="3">
<bibRefCitation id="71494D44FD6DDDF3B34275538B9E3F5E" author="Meek, S. E. &amp; Hildebrand, S. F." pageNumber="3" pagination="77 - 91" refId="ref36487" refString="Meek, S. E. &amp; Hildebrand, S. F. (1913) New species of fishes from Panama. Field Columbian Museum of Natural History Publications, Zoological Series, 166, 77 - 91." type="journal article" year="1913">Meek &amp; Hildebrand, 1913</bibRefCitation>
</td>
<td id="7174258FD400AB6B079BE24E74AB5CC7" pageNumber="3">
<bibRefCitation id="4A5DB8FC783B7A2A254B5C8394DCA48A" author="Kullander, S. O." pageNumber="3" pagination="605 - 654" refId="ref35967" refString="Kullander, S. O. (2003) Family Cichlidae (Cichlids). In: Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J., Jr. (eds.). Checklist of the Freshwater Fishes of South and Central America. Edipucrs, Porto Alegre, pp. 605 - 654." type="book chapter" year="2003">Kullander 2003</bibRefCitation>
</td>
<td id="811B92734AF439EFA064CAE2DD131112" pageNumber="3">
<taxonomicName id="1A97E7A300918BAACFBCC04D8CE5DA29" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="panamensis">Cryptoheros panamensis</taxonomicName>
</td>
</tr>
<tr id="7542E4A3D77977A5B3B99843FC6D8FE5" pageNumber="3">
<taxonomicName id="04CA71AF71E9DBC43FA97888C54FBC9A" LSID-ZBK="urn:lsid:zoobank.org:act:AA1787B8-6C57-46B3-B745-27ED2F0BC0D8" authority="Fowler, 1932" authorityName="Fowler" authorityYear="1932" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma cutteri Fowler 1932:380" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">
<td id="15F4220E0BFFE60E527F2F35D4158D37" pageNumber="3">Cichlasoma cutteri</td>
<td id="DA5BBF90464EFEAC194A11F13F861964" pageNumber="3">
<bibRefCitation id="2FCCE41A7A7F34FADA7D71E3475B8FCA" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler, 1932</bibRefCitation>
</td>
</taxonomicName>
<td id="8F1903D3109B4F4136805BB9172A708A" pageNumber="3">
<taxonomicName id="8546473600DE3A1927DB40DAF29B2F07" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">Cryptoheros cutteri</taxonomicName>
</td>
</tr>
<tr id="8746430D5887E28A21325DACD93E0B79" pageNumber="3">
<taxonomicName id="BC92414B88358A3D1B14E7F1B60B3E84" LSID-ZBK="urn:lsid:zoobank.org:act:66BF6A4B-52CB-4DF1-88E4-19D5998880F0" authority="Bussing, 1974" authorityName="Bussing" authorityYear="1974" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma sajica Bussing 1974:30" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="sajica">
<td id="54503333383C623787E1ED4A1165DC80" pageNumber="3">Cichlasoma sajica</td>
<td id="9F36D7EEA7F36C30ACFB4511F71A8DA4" pageNumber="3">
<bibRefCitation id="9F08788BF956751C86023BB513720204" author="Bussing, W. A." pageNumber="3" pagination="29 - 49" refId="ref34626" refString="Bussing, W. A. (1974) Two new species of cichlid fishes, Cichlasoma sajica and C. diquis, from southeastern Costa Rica. Revista de Biologia Tropical, 22, 29 - 49." type="journal article" year="1974">Bussing, 1974</bibRefCitation>
</td>
</taxonomicName>
<td id="424CE0CAF1FE4ACAFABF255706473254" pageNumber="3">
<bibRefCitation id="A7F4EA9B4F8179D353844CDD287AF74E" author="Bussing, W. A." pageNumber="3" pagination="1 - 468" refId="ref34754" refString="Bussing, W. A. (1998) Peces de las aguas continentales de Costa Rica. Revista de Biologia Tropical, 46, 1 - 468." type="journal article" year="1998">Bussing 1998</bibRefCitation>
</td>
<td id="7062618669CBBB3BE86806D05DF14D07" pageNumber="3">
<taxonomicName id="361F3D0F4F48848962F330AA42E95D69" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="sajica">Cryptoheros sajica</taxonomicName>
</td>
</tr>
<tr id="F9A691600622BE5BC9DCA0FEC07722DF" pageNumber="3">
<taxonomicName id="63DAF8AF555364257389DEB5FF21C015" LSID-ZBK="urn:lsid:zoobank.org:act:BE078628-41E1-4939-87E9-491357D791E8" authority="Allgayer, 1994" authorityName="Allgayer" authorityYear="1994" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" lsidName-ZBK="Archocentrus nanoluteus Allgayer 1994" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="nanoluteus">
<td id="8A822DEE19BFCB325F4C36944102229B" pageNumber="3">Archocentrus nanoluteus</td>
<td id="BE80515BE5C4A03141E6CA3C7B4A399B" pageNumber="3">
<bibRefCitation id="48EEEA12922A47A815F9CC1FBFDA0DF5" author="Allgayer, R." pageNumber="3" pagination="6 - 24" refId="ref34494" refString="Allgayer, R. (1994) Description d’une espece nouvelle du genre Archocentrus. Revue Francaise des Cichlidophiles, 135, 6 - 24." type="journal article" year="1994">Allgayer, 1994</bibRefCitation>
</td>
</taxonomicName>
<td id="A14BB54D53451756C63C167D11A3BC82" pageNumber="3">Original author</td>
<td id="0FB624FEE79735DB4FCBE0A83AA73200" pageNumber="3">
<taxonomicName id="DA65A2B163923A87416EA8005BF957DF" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="nanoluteus">Cryptoheros nanoluteus</taxonomicName>
</td>
</tr>
<tr id="D9E044CB64EBEF3F4C8B0A4C0BCB9707" pageNumber="3">
<taxonomicName id="4CC6F1100DF52505BDBE9895EE005623" LSID-ZBK="urn:lsid:zoobank.org:act:F542B413-FA04-4C6E-8A37-46EBA4344080" authority="Loiselle, 1997" authorityName="Loiselle" authorityYear="1997" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" lsidName-ZBK="Archocentrus myrnae Loiselle 1997:3" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="myrnae">
<td id="34EBD658753BA96E8FF65CB1C850360A" pageNumber="3">Archocentrus myrnae</td>
<td id="687830A06E81C32F81E8FF2EC20A11DC" pageNumber="3">
<bibRefCitation id="6EDBCF5839EF5C89AEF94BCD028DDDB3" author="Loiselle, P. V." pageNumber="3" pagination="1 - 8" refId="ref36179" refString="Loiselle, P. V. (1997) Diagnosis of two new cichlids from the Rio Sixaola drainage, Costa Rica. Buntbarsche Bulletin, 180, 1 - 8." type="journal article" year="1997">Loiselle, 1997</bibRefCitation>
</td>
</taxonomicName>
<td id="B022522566016DAABAF594505A4E74A8" pageNumber="3">Original author</td>
<td id="6D2BEEF991F8D3B7E6D6367F94199530" pageNumber="3">
<taxonomicName id="A88B9CF10D2AD9C2A14652752D23870C" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="myrnae">Cryptoheros myrnae</taxonomicName>
</td>
</tr>
<tr id="38089CA3120EACA9A17B371E09F9FDA7" pageNumber="3">
<taxonomicName id="2D138980DA7E4DEC635BB2F0F805B158" LSID-ZBK="urn:lsid:zoobank.org:act:B8A94766-C505-4E59-98D6-19AEF7E7B80A" authority="Allgayer, 2001" authorityName="Allgayer" authorityYear="2001" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" lsidName-ZBK="Cryptoheros altoflavus Allgayer 2001:16" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="altoflavus">
<td id="DDD4263E959B38954D2EF3C033825483" pageNumber="3">Cryptoheros altoflavus</td>
<td id="645F4CB178647EDB709B45335AAF1873" pageNumber="3">
<bibRefCitation id="E1D47DA23E9F8DC29F5CC061DCAAC046" author="Allgayer, R." pageNumber="3" pagination="13 - 20" refId="ref34520" refString="Allgayer, R. (2001) Description d'un genre nouveau, Cryptoheros, d'Amerique Centrale et d'une espece nouvelle du Panama (Pisces: Cichlidae). L’An Cichlide, 1, 13 - 20." type="journal article" year="2001">Allgayer, 2001</bibRefCitation>
</td>
</taxonomicName>
<td id="A27B8AFAF0798BD9D5605FD5C2A9119D" pageNumber="3">
<bibRefCitation id="195E985A606479B34831DDEF32C507D7" author="Kullander, S. O." pageNumber="3" pagination="605 - 654" refId="ref35967" refString="Kullander, S. O. (2003) Family Cichlidae (Cichlids). In: Reis, R. E., Kullander, S. O. &amp; Ferraris, C. J., Jr. (eds.). Checklist of the Freshwater Fishes of South and Central America. Edipucrs, Porto Alegre, pp. 605 - 654." type="book chapter" year="2003">Kullander 2003</bibRefCitation>
</td>
<td id="1DED5182D922DAE88AB8936561B322E5" pageNumber="3">Valid</td>
</tr>
</table>
</paragraph>
<paragraph id="D291BDF07D0018CE87DDE502B33E0FAB" pageNumber="6">
<pageBreakToken id="476307E625F045E3E074D6BD4F14E42D" pageNumber="3">To</pageBreakToken>
complicate things further, variation within some
<taxonomicName id="A361AA7F015F8890DAA055DC6881CF6E" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
species has not been evaluated before.
<bibRefCitation id="AD45B7423D43A63081378585D54102D6" author="Hubbs, C. L." pageNumber="3" pagination="157 - 287" refId="ref35613" refString="Hubbs, C. L. (1936) Fishes of the Yucatan Peninsula. Carnegie Institution of Washington Publications, 457, 157 - 287." type="journal article" year="1936">Hubbs (1936)</bibRefCitation>
observed tantalizing meristic and morphometric differences among populations of
<taxonomicName id="F21426A2FB8DD1DCF74F087AAEF67DAE" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatum">Cichlasoma octofasciatum</taxonomicName>
, but he chose not to describe any subspecies because, in his material, geographic differences were confounded with differences in body size.
</paragraph>
<paragraph id="ECD83B2236CB010DE1B77770D5194073" pageNumber="6">
Considering the potential for undetected biodiversity within presently recognized species, especially those more widely distributed, lectotypes are designated in the present work:
<taxonomicName id="DDD6AFBE03A2735F7B08D19E434C912C" family="Cichlidae" genus="Amatitlania" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="nigrofasciata">Amatitlania nigrofasciata</taxonomicName>
,
<taxonomicName id="71AD0FE77EF82CCD7754457CBAABBB63" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spinosissimus">Archocentrus spinosissimus</taxonomicName>
,
<taxonomicName id="8CD3F2643B2806EE40A82E09ED7ABB85" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="septemfasciatus">Cryptoheros septemfasciatus</taxonomicName>
,
<taxonomicName id="79ABAAC6990392E7155A900852089F3F" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cr. spilurus</taxonomicName>
, and
<taxonomicName id="044C81B3B5D79B74968DA492B506F46B" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">Rocio octofasciata</taxonomicName>
.
<taxonomicName id="58FCD0FE135973FD3BE46E0050531398" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="immaculatum">Cichlasoma immaculatum</taxonomicName>
is shown to be a synonym of
<taxonomicName id="BA65A09E5E8D81C7617194216B3E531E" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cryptoheros spilurus</taxonomicName>
and not of
<taxonomicName id="6472A16DFAA2AC36E4CF04CEC9316FCC" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spinosissimus">Archocentrus spinosissimus</taxonomicName>
, although it was first described as a “variety” of the latter.
</paragraph>
<paragraph id="30F0A2C534BAA120867815C86BE3B827" pageNumber="6">
The present revision treats all nominal species ever assigned to the genus
<taxonomicName id="D2D2DFEB45AF1A39844724F36DECC73D" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
, as well as species that have appeared in the same clade as (or as sister group to)
<taxonomicName id="BD37C24F86C690902236F3DFFAD5DC11" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="centrarchus">Archocentrus centrarchus</taxonomicName>
in relevant phylogenetic analyses (
<bibRefCitation id="DA817A04FE30BBECD07734D13DD7B833" author="Roe, K. J. &amp; Conkel, D. &amp; Lydeard, C." pageNumber="3" pagination="366 - 376" refId="ref37128" refString="Roe, K. J., Conkel, D. &amp; Lydeard, C. (1997) Molecular systematics of Middle American cichlid fishes and the evolution of trophic-types in ‘ Cichlasoma (Amphilophus) ’ and ‘ C. (Thorichthys) ’. Molecular Phylogenetics and Evolution, 7, 366 - 376." type="journal article" year="1997">Roe et al. 1997</bibRefCitation>
,
<bibRefCitation id="5DC6F934DB104E73BCA89BAA9BF0FA23" author="Martin, P. A. &amp; Bermingham, E." pageNumber="3" pagination="192 - 203" refId="ref36305" refString="Martin, P. A. &amp; Bermingham, E. (1998) Systematics and evolution of lower Central American cichlids inferred from analysis of cytochrome b gene sequences. Molecular Phylogenetics and Evolution, 9, 192 - 203." type="journal article" year="1998">Martin &amp; Bermingham 1998</bibRefCitation>
,
<bibRefCitation id="B7A917F7718047756A6E9452F4A448DC" author="Farias, L. P. &amp; Orti, G. &amp; Meyer, A." pageNumber="3" pagination="76 - 92" refId="ref35110" refString="Farias, L. P., Orti, G. &amp; Meyer, A. (2000) Total evidence: molecules, morphology, and the phylogenetics of cichlid fishes. Journal of Experimental Zoology, 288, 76 - 92." type="journal article" year="2000">Farias et al. 2000</bibRefCitation>
). Six new species are described and a seventh is resurrected from synonymy.
<taxonomicName id="0B1CF5A27E5C0B23164C3BA5A9A01A8F" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
,
<taxonomicName id="FCC5EAC1A980F5810CBD361DBEB94323" LSID-ZBK="urn:lsid:zoobank.org:act:273F29E5-91F4-45BE-8447-659BDE8939E2" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" lsidName-ZBK="Cryptoheros Allgayer 2001" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Cryptoheros</taxonomicName>
, and
<taxonomicName id="DFD9266C27832B87B94219631A67ADDB" LSID-ZBK="urn:lsid:zoobank.org:act:7A9BA3FB-1462-4C73-A40A-F9A660B3AA90" class="Malacostraca" family="Homolidae" genus="Hypsophrys" kingdom="Animalia" lsidName-ZBK="Hypsophrys Agassiz 1859:408" order="Decapoda" pageNumber="3" phylum="Arthropoda" rank="genus">Hypsophrys</taxonomicName>
are redefined, and two new genera (
<taxonomicName id="7731F8DEAC84A72B2AA3C2B2D8A7961C" family="Cichlidae" genus="Amatitlania" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Amatitlania</taxonomicName>
and
<taxonomicName id="375F7B2A5FE07D6255760D93E78103F1" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Rocio</taxonomicName>
) are diagnosed.
</paragraph>
<paragraph id="ECE6EF7A8BA817110B10946118D0DFE9" pageNumber="6">Genus-level decisions in this work are supported by a phylogeny (Schmitter-Soto, in press), published separately, which includes most character illustrations and analysis.</paragraph>
</subSubSection>
</treatment>
</document>
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