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<document id="E3E1302F187E4374E718AC744E43671F" ID-DOI="10.5281/zenodo.212283" ID-GBIF-Dataset="87c1b7d7-8ba9-48b9-8069-226863c4818d" ID-ISSN="1175-5326" ID-Zenodo-Dep="212283" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460524491036" checkinUser="plazi" docAuthor="Khramov, Alexander V." docDate="2012" docId="46624403FFFCC05DA79F59D0FB4CFD11" docLanguage="en" docName="zt03478p308.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Litholingia longa Khramov, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="302" masterDocId="BA5B3C7BFFF8C058A708590EFFF4FFA6" masterDocTitle="The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family" masterLastPageNumber="308" masterPageNumber="297" pageNumber="301" updateTime="1698314878310" updateUser="plazi">
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<mods:title id="B614B62CE4964BE20943FF8EFF762F78">The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family</mods:title>
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<mods:namePart id="38A48170C66915EFFEDA7C45F643339D">Khramov, Alexander V.</mods:namePart>
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<mods:date id="96FF34709B0CB26FA48ED32D5FE045DA">2012</mods:date>
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<mods:identifier id="057FC751C010D4DAA5601D5801FADD27" type="DOI">10.5281/zenodo.212283</mods:identifier>
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<treatment id="46624403FFFCC05DA79F59D0FB4CFD11" ID-DOI="http://doi.org/10.5281/zenodo.6180096" ID-GBIF-Taxon="119526082" ID-Zenodo-Dep="6180096" LSID="urn:lsid:plazi:treatment:46624403FFFCC05DA79F59D0FB4CFD11" httpUri="http://treatment.plazi.org/id/46624403FFFCC05DA79F59D0FB4CFD11" lastPageId="5" lastPageNumber="302" pageId="4" pageNumber="301">
<subSubSection id="86D1A69EFFFCC05CA79F59D0FE3BFF5E" box="[151,463,222,248]" pageId="4" pageNumber="301" type="nomenclature">
<paragraph id="CE74F515FFFCC05CA79F59D0FE3BFF5E" blockId="4.[151,463,222,248]" box="[151,463,222,248]" pageId="4" pageNumber="301">
<heading id="953C4279FFFCC05CA79F59D0FE3BFF5E" bold="true" box="[151,463,222,248]" fontSize="11" level="1" pageId="4" pageNumber="301" reason="1">
<emphasis id="FCBF2907FFFCC05CA79F59D0FE3BFF5E" bold="true" box="[151,463,222,248]" pageId="4" pageNumber="301">
<taxonomicName id="09CB8E96FFFCC05CA79F59D0FE9DFF5E" ID-CoL="6QKK5" box="[151,361,222,248]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="longa" status="sp. nov.">
<emphasis id="FCBF2907FFFCC05CA79F59D0FE9DFF5E" bold="true" box="[151,361,222,248]" italics="true" pageId="4" pageNumber="301">Litholingia longa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E78C947CFFFCC05CA67959D0FE3BFF5E" box="[369,463,222,248]" pageId="4" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86D1A69EFFFCC05CA79F582DFAC2F8C5" pageId="4" pageNumber="301" type="description">
<paragraph id="CE74F515FFFCC05CA79F582DFC29FE60" blockId="4.[151,1436,291,454]" pageId="4" pageNumber="301">
<emphasis id="FCBF2907FFFCC05CA79F582DFED5FE9A" bold="true" box="[151,289,291,316]" pageId="4" pageNumber="301">Description</emphasis>
.
<typeStatus id="11704BB7FFFCC05CA638582AFE62FE9A" box="[304,406,292,316]" pageId="4" pageNumber="301" type="holotype">
<emphasis id="FCBF2907FFFCC05CA638582AFE62FE9A" box="[304,406,292,316]" italics="true" pageId="4" pageNumber="301">Holotype</emphasis>
</typeStatus>
. Forewing (
<figureCitation id="56F0E990FFFCC05CA528582AFD92FE9A" box="[544,614,292,316]" captionStart="FIGURE 4" captionStartId="4.[151,250,1525,1548]" captionTargetBox="[183,1393,512,1524]" captionTargetId="figure@4.[151,1436,426,1566]" captionTargetPageId="4" captionText="FIGURE 4. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. (A). The photograph of the proximal part of the forewing. Holotype, PIN № 1526 / 1. (B) The drawing of the holotype, PIN № 1526 / 1. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212287/files/figure.png" pageId="4" pageNumber="301">Fig. 4</figureCitation>
). Length
<quantity id="093358F0FFFCC05CA5DA582AFCD6FE9A" box="[722,802,292,316]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="301" unit="mm" value="35.0">35 mm</quantity>
as preserved (estimated complete length
<quantity id="093358F0FFFCC05CA3FD582AFABCFE9A" box="[1269,1352,292,316]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="301" unit="mm" value="40.0">40 mm</quantity>
), width
<quantity id="093358F0FFFCC05CA79F5848FF10FEF8" box="[151,228,326,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="301" unit="mm" value="11.0">11 mm</quantity>
(estimated width
<quantity id="093358F0FFFCC05CA6A45848FDDDFEF8" box="[428,553,326,350]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="4" pageNumber="301" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
). There are two rows of cells along entire length of the wing. Longitudinal vein b located between the stem of Rs and basal part of MP and fused with Rs near to origin MA from Rs. MP forked after origin MA from Rs. Distal part of MP2 deeply dichotomously branched. CuP pectinately branched, the first of its 4 branches forked. Branches of A1 connected by crossveins and forked.
</paragraph>
<caption id="9AB4A59DFFFCC05CA79F5CFBFE5DF9EB" httpUri="https://zenodo.org/record/212287/files/figure.png" pageId="4" pageNumber="301" targetBox="[183,1393,512,1524]" targetPageId="4">
<paragraph id="CE74F515FFFCC05CA79F5CFBFE5DF9EB" blockId="4.[151,1436,1525,1613]" pageId="4" pageNumber="301">
<emphasis id="FCBF2907FFFCC05CA79F5CFBFEE6F9AD" bold="true" box="[151,274,1525,1548]" pageId="4" pageNumber="301">FIGURE 4.</emphasis>
<taxonomicName id="09CB8E96FFFCC05CA6115CF8FE3DF9AA" box="[281,457,1526,1548]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="301" phylum="Arthropoda" rank="species" species="longa" status="sp. nov.">
<emphasis id="FCBF2907FFFCC05CA6115CF8FE3DF9AA" box="[281,457,1526,1548]" italics="true" pageId="4" pageNumber="301">Litholingia longa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E78C947CFFFCC05CA6C75CF8FDD4F9AD" box="[463,544,1526,1547]" pageId="4" pageNumber="301" rank="species">sp. nov.</taxonomicNameLabel>
, Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. (A). The photograph of the proximal part of the forewing. Holotype, PIN № 1526/1. (B) The drawing of the holotype, PIN № 1526/1. Scale bar represents 5 mm.
</paragraph>
</caption>
<paragraph id="CE74F515FFFCC05CA7CF5F75FAC2F8C5" blockId="4.[151,1436,1659,2030]" pageId="4" pageNumber="301">
Specimens
<emphasis id="FCBF2907FFFCC05CA64E5F72FE87F935" box="[326,371,1660,1683]" italics="true" pageId="4" pageNumber="301">PIN</emphasis>
<emphasis id="FCBF2907FFFCC05CA6935F75FDF8F935" box="[411,524,1659,1683]" italics="true" pageId="4" pageNumber="301">2389/526,</emphasis>
<emphasis id="FCBF2907FFFCC05CA53B5F75FD50F935" box="[563,676,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/840,</emphasis>
<emphasis id="FCBF2907FFFCC05CA5C35F75FCC8F935" box="[715,828,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/842,</emphasis>
<emphasis id="FCBF2907FFFCC05CA46B5F75FC20F935" box="[867,980,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/453,</emphasis>
<emphasis id="FCBF2907FFFCC05CA4F35F75FB98F935" box="[1019,1132,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/447,</emphasis>
<emphasis id="FCBF2907FFFCC05CA39B5F75FAF0F935" box="[1171,1284,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/446,</emphasis>
<emphasis id="FCBF2907FFFCC05CA2235F75FA60F935" box="[1323,1428,1659,1683]" italics="true" pageId="4" pageNumber="301">3073/448</emphasis>
. Forewings (
<figureCitation id="56F0E990FFFCC05CA6155F90FEAFF910" box="[285,347,1694,1718]" captionStart="FIGURE 5" captionStartId="5.[151,250,1892,1915]" captionTargetBox="[185,1388,814,1891]" captionTargetId="figure@5.[151,1436,780,1917]" captionTargetPageId="5" captionText="FIGURE 5. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. The drawings of proximal parts of the forewings. A, PIN № 2389 / 526; B, PIN № 3073 / 840; C, PIN № 3073 / 842; D, PIN № 3073 / 453; E, PIN № 3073 / 447; F, PIN 3073 / 446; G, PIN 3073 / 448. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212288/files/figure.png" pageId="4" pageNumber="301">Fig.5</figureCitation>
). They slightly vary in size and belong to wings of medium length (
<quantity id="093358F0FFFCC05CA3585F90FB3AF910" box="[1104,1230,1694,1718]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="4" pageNumber="301" unit="mm" value="40.0" valueMax="45.0" valueMin="35.0">3545 mm</quantity>
). Their costal area contains two rows of cells. All specimens have the distinct oblique or longitudinal vein b between the stem of Rs and the basal part of MP. In the most of specimens vein b fused with Rs at the point of the separation MA from Rs, but also could join it distally, as in the fragment PIN № 3073/453 (
<figureCitation id="56F0E990FFFCC05CA4735E08FC49F8B8" box="[891,957,1798,1822]" captionStart="FIGURE 5" captionStartId="5.[151,250,1892,1915]" captionTargetBox="[185,1388,814,1891]" captionTargetId="figure@5.[151,1436,780,1917]" captionTargetPageId="5" captionText="FIGURE 5. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. The drawings of proximal parts of the forewings. A, PIN № 2389 / 526; B, PIN № 3073 / 840; C, PIN № 3073 / 842; D, PIN № 3073 / 453; E, PIN № 3073 / 447; F, PIN 3073 / 446; G, PIN 3073 / 448. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212288/files/figure.png" pageId="4" pageNumber="301">Fig. 5</figureCitation>
D). MP commonly forked slightly after the separation MA from Rs, but also could forked at the level of it (as in PIN 3073/448,
<figureCitation id="56F0E990FFFCC05CA37A5E26FB42F8E6" box="[1138,1206,1832,1856]" captionStart="FIGURE 5" captionStartId="5.[151,250,1892,1915]" captionTargetBox="[185,1388,814,1891]" captionTargetId="figure@5.[151,1436,780,1917]" captionTargetPageId="5" captionText="FIGURE 5. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. The drawings of proximal parts of the forewings. A, PIN № 2389 / 526; B, PIN № 3073 / 840; C, PIN № 3073 / 842; D, PIN № 3073 / 453; E, PIN № 3073 / 447; F, PIN 3073 / 446; G, PIN 3073 / 448. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212288/files/figure.png" pageId="4" pageNumber="301">Fig. 5</figureCitation>
G). The anal area is preserved in specimen PIN № 2389/526 (
<figureCitation id="56F0E990FFFCC05CA56F5E45FD5DF8C5" box="[615,681,1867,1891]" captionStart="FIGURE 5" captionStartId="5.[151,250,1892,1915]" captionTargetBox="[185,1388,814,1891]" captionTargetId="figure@5.[151,1436,780,1917]" captionTargetPageId="5" captionText="FIGURE 5. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. The drawings of proximal parts of the forewings. A, PIN № 2389 / 526; B, PIN № 3073 / 840; C, PIN № 3073 / 842; D, PIN № 3073 / 453; E, PIN № 3073 / 447; F, PIN 3073 / 446; G, PIN 3073 / 448. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212288/files/figure.png" pageId="4" pageNumber="301">Fig. 5</figureCitation>
A): A2 terminates by three short branches, A3 is a simple.
</paragraph>
</subSubSection>
<subSubSection id="86D1A69EFFFCC05CA7CF5E60FB49F848" pageId="4" pageNumber="301" type="materials_examined">
<paragraph id="CE74F515FFFCC05CA7CF5E60FAE8F80E" blockId="4.[151,1436,1659,2030]" pageId="4" pageNumber="301">
<emphasis id="FCBF2907FFFCC05CA7CF5E60FE87F820" bold="true" box="[199,371,1901,1926]" pageId="4" pageNumber="301">
<typeStatus id="11704BB7FFFCC05CA7CF5E60FEF6F820" box="[199,258,1902,1926]" pageId="4" pageNumber="301">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="11704BB7FFFCC05CA68E5E60FE1AF820" box="[390,494,1902,1926]" pageId="4" pageNumber="301" type="holotype">Holotype</typeStatus>
<collectionCode id="A8DA6DD0FFFCC05CA6F35E61FDDCF820" box="[507,552,1903,1926]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="301">PIN</collectionCode>
№ 1526/1, an incomplete forewing.
<collectingCountry id="B6DCB585FFFCC05CA4ED5E61FB9EF821" box="[997,1130,1903,1927]" name="Kyrgyzstan" pageId="4" pageNumber="301">Kyrgyzstan</collectingCountry>
: Batken Province, Batken district on the border with
<collectingCountry id="B6DCB585FFFCC05CA6C85E9EFDC6F80E" box="[448,562,1936,1960]" name="Tajikistan" pageId="4" pageNumber="301">Tajikistan</collectingCountry>
; Sai-Sagul locality, upper Lower Jurassic lower Middle Jurassic.
</paragraph>
<paragraph id="CE74F515FFFCC05CA7CF5EBCFB49F848" blockId="4.[151,1436,1659,2030]" pageId="4" pageNumber="301">
<emphasis id="FCBF2907FFFCC05CA7CF5EBCFE0DF86D" bold="true" box="[199,505,1970,1995]" pageId="4" pageNumber="301">Other material examined</emphasis>
. Specimens
<collectionCode id="A8DA6DD0FFFCC05CA5835EBAFD4CF86D" box="[651,696,1972,1995]" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="301">PIN</collectionCode>
№ 2389/526, № 3073/840, № 3073/842, № 3073/453, № 3073/ 447, № 3073/446, № 3073/448, all are fragments of the forewing proximal part. Same locality.
</paragraph>
</subSubSection>
<subSubSection id="86D1A69EFFFDC05DA7CF5996FC76FF09" box="[199,898,151,176]" pageId="5" pageNumber="302" type="etymology">
<paragraph id="CE74F515FFFDC05DA7CF5996FC76FF09" blockId="5.[151,1436,151,695]" box="[199,898,151,176]" pageId="5" pageNumber="302">
<emphasis id="FCBF2907FFFDC05DA7CF5996FEB2FF16" bold="true" box="[199,326,152,176]" pageId="5" pageNumber="302">Etymology</emphasis>
. From the Latin word
<emphasis id="FCBF2907FFFDC05DA54C5996FD4EFF16" box="[580,698,152,176]" italics="true" pageId="5" pageNumber="302">longus, - a</emphasis>
, meaning “long”.
</paragraph>
</subSubSection>
<subSubSection id="86D1A69EFFFDC05DA7CF59B4FB4CFD11" pageId="5" pageNumber="302" type="discussion">
<paragraph id="CE74F515FFFDC05DA7CF59B4FC68FE04" blockId="5.[151,1436,151,695]" pageId="5" pageNumber="302">
<emphasis id="FCBF2907FFFDC05DA7CF59B4FEBBFF75" bold="true" box="[199,335,186,211]" pageId="5" pageNumber="302">Comments.</emphasis>
The
<typeStatus id="11704BB7FFFDC05DA69859B4FE06FF74" box="[400,498,186,210]" pageId="5" pageNumber="302" type="holotype">holotype</typeStatus>
belongs to
<taxonomicName id="09CB8E96FFFDC05DA58859B5FD08FF75" box="[640,764,187,211]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="FCBF2907FFFDC05DA58859B5FD08FF75" box="[640,764,187,211]" italics="true" pageId="5" pageNumber="302">Litholingia</emphasis>
</taxonomicName>
, because A1 terminates on the posterior margin after the separation Rs1 from Rs, and CuA forked after the bifurcation of CuP. The imprint is slightly crumpled in transverse direction (R1 is not visible, the area of CuA is deformed).
<taxonomicName id="09CB8E96FFFDC05DA417580FFC88FEBE" box="[799,892,256,280]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="FCBF2907FFFDC05DA417580FFC88FEBE" box="[799,892,256,280]" italics="true" pageId="5" pageNumber="302">L. longa</emphasis>
</taxonomicName>
differs from other four species of
<taxonomicName id="09CB8E96FFFDC05DA3F1580EFA81FEBE" box="[1273,1397,256,280]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="FCBF2907FFFDC05DA3F1580EFA81FEBE" box="[1273,1397,256,280]" italics="true" pageId="5" pageNumber="302">Litholingia</emphasis>
</taxonomicName>
(
<taxonomicName id="09CB8E96FFFDC05DA28D580FFEA6FE9C" authority="Ren, 2002" authorityName="Ren" authorityYear="2002" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="rhora">
<emphasis id="FCBF2907FFFDC05DA28D580FFF22FE9D" italics="true" pageId="5" pageNumber="302">L. rhora</emphasis>
<bibRefCitation id="AA5A88E4FFFDC05DA7D5582DFEA6FE9C" author="Ren" box="[221,338,290,314]" pageId="5" pageNumber="302" refString="Ren, D. (2002) A new lacewing family (Neuroptera) from the Middle Jurassic of Inner Mongolia, China. Entomologia Sinica, 9, 53 - 67." type="journal article" year="2002">Ren, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="09CB8E96FFFDC05DA656582AFD9DFE9C" authority="Ren, 2002" authorityName="Ren" authorityYear="2002" box="[350,617,290,315]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="eumorpha">
<emphasis id="FCBF2907FFFDC05DA656582AFE19FE9D" box="[350,493,291,315]" italics="true" pageId="5" pageNumber="302">L. eumorpha</emphasis>
<bibRefCitation id="AA5A88E4FFFDC05DA6FC582DFD9DFE9C" author="Ren" box="[500,617,290,314]" pageId="5" pageNumber="302" refString="Ren, D. (2002) A new lacewing family (Neuroptera) from the Middle Jurassic of Inner Mongolia, China. Entomologia Sinica, 9, 53 - 67." type="journal article" year="2002">Ren, 2002</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="09CB8E96FFFDC05DA57C582AFC68FE9C" authority="Ren, 2002" authorityName="Ren" authorityYear="2002" box="[628,924,290,315]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="polychotoma">
<emphasis id="FCBF2907FFFDC05DA57C582AFCD7FE9D" box="[628,803,291,315]" italics="true" pageId="5" pageNumber="302">L. polychotoma</emphasis>
<bibRefCitation id="AA5A88E4FFFDC05DA421582DFC68FE9C" author="Ren" box="[809,924,290,314]" pageId="5" pageNumber="302" refString="Ren, D. (2002) A new lacewing family (Neuroptera) from the Middle Jurassic of Inner Mongolia, China. Entomologia Sinica, 9, 53 - 67." type="journal article" year="2002">Ren, 2002</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="09CB8E96FFFDC05DA4DA582AFB29FE9C" authority="Shi et al., 2011" authorityName="Shi et al." authorityYear="2011" box="[978,1245,290,315]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="ptesa">
<emphasis id="FCBF2907FFFDC05DA4DA582AFBDEFE9D" box="[978,1066,292,315]" italics="true" pageId="5" pageNumber="302">L. ptesa</emphasis>
Shi
<emphasis id="FCBF2907FFFDC05DA356582AFB7BFE9D" box="[1118,1167,291,315]" italics="true" pageId="5" pageNumber="302">et al</emphasis>
., 2011
</taxonomicName>
) by the pectinate CuP. Similar structure of CuP is observed in the forewing of
<taxonomicName id="09CB8E96FFFDC05DA45D584BFAFBFEFB" authority="Shi et al., 2011" authorityName="Shi et al." authorityYear="2011" box="[853,1295,325,349]" class="Insecta" family="Grammolingiidae" genus="Leptolingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="calonervis">
<emphasis id="FCBF2907FFFDC05DA45D584BFBA0FEFB" box="[853,1108,325,349]" italics="true" pageId="5" pageNumber="302">Leptolingia calonervis</emphasis>
Shi
<emphasis id="FCBF2907FFFDC05DA3835848FB4AFEFB" box="[1163,1214,325,349]" italics="true" pageId="5" pageNumber="302">et al</emphasis>
., 2011
</taxonomicName>
, whose CuP also has 4 branches. It corroborates the assumption of Shi
<emphasis id="FCBF2907FFFDC05DA42C5867FCA8FE26" box="[804,860,360,384]" italics="true" pageId="5" pageNumber="302">et al.</emphasis>
(2011), that CuP of
<taxonomicName id="09CB8E96FFFDC05DA3495869FAF2FED9" box="[1089,1286,359,383]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
might be not only dichotomously branched, but also slightly pectinately branched.
</paragraph>
<paragraph id="CE74F515FFFDC05DA7CF58A3FBABFDD4" blockId="5.[151,1436,151,695]" pageId="5" pageNumber="302">
The vein b, which is apparent in forewings of
<taxonomicName id="09CB8E96FFFDC05DA5DA58A0FCC4FE63" box="[722,816,429,453]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="FCBF2907FFFDC05DA5DA58A0FCC4FE63" box="[722,816,429,453]" italics="true" pageId="5" pageNumber="302">L. longa</emphasis>
</taxonomicName>
, may be interpreted as the remnant of the basal part of media, which anterior branch (MA) in the very beginning falls into Rs and immediately after that separates from it, like branches of Rs (consequently, the first vein, that separates from Rs is MA and only the second is Rs1) (
<bibRefCitation id="AA5A88E4FFFDC05DA7965B1BFEA6FD8B" author="Martynov" box="[158,338,533,557]" pageId="5" pageNumber="302" refString="Martynov, A. V. (1928) Order Neuroptera. In: Martynov, A. V., Permian fossil insects of North-East Europe. AN USSR Press, Leningrad, pp. 86 - 93." type="book chapter" year="1928">Martynov 1928</bibRefCitation>
). The remnant of the
<emphasis id="FCBF2907FFFDC05DA54C5B1BFDA7FD8B" bold="true" box="[580,595,533,557]" pageId="5" pageNumber="302">b</emphasis>
asal part of media (vein
<emphasis id="FCBF2907FFFDC05DA46D5B1BFC80FD8B" bold="true" box="[869,884,533,557]" pageId="5" pageNumber="302">b</emphasis>
) is usually visible only in hind wings (
<bibRefCitation id="AA5A88E4FFFDC05DA2245B1BFF27FDE9" author="Carpenter" pageId="5" pageNumber="302" refString="Carpenter, F. M. (1943) Osmylidae of the Florissant shales, Colorado (Insecta - Neuroptera). American Journal of Science, 241, 753 - 760." type="journal article" year="1943">Carpenter 1943</bibRefCitation>
). However, other specialists suppose, that b is only a modified crossvein (
<bibRefCitation id="AA5A88E4FFFDC05DA3175B39FB22FDE9" author="Comstock" box="[1055,1238,567,591]" pageId="5" pageNumber="302" refString="Comstock, J. H. (1918) The wings of insects. The Comstock Publishing Company, Ithaca, N. Y., 430 pp." type="book" year="1918">Comstock 1918</bibRefCitation>
,
<bibRefCitation id="AA5A88E4FFFDC05DA3EA5B39FA84FDE9" author="Tjeder" box="[1250,1392,567,591]" pageId="5" pageNumber="302" refString="Tjeder, B. (1957) Neuroptera-Planipennia. The Lacewings of Southern Africa. 1. Introduction and families Coniopterygidae, Sisyridae, and Osmylidae. In: Hanstrom B., Brinck P. &amp; Rudebec G. (Eds), South African Animal Life, 4. Swedish Natural Science Research Council, Stockholm, pp. 95 - 188." type="journal article" year="1957">Tjeder 1957</bibRefCitation>
). In any case, the presence of this vein in forewings of
<taxonomicName id="09CB8E96FFFDC05DA5C55B52FCDFFDD5" box="[717,811,603,627]" class="Insecta" family="Grammolingiidae" genus="Litholingia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="FCBF2907FFFDC05DA5C55B52FCDFFDD5" box="[717,811,603,627]" italics="true" pageId="5" pageNumber="302">L. longa</emphasis>
</taxonomicName>
is an important peculiarity.
</paragraph>
<paragraph id="CE74F515FFFDC05DA7CF5B73FB4CFD11" blockId="5.[151,1436,151,695]" pageId="5" pageNumber="302">
The comparison of seven described specimens (
<figureCitation id="56F0E990FFFDC05DA4035B73FCA0FD33" box="[779,852,637,661]" captionStart="FIGURE 5" captionStartId="5.[151,250,1892,1915]" captionTargetBox="[185,1388,814,1891]" captionTargetId="figure@5.[151,1436,780,1917]" captionTargetPageId="5" captionText="FIGURE 5. Litholingia longa sp. nov., Sai Sagul locality, Kyrgyzstan, upper Lower Jurassic lower Middle Jurassic. The drawings of proximal parts of the forewings. A, PIN № 2389 / 526; B, PIN № 3073 / 840; C, PIN № 3073 / 842; D, PIN № 3073 / 453; E, PIN № 3073 / 447; F, PIN 3073 / 446; G, PIN 3073 / 448. Scale bar represents 5 mm." httpUri="https://zenodo.org/record/212288/files/figure.png" pageId="5" pageNumber="302">Fig. 5</figureCitation>
) allows making conclusion about the individual variability of wings of
<taxonomicName id="09CB8E96FFFDC05DA69F5B91FDABFD11" box="[407,607,671,695]" class="Insecta" family="Grammolingiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="302" phylum="Arthropoda" rank="family">Grammolingiidae</taxonomicName>
, if these specimens really belong to the same species.
</paragraph>
</subSubSection>
</treatment>
</document>