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<document id="6689E9659406549BE89859D6C5C8C77B" ID-CLB-Dataset="31806" ID-DOI="10.5281/zenodo.893555" ID-GBIF-Dataset="4a2efe36-c48b-456b-9223-7b5793025a72" ID-ISSN="0035-418" ID-Zenodo-Dep="893555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508361162557" checkinUser="plazi" docAuthor="Peter J. Schwendinger" docDate="2017" docId="4C30A452FFFFFFE8BB55F88939D7FBDE" docLanguage="en" docName="rsz_124_2_11_Schwendinger.pdf" docOrigin="Revue suisse de Zoologie 124 (2)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Liphistius linang Peter J. Schwendinger 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="422" masterDocId="B009DC2AFFE4FFF7BA6FFFF13818FFE3" masterDocTitle="A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1" masterLastPageNumber="445" masterPageNumber="391" pageNumber="418" updateTime="1698469770161" updateUser="ExternalLinkService">
<mods:mods id="F0CABADFF48913FF8267AFD3A8D1CD24" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2B9B13FBA23B6FCC2FFA87CC91594C29">
<mods:title id="1042ADE1EFB99155751824542860A3FD">A revision of the trapdoor spider genus Liphistius (Mesothelae: Liphistiidae) in peninsular Malaysia; part 1</mods:title>
</mods:titleInfo>
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<mods:namePart id="00A632B8FD485D5D11355491E07D0AA8">Peter J. Schwendinger</mods:namePart>
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<mods:typeOfResource id="30AED8818A62F36D64B94C54135300C7">text</mods:typeOfResource>
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<mods:title id="08C421A9E58C64B590F12D2D9DFE8761">Revue suisse de Zoologie</mods:title>
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<mods:part id="0C9A01D2D8A1CC87A8DCE7CFF4759DB3">
<mods:date id="0E7E3A9E26C5846E06D81A5B33148C10">2017</mods:date>
<mods:detail id="98637D156B663EED58C92D06A070F9DE" type="pubDate">
<mods:number id="4EF482011205D99964BB5479B40F7DE7">2017-09-30</mods:number>
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<mods:number id="6869676EF87646B09575C2E7C937332B">124</mods:number>
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<mods:number id="1CE357D42F36B28D1B4B52BA5B3318C7">2</mods:number>
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<mods:start id="9D7E09BE0E9FD1EB56A2A4AFCA572102">391</mods:start>
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<mods:classification id="88C29A3E862692BB2BFB6315E0E21D58">journal article</mods:classification>
<mods:identifier id="505CD0763E0390EBB9F68DF6B9C691AB" type="CLB-Dataset">31806</mods:identifier>
<mods:identifier id="64F50F9EB5EB464B623524AE0114705D" type="DOI">10.5281/zenodo.893555</mods:identifier>
<mods:identifier id="66E6C946690DDA580F6C78DCA39FFE85" type="GBIF-Dataset">4a2efe36-c48b-456b-9223-7b5793025a72</mods:identifier>
<mods:identifier id="9F7B16B43EE5E7485212017A503044B4" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="7FF3AB04C5941596F2C2E0639C29E8E0" type="Zenodo-Dep">893555</mods:identifier>
</mods:mods>
<treatment id="4C30A452FFFFFFE8BB55F88939D7FBDE" ID-DOI="http://doi.org/10.5281/zenodo.6042372" ID-GBIF-Taxon="135450769" ID-Zenodo-Dep="6042372" LSID="urn:lsid:plazi:treatment:4C30A452FFFFFFE8BB55F88939D7FBDE" httpUri="http://treatment.plazi.org/id/4C30A452FFFFFFE8BB55F88939D7FBDE" lastPageId="31" lastPageNumber="422" pageId="27" pageNumber="418">
<subSubSection id="8C8346CFFFFFFFECBB55F88939E7F84D" pageId="27" pageNumber="418" type="nomenclature">
<paragraph id="C4261544FFFFFFECBB55F8893A4AF873" blockId="27.[314,594,1912,1966]" box="[314,594,1912,1936]" pageId="27" pageNumber="418">
<heading id="9F6EA228FFFFFFECBB55F8893A4AF873" bold="true" box="[314,594,1912,1936]" centered="true" fontSize="10" level="3" pageId="27" pageNumber="418" reason="4">
<emphasis id="F6EDC956FFFFFFECBB55F8893A4AF873" bold="true" box="[314,594,1912,1936]" pageId="27" pageNumber="418">
<taxonomicName id="03996EC7FFFFFFECBB55F88939EDF86C" ID-CoL="6QJ5F" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[314,501,1912,1935]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
<emphasis id="F6EDC956FFFFFFECBB55F88939EDF86C" bold="true" box="[314,501,1912,1935]" italics="true" pageId="27" pageNumber="418">Liphistius linang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFFFFECBB93F8893A4AF873" box="[508,594,1912,1936]" pageId="27" pageNumber="418" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C4261544FFFFFFECBBE3F86639E7F84D" blockId="27.[314,594,1912,1966]" box="[396,511,1943,1966]" pageId="27" pageNumber="418">
<figureCitation id="5CA209C1FFFFFFECBBE3F86639E7F84D" box="[396,511,1943,1966]" captionStart-0="Fig. 15" captionStart-1="Fig. 16" captionStartId-0="28.[153,188,1792,1813]" captionStartId-1="30.[151,186,1755,1776]" captionTargetBox-0="[154,1431,259,1731]" captionTargetBox-1="[153,1440,257,1712]" captionTargetId-0="figure@28.[148,1439,254,1752]" captionTargetId-1="figure@30.[149,1458,241,1736]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionText-0="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." captionText-1="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri-0="https://zenodo.org/record/1021969/files/figure.png" httpUri-1="https://zenodo.org/record/1021971/files/figure.png" pageId="27" pageNumber="418">Figs 15-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFFFFECBAF8F8373C2EFCD4" pageId="27" pageNumber="418" type="materials_examined">
<paragraph id="C4261544FFFFFFECBAF8F8373C2EFCD4" blockId="27.[151,756,1990,2078]" lastBlockId="27.[831,1437,702,824]" pageId="27" pageNumber="418">
<materialsCitation id="74F11F19FFFFFFECBAF8F8373C2EFCD4" ID-GBIF-Occurrence="1675315701" collectingDate="1999-01-12" collectingDateMax="2002-02-12" collectingDateMin="1999-01-12" collectionCode="MHNG" collectorName="P. J. Schwendinger." country="Malaysia" elevation="50" latitude="5.742778" location="Jeram Linang Waterfall" longLatPrecision="21" longitude="102.374725" pageId="27" pageNumber="418" specimenCount="11" specimenCount-female="4" specimenCount-male="4" stateProvince="Kelantan" typeStatus="holotype">
<emphasis id="F6EDC956FFFFFFECBAF8F83738FAF83D" bold="true" box="[151,226,1990,2014]" pageId="27" pageNumber="418">
<typeStatus id="1B22ABE6FFFFFFECBAF8F83738C5F83D" box="[151,221,1990,2014]" pageId="27" pageNumber="418">Types</typeStatus>
:
</emphasis>
<collectionCode id="A2888D81FFFFFFECBA85F8373927F83D" box="[234,319,1990,2014]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="27" pageNumber="418">MHNG</collectionCode>
; male
<typeStatus id="1B22ABE6FFFFFFECBBE8F83739FCF83E" box="[391,484,1990,2013]" pageId="27" pageNumber="418" type="holotype">holotype</typeStatus>
(matured end
<collectingDate id="A063CA6CFFFFFFECB8ECF8373AFFF83D" box="[643,743,1990,2014]" pageId="27" pageNumber="418" value="2001-12">
<date id="B0273384FFFFFFECB8ECF8373AFFF83D" box="[643,743,1990,2014]" pageId="27" pageNumber="418" value="2001-12">XII.2001</date>
</collectingDate>
),
<specimenCount id="D29FDECDFFFFFFECBAF8F81738F2F81F" box="[151,234,2021,2046]" pageId="27" pageNumber="418" type="male">4 male</specimenCount>
<typeStatus id="1B22ABE6FFFFFFECBA95F817397AF81D" box="[250,354,2022,2046]" pageId="27" pageNumber="418" type="paratype">paratypes</typeStatus>
(matured
<collectingDate id="A063CA6CFFFFFFECBB8AF8173AB0F81D" box="[485,680,2022,2046]" isEnumeration="true" pageId="27" pageNumber="418">
<date id="B0273384FFFFFFECBB8AF8173A35F81D" box="[485,557,2022,2046]" pageId="27" pageNumber="418" value="1999-09">14.IX.</date>
,
<date id="B0273384FFFFFFECB82DF8173AB0F81D" box="[578,680,2022,2046]" pageId="27" pageNumber="418" value="1999-10-01">1.X.1999</date>
</collectingDate>
, early
<collectingDate id="A063CA6CFFFFFFECBAF8F7F438ECF7FE" box="[151,244,2053,2077]" pageId="27" pageNumber="418" value="2000-11">
<date id="B0273384FFFFFFECBAF8F7F438ECF7FE" box="[151,244,2053,2077]" pageId="27" pageNumber="418" value="2000-11">XI.2000</date>
</collectingDate>
, end
<collectingDate id="A063CA6CFFFFFFECBB25F7F439B7F7FE" box="[330,431,2053,2077]" pageId="27" pageNumber="418" value="2001-12">
<date id="B0273384FFFFFFECBB25F7F439B7F7FE" box="[330,431,2053,2077]" pageId="27" pageNumber="418" value="2001-12">XII.2001</date>
</collectingDate>
) and
<specimenCount id="D29FDECDFFFFFFECB866F7F43A6EF7FF" box="[521,630,2053,2077]" pageId="27" pageNumber="418" type="female">4 female</specimenCount>
<typeStatus id="1B22ABE6FFFFFFECB8E4F7F73AEBF7FD" box="[651,755,2054,2078]" pageId="27" pageNumber="418" type="paratype">paratypes</typeStatus>
(including
<typeStatus id="1B22ABE6FFFFFFECB9ADFD4F3C05FD36" box="[962,1053,702,725]" pageId="27" pageNumber="418" type="allotype">allotype</typeStatus>
, moulted
<collectingDate id="A063CA6CFFFFFFECBEF1FD4E3D0BFD34" box="[1182,1299,703,727]" pageId="27" pageNumber="418" value="2002-02-12">
<date id="B0273384FFFFFFECBEF1FD4E3D0BFD34" box="[1182,1299,703,727]" pageId="27" pageNumber="418" value="2002-02-12">12.II.2002</date>
</collectingDate>
);
<collectingCountry id="BC8E55D4FFFFFFECBF5DFD4F3D80FD36" box="[1330,1432,702,725]" name="Malaysia" pageId="27" pageNumber="418">Malaysia</collectingCountry>
,
<collectingRegion id="065DDBA6FFFFFFECB950FD2E3BBBFD15" box="[831,931,735,758]" country="Malaysia" name="Kelantan" pageId="27" pageNumber="418">Kelantan</collectingRegion>
, about
<locationDeviation id="D5940C66FFFFFFECB994FD2E3CEDFD15" box="[1019,1269,735,759]" pageId="27" pageNumber="418">
<quantity id="0361B8A1FFFFFFECB994FD2E3C5BFD15" box="[1019,1091,735,758]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="27" pageNumber="418" unit="km" value="15.0">15 km</quantity>
E of Machang
</locationDeviation>
,
<location id="C146439FFFFFFFECBF6BFD113BBDFCF5" LSID="urn:lsid:plazi:treatment:4C30A452FFFFFFE8BB55F88939D7FBDE:C146439FFFFFFFECBF6BFD113BBDFCF5" country="Malaysia" latitude="5.742778" longLatPrecision="21" longitude="102.374725" name="Jeram Linang Waterfall" pageId="27" pageNumber="418" stateProvince="Kelantan">Jeram Linang Waterfall</location>
,
<geoCoordinate id="A1AD7383FFFFFFECB9DAFD0E3C36FCF5" box="[949,1070,767,790]" direction="north" orientation="latitude" pageId="27" pageNumber="418" precision="15" value="5.742778">5°44’34”N</geoCoordinate>
,
<geoCoordinate id="A1AD7383FFFFFFECBE51FCF13CD6FCFB" box="[1086,1230,768,792]" direction="east" orientation="longitude" pageId="27" pageNumber="418" precision="15" value="102.374725">102°22’29”E</geoCoordinate>
,
<elevation id="4FB4F277FFFFFFECBEB0FD0E3D06FCF4" box="[1247,1310,767,791]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="418" unit="m" value="50.0">
<quantity id="0361B8A1FFFFFFECBEB0FD0E3D06FCF4" box="[1247,1310,767,791]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="418" unit="m" value="50.0">50 m</quantity>
</elevation>
;
<collectingDate id="A063CA6CFFFFFFECBF41FCF13D80FCFB" box="[1326,1432,768,792]" pageId="27" pageNumber="418" value="1999-01-12">
<date id="B0273384FFFFFFECBF41FCF13D80FCFB" box="[1326,1432,768,792]" pageId="27" pageNumber="418" value="1999-01-12">12.I.1999</date>
</collectingDate>
; leg.
<collectorName id="696C7092FFFFFFECB902FCD03C2EFCD4" box="[877,1078,800,824]" pageId="27" pageNumber="418">P.J. Schwendinger.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFFFFECB950FCA13C1EFC4B" pageId="27" pageNumber="418" type="etymology">
<paragraph id="C4261544FFFFFFECB950FCA13C1EFC4B" blockId="27.[831,1436,848,936]" pageId="27" pageNumber="418">
<emphasis id="F6EDC956FFFFFFECB950FCA13BDCFC8B" bold="true" box="[831,964,848,872]" pageId="27" pageNumber="418">Etymology:</emphasis>
Name in apposition, taken from the Malay name of the
<typeStatus id="1B22ABE6FFFFFFECB9B0FC833C16FC69" box="[991,1038,882,906]" pageId="27" pageNumber="418">type</typeStatus>
locality: jeram (= rapids),
<taxonomicName id="03996EC7FFFFFFECBF36FC813D84FC64" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1369,1436,880,903]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="linang">linang</taxonomicName>
(tearful, weeping).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFFFFECB950FC303D88FA21" pageId="27" pageNumber="418" type="diagnosis">
<paragraph id="C4261544FFFFFFECB950FC303D88FA21" blockId="27.[831,1437,961,1475]" pageId="27" pageNumber="418">
<emphasis id="F6EDC956FFFFFFECB950FC303BAFFC3B" bold="true" box="[831,951,961,984]" pageId="27" pageNumber="418">Diagnosis:</emphasis>
Medium-sized, bicoloured species in both sexes. Similar to
<taxonomicName id="03996EC7FFFFFFECBE7EFC123C98FC19" authorityName="Platnick &amp; Sedgwick" authorityYear="1984" box="[1041,1152,995,1018]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="418" phylum="Arthropoda" rank="species" species="tioman">
<emphasis id="F6EDC956FFFFFFECBE7EFC123C98FC19" box="[1041,1152,995,1018]" italics="true" pageId="27" pageNumber="418">L. tioman</emphasis>
</taxonomicName>
, males distinguished by retroventral part of bipartite tibial apophysis carrying longer and bent megaspines; retrodorsal part of tibial apophysis developed as a small tubercle carrying a weaker megaspine (
<figureCitation id="5CA209C1FFFFFFECBE4FFB953C9FFB98" box="[1056,1159,1124,1147]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15A</figureCitation>
, G-I cf.
<figureCitation id="5CA209C1FFFFFFECBE9FFB953D4EFB9E" box="[1264,1366,1124,1149]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11A</figureCitation>
, G-I); tegulum with wide, distinctly dentate proximal edge (
<figureCitation id="5CA209C1FFFFFFECB928FB543BB1FB5F" box="[839,937,1189,1212]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15D</figureCitation>
cf.
<figureCitation id="5CA209C1FFFFFFECB9BBFB543C2DFB5D" box="[980,1077,1189,1214]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11B</figureCitation>
); contrategulum without (or with very indistinct) ventral process, its distal edge finely dentate instead of ribbed, prodorsally with a wide gap before reaching hook-shaped dorsal apex (
<figureCitation id="5CA209C1FFFFFFECBF68FAF63D70FAFD" box="[1287,1384,1287,1310]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15B</figureCitation>
, E-F cf.
<figureCitation id="5CA209C1FFFFFFECB90CFAD93BFAFAA2" box="[867,994,1320,1345]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11C, F</figureCitation>
); para-embolic plate large, distally wide, with coarsely dentate margins (
<figureCitation id="5CA209C1FFFFFFECBEFDFAB83CC5FA83" box="[1170,1245,1353,1376]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15</figureCitation>
A-F cf.
<figureCitation id="5CA209C1FFFFFFECBF59FAB83D98FA82" box="[1334,1408,1353,1377]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11</figureCitation>
C- E); dorsal wall of sclerotised part of embolus proper not ending in protruding lobe, as wide as ventral wall (
<figureCitation id="5CA209C1FFFFFFECBF1EFA7B3B76FA21" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="27" pageNumber="418">Fig. 15B</figureCitation>
, E-F cf.
<figureCitation id="5CA209C1FFFFFFECB9A5FA5A3C0BFA20" box="[970,1043,1451,1475]" captionStart="Fig. 11" captionStartId="21.[151,186,1705,1726]" captionTargetBox="[158,1428,255,1634]" captionTargetId="figure@21.[148,1439,254,1658]" captionTargetPageId="21" captionText="Fig. 11. Liphistius tioman, details of left and right palp of a male (matured 3. I. 2002) from the type locality; left palp (A-C, E-H) and right palp (D, I). (A) Distal part of tibia and base of tarsus, retroventral view. (B) Palpal organ and tips of bristles on cumulus, retrolateral view. (C) Distal part of cymbium and palpal organ, prolateral view. (D) Palpal organ, retrolateral view. (E) Same, retroventral view. (F) Same, distal view (dorsal side at 1 o’clock). (G) Tibial apophysis, ventral view. (H) Same, retroventral view. (I) Same, retrolateral and slightly proximal view. Abbreviations: e - embolus proper; pe - para-embolic plate; rd - retrodorsal part of tibial apophysis; rv - retroventral part of tibial apophysis; vp - ventral process of contrategulum. Scale lines: 1.0 mm (A, G-H; B-E, I; F)." httpUri="https://zenodo.org/record/1021959/files/figure.png" pageId="27" pageNumber="418">Fig. 11</figureCitation>
B-C, F); scopula on tarsi IV absent.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFFFFEAB950FA2B3B66F8FD" lastPageId="29" lastPageNumber="420" pageId="27" pageNumber="418" type="description">
<paragraph id="C4261544FFFFFFEAB950FA2B38FBFE5B" blockId="27.[831,1437,1498,2077]" lastBlockId="29.[151,757,258,2000]" lastPageId="29" lastPageNumber="420" pageId="27" pageNumber="418">
<emphasis id="F6EDC956FFFFFFECB950FA2B3CA5FA12" bold="true" box="[831,1213,1498,1523]" pageId="27" pageNumber="418">
Description of male (
<typeStatus id="1B22ABE6FFFFFFECBE28FA2B3CA9FA12" box="[1095,1201,1498,1521]" pageId="27" pageNumber="418" type="holotype">holotype</typeStatus>
):
</emphasis>
<emphasis id="F6EDC956FFFFFFECBEA3FA2B3D84FA12" box="[1228,1436,1498,1523]" italics="true" pageId="27" pageNumber="418">Colour in alcohol</emphasis>
(much darker in life): Carapace mostly light brown; pars cephalica brown except for indistinct grey-brown W-shaped pattern behind eye mound and light brown medial patch between eye mound and fovea; brown area on pars cephalica connected to brown flower-shaped area around fovea; long brown patches along lateral (wide) and posterior (very narrow) margins of pars thoracica and between coxal elevations. Chelicerae with proximal portion cream-coloured, distal portion dark brown. Palps in proximal half of femur and tibia and in distal half of patella light brown, mottled with dark spots, other parts dark brown; cymbium entirely very dark reddish brown. Legs mostly dark brown except for light brown distal half of all patellae and light brown proximal half of all femora (mottled with dark spots); leg tibiae (in contrast to those of palp) to tarsi entirely dark brown. Opisthosoma mostly light brown, mottled with dark spots laterally; tergites I-II almost entirely dark brown, tergite III light brown with extensive dark lateral and median patches, following tergites with distinct dark lateral spots and increasingly indistinct median spots, last two tergites tiny and entirely dark brown.
</paragraph>
<caption id="90E645CCFFF8FFEBBAF6F8F13949F845" httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="28" pageNumber="419" targetBox="[154,1431,259,1731]" targetPageId="28">
<paragraph id="C4261544FFF8FFEBBAF6F8F13949F845" blockId="28.[153,1439,1792,1958]" pageId="28" pageNumber="419">
Fig. 15.
<taxonomicName id="03996EC7FFF8FFEBBA81F8F1398AF8F6" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[238,402,1792,1813]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="419" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
<emphasis id="F6EDC956FFF8FFEBBA81F8F1398AF8F6" box="[238,402,1792,1813]" italics="true" pageId="28" pageNumber="419">Liphistius linang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFF8FFEBBBF5F8F039F1F8F5" box="[410,489,1793,1814]" pageId="28" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
, details of left palp of two males; holotype (A-E, G-H), paratype matured 14.IX.1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I).
</paragraph>
</caption>
<paragraph id="C4261544FFF9FFEABAF8FE303A1CFDD4" blockId="29.[151,757,258,2000]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8FE303998FE3A" box="[151,384,449,473]" italics="true" pageId="29" pageNumber="420">Bristles on carapace</emphasis>
: Few short, weak bristles along all margins except posterior one, on coxal elevations, in front and behind fovea; longer, stronger bristles on, behind and in front of eye mound.
</paragraph>
<paragraph id="C4261544FFF9FFEABAF8FDB13A0AFD94" blockId="29.[151,757,258,2000]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8FDB13956FDB4" box="[151,334,576,599]" italics="true" pageId="29" pageNumber="420">Cheliceral teeth</emphasis>
: Ten small ones on promargin of cheliceral groove of each chelicera.
</paragraph>
<paragraph id="C4261544FFF9FFEABAF8FD8E3A61FCD6" blockId="29.[151,757,258,2000]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8FD8E38F5FD75" box="[151,237,639,662]" italics="true" pageId="29" pageNumber="420">Scopula</emphasis>
: Tarsi I-III with moderately dense scopula in distal two-thirds of ventral side, only distally divided by a short median stripe; tarsus IV entirely without scopula.
<emphasis id="F6EDC956FFF9FFEABAF8FD2E38C2FD15" box="[151,218,735,758]" italics="true" pageId="29" pageNumber="420">Claws</emphasis>
: Paired tarsal claws on anterior legs with 3-5 denticles, on posterior legs with 4-5 denticles; unpaired claws with one indistinct denticle or without.
</paragraph>
<paragraph id="C4261544FFF9FFEABAF8FCCF3AD1F973" blockId="29.[151,757,258,2000]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8FCCF38D1FCB6" box="[151,201,830,853]" italics="true" pageId="29" pageNumber="420">Palp</emphasis>
: Tibial apophysis situated distally, only slightly set back from anterior margin of palp (
<figureCitation id="5CA209C1FFF9FFEAB842FCAF3A8CFC96" box="[557,660,862,885]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15A</figureCitation>
), deeply divided: (a) retroventral part long and deep, pointing slightly away from axis of tibia, carrying three mediumsized, bent megaspines; (b) retrodorsal part much smaller and shorter, shaped like a low tubercle, carrying a single bent, relatively weak spine (i.e. a reduced megaspine); long bristle retroventrally below tibial apophysis weak (
<figureCitation id="5CA209C1FFF9FFEABAF0FBCC3915FBB7" box="[159,269,1085,1108]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15A</figureCitation>
, G-H). Distal margin of cymbium with indistinct lobes (
<figureCitation id="5CA209C1FFF9FFEABB37FBAD39DBFB90" box="[344,451,1116,1139]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15B</figureCitation>
). Paracymbium quite long and moderately deep, with an indistinct retrolateralproximal heel (
<figureCitation id="5CA209C1FFF9FFEABB58FB6D39A0FB56" box="[311,440,1180,1205]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15A, G</figureCitation>
); cumulus indistinct, carrying a moderately compact group of long stiff bristles (
<figureCitation id="5CA209C1FFF9FFEAB8A6FB4D38EFFB17" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15A, G</figureCitation>
). Subtegulum without apophysis. Tegulum with wide, distinctly dentate proximal edge (
<figureCitation id="5CA209C1FFF9FFEAB8EDFB0A3AF0FAF1" box="[642,744,1275,1298]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15D</figureCitation>
). Contrategulum without recognizable ventral process; prolateral surface with a few ribs, distal edge sharp in proventral part, finely dentate in prolateral part, with a deep gap in prodorsal part before reaching very narrow, slightly hook-shaped dorsal apex (
<figureCitation id="5CA209C1FFF9FFEAB875FA6B3ABCFA50" box="[538,676,1434,1459]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15B, E</figureCitation>
). Paraembolic plate large, its distal margin wide, strongly dentate and directed proventrad-distad (
<figureCitation id="5CA209C1FFF9FFEAB80CFA2B3AA0FA12" box="[611,696,1498,1521]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15</figureCitation>
B-E); embolus proper relatively narrow, inclined prolaterad (
<figureCitation id="5CA209C1FFF9FFEABAF0F9EB38F2F9D2" box="[159,234,1562,1585]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15</figureCitation>
A-D), dorsal and ventral walls of sclerotised part equally wide and lying close to each other, retrolateral wall enforced by five weak ridges (
<figureCitation id="5CA209C1FFF9FFEABB93F9A83A41F993" box="[508,601,1625,1648]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15E</figureCitation>
); membranous part of embolus proper narrow, indistinct (
<figureCitation id="5CA209C1FFF9FFEAB834F9883AA4F973" box="[603,700,1657,1680]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="29" pageNumber="420">Fig. 15B</figureCitation>
).
</paragraph>
<paragraph id="C4261544FFF9FFEABAF8F96B38CEF833" blockId="29.[151,757,258,2000]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8F96B392AF952" box="[151,306,1690,1713]" italics="true" pageId="29" pageNumber="420">Measurements</emphasis>
: Total length 14.22; carapace 5.83 long, 5.05 wide; opisthosoma 6.86 long, 4.31 wide; eye mound 0.91 long, 1.12 wide; palpal coxae 1.76 long, 1.23 wide; labium 0.49 long, 1.08 wide; sternum 2.55 long, 1.81 wide (1.03 on ventral surface); palp 9.66 long (2.55 + 1.91 + 3.53 + 1.67); leg I 17.64 long (4.80 + 2.30 + 3.82 + 4.61 + 2.11); leg II 18.67 long (4.80 + 2.30 + 4.02 + 5.20 + 2.35); leg III 20.48 long (4.95 + 2.35 + 4.26 + 6.27 + 2.65); leg IV 26.72 long (6.27 + 2.65 + 5.74 + 8.58 + 3.48).
</paragraph>
<paragraph id="C4261544FFF9FFEABAF8F8173CBAFDD9" blockId="29.[151,756,2022,2078]" lastBlockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEABAF8F8173A4EF81E" bold="true" box="[151,598,2022,2045]" pageId="29" pageNumber="420">
Description of female (
<typeStatus id="1B22ABE6FFF9FFEABB8BF8173A51F81E" box="[484,585,2022,2045]" pageId="29" pageNumber="420" type="allotype">allotype</typeStatus>
):
</emphasis>
<emphasis id="F6EDC956FFF9FFEAB81CF81738F0F7FE" italics="true" pageId="29" pageNumber="420">Colour in alcohol</emphasis>
(much darker in life): As in male, except for carapace with more extensive dark areas, all of them interconnected; W-shaped pattern behind eye mound distinct; metatarsi of legs and palps mostly dark, with a small light zone at base; tibiae with dark proximal and subdistal annulations; light brown femora with indistinct (in comparison to tibiae) dark proximal and subdistal annulations; opisthosomal tergites III-VII with more extensive dark median patches; genital area darker than surrounding parts of genital sternite, with a white, bellshaped posterior zone (
<figureCitation id="5CA209C1FFF9FFEABE58FDD23C8DFDD9" box="[1079,1173,547,570]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="29" pageNumber="420">Fig. 16F</figureCitation>
).
</paragraph>
<paragraph id="C4261544FFF9FFEAB950FDB23CDEFD99" blockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950FDB23C3CFDB8" box="[831,1060,579,603]" italics="true" pageId="29" pageNumber="420">Bristles on carapace</emphasis>
: Stiff bristles as in male, plus an additional pair just anterior to fovea.
</paragraph>
<paragraph id="C4261544FFF9FFEAB950FD723CA1FD59" blockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950FD723BF5FD79" box="[831,1005,643,666]" italics="true" pageId="29" pageNumber="420">Cheliceral teeth</emphasis>
: Nine strong teeth on promargin of left cheliceral groove, ten on right side.
</paragraph>
<paragraph id="C4261544FFF9FFEAB950FD323C3DFCB8" blockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950FD323B9AFD39" box="[831,898,707,730]" italics="true" pageId="29" pageNumber="420">Claws</emphasis>
: Palpal claw with three denticles on right side, none on left side. Paired claws with two denticles on anterior legs, 2-3 denticles on posterior legs; unpaired claws with 1-2 denticles on legs I-III, none on leg IV. All tarsi without scopula.
</paragraph>
<paragraph id="C4261544FFF9FFEAB950FC953CEAFA3D" blockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950FC953B63FC98" box="[831,891,868,891]" italics="true" pageId="29" pageNumber="420">Vulva</emphasis>
: Posterior margin of genital sterite invaginated (
<figureCitation id="5CA209C1FFF9FFEAB928FC753BABFC78" box="[839,947,900,923]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="29" pageNumber="420">Fig. 16F</figureCitation>
). Vulval plate (
<figureCitation id="5CA209C1FFF9FFEABE11FC753CCEFC78" box="[1150,1238,900,923]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="29" pageNumber="420">Fig. 16</figureCitation>
A-E, illustrations of
<typeStatus id="1B22ABE6FFF9FFEAB905FC543BCAFC5E" box="[874,978,933,957]" pageId="29" pageNumber="420" type="paratype">paratypes</typeStatus>
) with pigmentation and sclerotisation completely lost in median zone. Posterior stalk reduced to a small isolated, ventrad-bent posterior sclerite at some distance from or on posterior margin of genital sternite; posterior stalk and poreplate widely separated from each other. Genital atrium slightly sunken below level of poreplate, carrying numerous hairs on both sides of remnant of posterior stalk. Poreplate with indistinctly outlined lateral and posterior margins, these not bulging from ventral side of plate; anterior margin distinctly and widely invaginated, forming two more or less pronounced anterolateral lobes. CDO relatively small, wider than long, transversally slit-like or triangular, situated in unpigmented or weakly pigmented area of poreplate (
<figureCitation id="5CA209C1FFF9FFEAB928FA943BB4FA9F" box="[839,940,1381,1404]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="29" pageNumber="420">Fig. 16A</figureCitation>
, C-D,
<typeStatus id="1B22ABE6FFF9FFEAB999FA973C79FA9D" box="[1014,1121,1382,1406]" pageId="29" pageNumber="420" type="paratype">paratypes</typeStatus>
). Ventral receptacular cluster quite large and racemose, longitudinally divided into two lateral subclusters, its individual vesicles globular, about as long as wide (
<figureCitation id="5CA209C1FFF9FFEAB99DFA373C68FA3D" box="[1010,1136,1478,1502]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="29" pageNumber="420">Fig. 16B, E</figureCitation>
,
<typeStatus id="1B22ABE6FFF9FFEABE14FA373CFEFA3D" box="[1147,1254,1478,1502]" pageId="29" pageNumber="420" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="C4261544FFF9FFEAB950FA163B66F8FD" blockId="29.[831,1437,258,1822]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950FA163BC2FA1D" box="[831,986,1511,1534]" italics="true" pageId="29" pageNumber="420">Measurements</emphasis>
: Total length 17.25; carapace 5.93 long, 5.00 wide; opisthosoma 7.89 long, 6.13 wide; eye mound 0.82 long, 1.04 wide; palpal coxae 1.91 long, 1.37 wide; labium 0.64 long, 1.37 wide; sternum 2.65 long, 2.11 wide (1.27 on ventral surface); palp 10.00 long (3.24 + 1.86 + 2.30 + 2.60); leg I 12.40 long (3.77 + 2.16 + 2.50 + 2.50 + 1.47); leg II 12.90 long (3.82 + 2.21 + 2.55 + 2.75 + 1.57); leg III 13.73 long (3.92 + 2.25 + 2.60 + 3.24 + 1.72); leg IV 19.71 long (5.10 + 2.55 + 4.07 + 5.54 + 2.45).
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFF9FFEAB950F8C63D48F84E" pageId="29" pageNumber="420" type="discussion">
<paragraph id="C4261544FFF9FFEAB950F8C63D48F84E" blockId="29.[831,1437,1846,1966]" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950F8C63BBFF8AC" bold="true" box="[831,935,1847,1871]" pageId="29" pageNumber="420">Remark:</emphasis>
In the female
<typeStatus id="1B22ABE6FFF9FFEABE31F8C73CADF8AE" box="[1118,1205,1846,1869]" pageId="29" pageNumber="420" type="allotype">allotype</typeStatus>
the long spinnerets (anterior and posterior laterals) are bent anteriad, which is an artefact resulting from having been pressed against the wall of the vial in that position for some time.
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFF9FFE9B950F834392AF7FA" lastPageId="30" lastPageNumber="421" pageId="29" pageNumber="420" type="description">
<paragraph id="C4261544FFF9FFE9B950F834392AF7FA" blockId="29.[831,1436,1989,2078]" lastBlockId="30.[151,757,1893,2073]" lastPageId="30" lastPageNumber="421" pageId="29" pageNumber="420">
<emphasis id="F6EDC956FFF9FFEAB950F8343BAEF83F" bold="true" box="[831,950,1989,2012]" pageId="29" pageNumber="420">Variation:</emphasis>
Carapace lengths in males (n=5) 4.91- 6.15, carapace widths 4.19-5.09; in females with welldeveloped vulval plates (n=4) 5.53-5.90 and 4.29-5.03, respectively. All males lack a scopula on tarsus IV. Variation in the shape of the retrolateral tibial apophysis of the male palp is shown in
<figureCitation id="5CA209C1FFFAFFE9BBBFF8523A04F859" box="[464,540,1955,1978]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="30" pageNumber="421">Fig. 15</figureCitation>
G-I; variation in the shape of the prolateral-distal edge of the contrategulum in
<figureCitation id="5CA209C1FFFAFFE9BADBF8133918F81A" box="[180,256,2018,2041]" captionStart="Fig. 15" captionStartId="28.[153,188,1792,1813]" captionTargetBox="[154,1431,259,1731]" captionTargetId="figure@28.[148,1439,254,1752]" captionTargetPageId="28" captionText="Fig. 15. Liphistius linang sp. nov., details of left palp of two males; holotype (A-E, G-H), paratype matured 14. IX. 1999 (F, I). (A) Distal part of palp, ventral view. (B) Distal part of cymbium and palpal organ, prolateral view (arrow indicating gap in distal edge of contrategulum). (C) Palpal organ, proventral and slightly proximal view. (D) Same, retrodorsal view. (E-F) Same, distal view (dorsal side up). (G) Paracymbium and tibial apophysis, retrolateral view. (H-I) Tibial apophysis, retrolateral and slightly proximal view. Abbreviations: a - dorsal apex of contrategulum; pe - para-embolic plate. Scale lines: 1.0 mm (A, G; B-F; H-I)." httpUri="https://zenodo.org/record/1021969/files/figure.png" pageId="30" pageNumber="421">Fig. 15</figureCitation>
E-F; variation in the shape of the vulval plate in
<figureCitation id="5CA209C1FFFAFFE9BADCF7F338E6F7FA" box="[179,254,2050,2073]" captionStart="Fig. 16" captionStartId="30.[151,186,1755,1776]" captionTargetBox="[153,1440,257,1712]" captionTargetId="figure@30.[149,1458,241,1736]" captionTargetPageId="30" captionText="Fig. 16. Liphistius linang sp. nov., vulval plate of three female paratypes (A-E): specimen moulted 30. VII. 1999 (A-B), moulted 10. III. 2000 (C), exuvia, moult 14. IX. 1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F)." httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="30" pageNumber="421">Fig. 16</figureCitation>
A-E.
</paragraph>
</subSubSection>
<caption id="90E645CCFFFAFFE9BAF8F92A3A95F8C9" httpUri="https://zenodo.org/record/1021971/files/figure.png" pageId="30" pageNumber="421" targetBox="[153,1440,257,1712]" targetPageId="30">
<paragraph id="C4261544FFFAFFE9BAF8F92A3A95F8C9" blockId="30.[151,1431,1755,1834]" pageId="30" pageNumber="421">
Fig. 16.
<taxonomicName id="03996EC7FFFAFFE9BA82F92A398AF913" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[237,402,1755,1776]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="421" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
<emphasis id="F6EDC956FFFAFFE9BA82F92A398AF913" box="[237,402,1755,1776]" italics="true" pageId="30" pageNumber="421">Liphistius linang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFAFFE9BBF4F92D39F4F912" box="[411,492,1756,1777]" pageId="30" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
, vulval plate of three female paratypes (A-E): specimen moulted 30.VII.1999 (A-B), moulted 10.III.2000 (C), exuvia, moult 14.IX.1999 (D-E); undissected genital sternite of allotype (F). (A, C-D) Dorsal view. (B, E, F) Ventral view. Scale lines: 1.0 mm (A-E; F).
</paragraph>
</caption>
<subSubSection id="8C8346CFFFFAFFE8B950F89438DDFE9A" lastPageId="31" lastPageNumber="422" pageId="30" pageNumber="421" type="distribution">
<paragraph id="C4261544FFFAFFE8B950F89438DDFE9A" blockId="30.[831,1437,1893,2075]" lastBlockId="31.[151,756,258,377]" lastPageId="31" lastPageNumber="422" pageId="30" pageNumber="421">
<emphasis id="F6EDC956FFFAFFE9B950F8943BCFF89F" bold="true" box="[831,983,1893,1916]" pageId="30" pageNumber="421">Distribution:</emphasis>
Known only from the
<typeStatus id="1B22ABE6FFFAFFE9BF66F8973D20F89D" box="[1289,1336,1894,1918]" pageId="30" pageNumber="421">type</typeStatus>
locality (
<figureCitation id="5CA209C1FFFAFFE9B927F8743B8AF87E" box="[840,914,1925,1949]" captionStart="Fig. 1" captionStartId="1.[151,186,1392,1413]" captionTargetBox="[153,758,625,1349]" captionTargetId="figure@1.[149,499,895,1170]" captionTargetPageId="1" captionText="Fig. 1. Localities of Liphistius species of the malayanus - group, tioman - group, linang - group and batuenis - group in peninsular Malaysia and southern Thailand (coast of Sumatra omitted): 1 - Gunung Angsi (type locality of L. malayanus); 2 - Templer Park and Gua Anak Takun (L. malayanus, L. batuensis); 3 - Genting Highlands (L. malayanus); 4 - Fraser’s Hill (L. malayanus); 5 - Cameron Highlands (type locality of L. malayanus cameroni); 6 - Sungai Jasin in Endau Rompin National Park (type locality of L. endau); 7 - Gunung Belumut (L. endau); 8 - Gunung Muntahak (L. endau; type locality of L. gracilis sp. nov.); 9 - Sungai Rengit (type locality of L. johore); 10 - Gunung Kajang on Tioman Island (type locality of L. tioman); 11 - Gua Charas in Bukit Charas (type locality of L. panching); 12 - Nusa Camp in Taman Negara (type locality of L. negara sp. nov.); 13 - Jeram Linang Waterfall (type locality of L. linang sp. nov.); 14 - Sankalakhierie Mountains (type locality of L. indra); 15 - Batu Caves (type locality of L. batuensis); 16 - Gua Tempurung (type locality of L. tempurung); 17 - Gua Cicak (L. tempurung); 18 - Gua Keris (type locality of L. priceae sp. nov). Localities with conspecific populations are encircled. Colours distinguish species groups." httpUri="https://zenodo.org/record/1021937/files/figure.png" pageId="30" pageNumber="421">Fig. 1</figureCitation>
, locality 13) in the northeastern corner of peninsular
<collectingCountry id="BC8E55D4FFFAFFE9B9D6F8553C07F858" box="[953,1055,1956,1979]" name="Malaysia" pageId="30" pageNumber="421">Malaysia</collectingCountry>
. At the same locality, only metres away from burrows of
<taxonomicName id="03996EC7FFFAFFE9BE38F8343CD9F838" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[1111,1217,1988,2012]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="421" phylum="Arthropoda" rank="species" species="linang" status="sp. nov.">
<emphasis id="F6EDC956FFFAFFE9BE38F8343CD9F838" box="[1111,1217,1988,2012]" italics="true" pageId="30" pageNumber="421">L. linang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFAFFE9BEBFF8343D36F83F" box="[1232,1326,1989,2012]" pageId="30" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
, a single female of another, much larger, dark-coloured
<taxonomicName id="03996EC7FFFAFFE9BF5CF8123D84F819" authorityName="Schiodte" authorityYear="1849" box="[1331,1436,2019,2042]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFFAFFE9BF5CF8123D84F819" box="[1331,1436,2019,2042]" italics="true" pageId="30" pageNumber="421">Liphistius</emphasis>
</taxonomicName>
species was found. This female (mentioned under
<taxonomicName id="03996EC7FFFBFFE8BAF8FEF3391AFEFA" authorityName="Schiodte" authorityYear="1849" box="[151,258,258,281]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFFBFFE8BAF8FEF3391AFEFA" box="[151,258,258,281]" italics="true" pageId="31" pageNumber="422">Liphistius</emphasis>
</taxonomicName>
sp. at the end of the presentation of the
<taxonomicName id="03996EC7FFFBFFE8BAF8FED03913FEDB" authorityName="Abraham" authorityYear="1923" box="[151,267,289,312]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="422" phylum="Arthropoda" rank="species" species="malayanus">
<emphasis id="F6EDC956FFFBFFE8BAF8FED03913FEDB" box="[151,267,289,312]" italics="true" pageId="31" pageNumber="422">malayanus</emphasis>
</taxonomicName>
-group) appears to be more closely related to
<taxonomicName id="03996EC7FFFBFFE8BAF8FEB338EDFEBB" authorityName="Sedgwick &amp; Platnick" authorityYear="1987" box="[151,245,321,345]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="422" phylum="Arthropoda" rank="species" species="endau">
<emphasis id="F6EDC956FFFBFFE8BAF8FEB338EDFEBB" box="[151,245,321,345]" italics="true" pageId="31" pageNumber="422">L. endau</emphasis>
</taxonomicName>
than to the geographically closer
<taxonomicName id="03996EC7FFFBFFE8B80AFEB33AD6FEBA" authority="Peter J. Schwendinger, 2017" authorityName="Peter J. Schwendinger" authorityYear="2017" box="[613,718,322,345]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="422" phylum="Arthropoda" rank="species" species="negara" status="sp. nov.">
<emphasis id="F6EDC956FFFBFFE8B80AFEB33AD6FEBA" box="[613,718,322,345]" italics="true" pageId="31" pageNumber="422">L. negara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EDDE742DFFFBFFE8B8B9FEB338DDFE9A" pageId="31" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8C8346CFFFFBFFE8BAF8FE7E39D7FBDE" pageId="31" pageNumber="422" type="biology_ecology">
<paragraph id="C4261544FFFBFFE8BAF8FE7E39D3FD60" blockId="31.[151,757,399,1085]" pageId="31" pageNumber="422">
<emphasis id="F6EDC956FFFBFFE8BAF8FE7E38EFFE45" bold="true" box="[151,247,399,422]" pageId="31" pageNumber="422">Biology:</emphasis>
Spiders of this species were quite abundant in a lush rain forest along the stream above the Jeram Linang Waterfall. The burrows had a single trapdoor and mostly nine signal lines (up to
<quantity id="0361B8A1FFFBFFE8B847FE1E3A79FDE5" box="[552,609,495,519]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="422" unit="cm" value="4.0">4 cm</quantity>
long) spread over soil surface. Trapdoors of penultimate males (n=5) were 1.2-2.0 cm long and
<quantity id="0361B8A1FFFBFFE8BBA3FDDF3A5DFDA6" box="[460,581,558,582]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="31" pageNumber="422" unit="cm" value="2.05" valueMax="2.3" valueMin="1.8">1.8-2.3 cm</quantity>
wide, those of reproductive females (with egg case; n=5)
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and
<quantity id="0361B8A1FFFBFFE8BAABFD9C3922FD67" box="[196,314,621,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="31" pageNumber="422" unit="cm" value="2.05" valueMax="2.3" valueMin="1.8">1.8-2.3 cm</quantity>
, respectively.
</paragraph>
<paragraph id="C4261544FFFBFFE8BAF8FD7C39D7FBDE" blockId="31.[151,757,399,1085]" pageId="31" pageNumber="422">
Two males matured in mid-September and at the beginning of October of the year when they were collected. The three other males were raised from eggs (hatched in late May); one of them matured in November of the following year, the other two one year later in December. Less than one and a half years from hatching to maturation is exceptionally short for a
<taxonomicName id="03996EC7FFFBFFE8B825FCB83AABFC83" authorityName="Schiodte" authorityYear="1849" box="[586,691,841,864]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="422" phylum="Arthropoda" rank="genus">
<emphasis id="F6EDC956FFFBFFE8B825FCB83AABFC83" box="[586,691,841,864]" italics="true" pageId="31" pageNumber="422">Liphistius</emphasis>
</taxonomicName>
male, and this may be due to conditions in captivity in
<collectingRegion id="065DDBA6FFFBFFE8B8F3FC983AE9FC62" box="[668,753,873,897]" country="Switzerland" name="Geneve" pageId="31" pageNumber="422">Geneva</collectingRegion>
. Three females built egg cases in captivity between early February and early March, two of them were
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diameter,
<quantity id="0361B8A1FFFBFFE8BA90FC363967FC3C" box="[255,383,967,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.3" metricValueMin="1.25" pageId="31" pageNumber="422" unit="cm" value="1.275" valueMax="1.3" valueMin="1.25">1.25-1.3 cm</quantity>
high, and contained 40 and 84 light beige-yellow eggs suspended on a thin mesh of fine silk threads. The first generation females in captivity moulted in April, July and September.
</paragraph>
</subSubSection>
</treatment>
</document>
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