treatments-xml/data/3A/6C/87/3A6C87E6FF944C7FFF0DDE0D051B04D1.xml
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<document id="3791B3288A735C2A1B6271A6843D0E62" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF944C7FFF0DDE0D051B04D1" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes dubiosa Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="41" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="41" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="BF1FB9CB2B59DC75C2FCF56A1B3D18A6">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="67FDB3FC55C5A52B4567A8C1F1275808">Irmler, Ulrich</mods:namePart>
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<mods:title id="F12CC835AE021893041B7B22DB502D53">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="1B9863E680643B5FCFF5F5BEA95649BB">
<mods:date id="14B2A0AAA403171506AAE6D6B55828FA">2021</mods:date>
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<mods:number id="776E7C9FC9A193A8DCA66348C07430D5">2021-06-30</mods:number>
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<mods:identifier id="7374024D7A2BE4A3D4B77E81A477988C" type="ISSN">0005-805X</mods:identifier>
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<treatment id="3A6C87E6FF944C7FFF0DDE0D051B04D1" ID-DOI="http://doi.org/10.5281/zenodo.5743062" ID-Zenodo-Dep="5743062" LSID="urn:lsid:plazi:treatment:3A6C87E6FF944C7FFF0DDE0D051B04D1" httpUri="http://treatment.plazi.org/id/3A6C87E6FF944C7FFF0DDE0D051B04D1" lastPageNumber="41" pageId="12" pageNumber="41">
<subSubSection id="FADF657BFF944C7FFF0DDE0D02D6079F" pageId="12" pageNumber="41" type="nomenclature">
<paragraph id="B27A36F0FF944C7FFF0DDE0D021C07BE" blockId="12.[227,752,1395,1487]" box="[227,570,1395,1422]" pageId="12" pageNumber="41">
<heading id="E932819CFF944C7FFF0DDE0D021C07BE" box="[227,570,1395,1422]" fontSize="12" level="2" pageId="12" pageNumber="41" reason="2">
<taxonomicName id="75C54D73FF944C7FFF0DDE0D01EC07BE" ID-CoL="8Y63F" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[227,458,1395,1422]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="dubiosa" status="sp. nov.">
<emphasis id="80B1EAE2FF944C7FFF0DDE0D01EC07BE" box="[227,458,1395,1422]" italics="true" pageId="12" pageNumber="41">Lithocharodes dubiosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B825799FF944C7FFE3FDE0D021C07BE" box="[465,570,1395,1422]" pageId="12" pageNumber="41" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="B27A36F0FF944C7FFF0DDEE502D6079F" blockId="12.[227,752,1395,1487]" box="[227,752,1435,1455]" pageId="12" pageNumber="41">
<uri id="C6543AF2FF944C7FFF0DDEE502D6079F" box="[227,752,1435,1455]" pageId="12" pageNumber="41">
urn:lsid:zoobank.org:act:
<uuid id="C6630C25FF944C7FFE7DDEE502D6079F" box="[403,752,1435,1455]" pageId="12" pageNumber="41">A9BFC64A-5AEA-499E-ABB9-94BE1F8F05B6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFF0DDEC501A707FF" box="[227,385,1467,1487]" pageId="12" pageNumber="41" type="description">
<paragraph id="B27A36F0FF944C7FFF0DDEC501A707FF" blockId="12.[227,752,1395,1487]" box="[227,385,1467,1487]" pageId="12" pageNumber="41">
<figureCitation id="2AFE2A75FF944C7FFF0DDEC5012907FF" box="[227,271,1467,1487]" captionStart="Fig" captionStartId="39.[151,184,1871,1893]" captionTargetBox="[170,1369,260,1835]" captionTargetId="figure-108@39.[151,1398,232,1862]" captionTargetPageId="39" captionText="Fig. 27: L.-verhaaghi-group: Lithocharodes verhaaghi (2), L. pampana (3), L. silvicola (4), L. trinitatis (5), L. dubiosa (6), L. peruana (7); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A D: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743007" httpUri="https://zenodo.org/record/5743007/files/figure.png" pageId="12" pageNumber="41">Figs 6</figureCitation>
a-d, 45 C, 49 F
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFF53DE9701C404B1" pageId="12" pageNumber="41" type="materials_examined">
<paragraph id="B27A36F0FF944C7FFF53DE9701C404B1" blockId="12.[189,795,1513,1665]" pageId="12" pageNumber="41">
<emphasis id="80B1EAE2FF944C7FFF53DE97017E0431" bold="true" box="[189,344,1513,1537]" pageId="12" pageNumber="41">Type material</emphasis>
:
<materialsCitation id="02AD3CADFF944C7FFE86DE9701F804B1" collectingDate="1989-06-26" collectionCode="KNHM" collectorName="Plot &amp; R. Leschen" country="Peru" elevation="200" latitude="-12.55" location="Amazonico" longLatPrecision="1293" longitude="-69.05" municipality="Tambopata Prov." pageId="12" pageNumber="41" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
male,
<typeStatus id="6D7E8852FF944C7FFE42DE97022B0431" box="[428,525,1513,1537]" pageId="12" pageNumber="41" type="holotype">holotype</typeStatus>
:
<collectingCountry id="CAD27660FF944C7FFDF4DE9402680431" box="[538,590,1514,1537]" name="Peru" pageId="12" pageNumber="41">Peru</collectingCountry>
,
<collectingMunicipality id="521EAC8AFF944C7FFDB2DE9703310431" box="[604,791,1513,1537]" pageId="12" pageNumber="41">Tambopata Prov.</collectingMunicipality>
,
<collectingRegion id="7001F812FF944C7FFF53DD77014E0411" box="[189,360,1545,1569]" country="Peru" name="Madre de Dios" pageId="12" pageNumber="41">Madre de Dios</collectingRegion>
Dpto.,
<locationDeviation id="A3C82FD2FF944C7FFE2ADD7703300411" box="[452,790,1545,1569]" location="Puerto Maldonado" pageId="12" pageNumber="41">
<quantity id="753D9B15FF944C7FFE2ADD7702210411" box="[452,519,1545,1569]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="41" unit="km" value="15.0">15 km</quantity>
NE Puerto Maldonado
</locationDeviation>
,
<collectingRegion id="7001F812FF944C7FFF53DD5701270471" box="[189,257,1577,1601]" country="Peru" name="Cusco" pageId="12" pageNumber="41">Cuzco</collectingRegion>
<location id="B71A602BFF944C7FFEFADD5701A90471" LSID="urn:lsid:plazi:treatment:3A6C87E6FF944C7FFF0DDE0D051B04D1:B71A602BFF944C7FFEFADD5701A90471" box="[276,399,1577,1601]" country="Peru" latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Tambopata Prov." name="Amazonico" pageId="12" pageNumber="41" stateProvince="Madre de Dios">Amazônico</location>
(
<geoCoordinate id="D7F15037FF944C7FFE44DD5701DD0471" box="[426,507,1577,1601]" degrees="12" direction="south" minutes="33" orientation="latitude" pageId="12" pageNumber="41" precision="925" value="-12.55">12°33'S</geoCoordinate>
,
<geoCoordinate id="D7F15037FF944C7FFDFCDD5702570471" box="[530,625,1577,1601]" degrees="69" direction="west" minutes="03" orientation="longitude" pageId="12" pageNumber="41" precision="925" value="-69.05">69°03'W</geoCoordinate>
),
<quantity id="753D9B15FF944C7FFD7EDD5702F40471" box="[656,722,1577,1601]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="41" unit="m" value="200.0">
<elevation id="39E8D1C3FF944C7FFD7EDD5702F40471" box="[656,722,1577,1601]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="41" unit="m" value="200.0">200 m</elevation>
</quantity>
elev.,
<collectorName id="1F305326FF944C7FFF53DD3700CE0451" box="[189,232,1609,1633]" pageId="12" pageNumber="41">Plot</collectorName>
#Z1 trail 26,
<collectingMethod id="6B844EE7FF944C7FFE61DD37024B0451" box="[399,621,1609,1633]" pageId="12" pageNumber="41">flight intercept trap</collectingMethod>
,
<date id="C67B1030FF944C7FFD93DD3702C30451" box="[637,741,1609,1633]" pageId="12" pageNumber="41" value="1989-06-26">
<collectingDate id="D63FE9D8FF944C7FFD93DD3702C30451" box="[637,741,1609,1633]" pageId="12" pageNumber="41" value="1989-06-26">26.6.1989</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF944C7FFF53DD14011504B1" box="[189,307,1641,1665]" pageId="12" pageNumber="41">R. Leschen</collectorName>
, #280 (
<collectionCode id="D4D4AE35FF944C7FFE90DD1701F004B1" box="[382,470,1641,1665]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="12" pageNumber="41">KNHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFF53DDD701A90591" pageId="12" pageNumber="41" type="diagnosis">
<paragraph id="B27A36F0FF944C7FFF53DDD701A90591" blockId="12.[189,794,1705,1953]" pageId="12" pageNumber="41">
<emphasis id="80B1EAE2FF944C7FFF53DDD7010C04F1" bold="true" box="[189,298,1705,1729]" pageId="12" pageNumber="41">Diagnosis</emphasis>
:
<taxonomicName id="75C54D73FF944C7FFEDBDDD7023A04F1" authorityName="Irmler" authorityYear="2021" box="[309,540,1705,1729]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="dubiosa">
<emphasis id="80B1EAE2FF944C7FFEDBDDD7023A04F1" box="[309,540,1705,1729]" italics="true" pageId="12" pageNumber="41">Lithocharodes dubiosa</emphasis>
</taxonomicName>
can hardly be separated from species with similar size. It can be separated from
<taxonomicName id="75C54D73FF944C7FFF53DD94011D0531" authorityName="Irmler" authorityYear="2021" box="[189,315,1770,1793]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="pampana">
<emphasis id="80B1EAE2FF944C7FFF53DD94011D0531" box="[189,315,1770,1793]" italics="true" pageId="12" pageNumber="41">L. pampana</emphasis>
</taxonomicName>
and
<taxonomicName id="75C54D73FF944C7FFE9DDD9401F90531" authorityName="Irmler" authorityYear="2021" box="[371,479,1769,1793]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="80B1EAE2FF944C7FFE9DDD9401F90531" box="[371,479,1769,1793]" italics="true" pageId="12" pageNumber="41">L. lescheni</emphasis>
</taxonomicName>
by sparser punctation of the head. The main separating character is the twisted band of the endophallus with larger teeth and lobes anteriorly and minute teeth posteriorly. The endophallus in
<taxonomicName id="75C54D73FF944C7FFD73DC34033C0551" authorityName="Irmler" authorityYear="2021" box="[669,794,1866,1889]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="pampana">
<emphasis id="80B1EAE2FF944C7FFD73DC34033C0551" box="[669,794,1866,1889]" italics="true" pageId="12" pageNumber="41">L. pampana</emphasis>
</taxonomicName>
has a spiral form and the endophallus of
<taxonomicName id="75C54D73FF944C7FFD84DC1402F005B1" authorityName="Irmler" authorityYear="2021" box="[618,726,1897,1921]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="80B1EAE2FF944C7FFD84DC1402F005B1" box="[618,726,1897,1921]" italics="true" pageId="12" pageNumber="41">L. lescheni</emphasis>
</taxonomicName>
forms a thick bulb at base.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFF53DCB703EE0451" pageId="12" pageNumber="41" type="description">
<paragraph id="B27A36F0FF944C7FFF53DCB702070A11" blockId="12.[189,794,1993,2082]" pageId="12" pageNumber="41">
<emphasis id="80B1EAE2FF944C7FFF53DCB7011905D1" bold="true" box="[189,319,1993,2017]" pageId="12" pageNumber="41">Description</emphasis>
: Length:
<quantity id="753D9B15FF944C7FFE4ADCB701DE05D1" box="[420,504,1993,2017]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="41" unit="mm" value="4.5">4.5 mm</quantity>
. Colouration: Brown; head darker brown than remaining body; legs and posterior abdominal segments light brown.
</paragraph>
<paragraph id="B27A36F0FF944C7FFCAEDB9703EE0451" blockId="12.[832,1437,233,1633]" pageId="12" pageNumber="41">
Head:
<quantity id="753D9B15FF944C7FFC62DB9703CC0331" box="[908,1002,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="12" pageNumber="41" unit="mm" value="0.79">0.79 mm</quantity>
long,
<quantity id="753D9B15FF944C7FFBD7DB9704BE0331" box="[1081,1176,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="41" unit="mm" value="0.65">0.65 mm</quantity>
wide; eyes moderately large, postocular sides twice as long as eyes; interantennal furrows convergent; reaching mid of eyes; setiferous punctation on anterior vertex denser than on posterior vertex; on average, interstices at least as wide as diameter of punctures; narrow midline impunctate; surface with extremely weak microsculpture; moderately shiny. Antennae with first antennomere half as long as length of head; second and third antennomere conical; approximately as long as apical width; subsequent antennomeres distinctly wider than long and increasing in width; forth antennomere 2.5 times as wide as long; tenth antennomere three times as long as wide; all antennomeres with long setae; fourth to eleventh antennomeres pubescent. Pronotum:
<quantity id="753D9B15FF944C7FFC57D9D7043E00F1" box="[953,1048,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="12" pageNumber="41" unit="mm" value="0.89">0.89 mm</quantity>
long,
<quantity id="753D9B15FF944C7FFBB4D9D7049E00F1" box="[1114,1208,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="12" pageNumber="41" unit="mm" value="0.58">0.58 mm</quantity>
wide; widest in anterior forth; shortly narrowed toward neck in convex curve; posteriad more strongly narrowed in middle third; nearly parallel in posterior third; posterior margin slightly convex; setiferous punctation moderately dense; on average, interstices as wide as diameter of punctures; close to posterior margin interstices slightly wider; moderately wide midline impunctate; surface with extremely weak microsculpture; moderately shiny. Elytra:
<quantity id="753D9B15FF944C7FFAEFD8D7054601F1" box="[1281,1376,937,961]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="12" pageNumber="41" unit="mm" value="0.87">0.87 mm</quantity>
long,
<quantity id="753D9B15FF944C7FFCAED8B703B801D1" box="[832,926,969,993]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="41" unit="mm" value="0.73">0.73 mm</quantity>
wide; humeral angles sub-rectangular; sides posteriad slightly divergent; posterior angles shortly rounded; posterior margin with wide triangular emargination; setiferous punctation dense; punctures larger than on head and pronotum; on average, interstices between punctures half as wide as diameter of punctures; surface with weak isodiametric microsculpture; less shiny than head and pronotum. Abdomen with setiferous punctation finer and sparser than on fore-body; surface with weak isodiametric microsculpture; posterior margin of sternite VII of male sinuate with short emargination on each side of central process; tergite VII of male straight; both sternite and tergite with longitudinal striae close to posterior margin; meso and metatibia with one subapical ctenidium. Aedeagus oval with sub-rectangular anterior part; endophallus forming one wide band with three torsions; lobes and teeth in anterior part large; tooth size decreasing posteriad; at posterior end of endophallus with minute teeth; paramere slender; curved in anterior third; at inner face with pairs of setae; setae at base longer than at apex.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF944C7FFCAEDDF7051B04D1" pageId="12" pageNumber="41" type="etymology">
<paragraph id="B27A36F0FF944C7FFCAEDDF7051B04D1" blockId="12.[832,1436,1673,1761]" pageId="12" pageNumber="41">
<emphasis id="80B1EAE2FF944C7FFCAEDDF703910491" bold="true" box="[832,951,1673,1697]" pageId="12" pageNumber="41">Etymology</emphasis>
: The species name is derived from the same Latin word (meaning doubtful) and refers to the extreme similarity with
<taxonomicName id="75C54D73FF944C7FFC0FDDB4046B04D1" authorityName="Irmler" authorityYear="2021" box="[993,1101,1737,1761]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="80B1EAE2FF944C7FFC0FDDB4046B04D1" box="[993,1101,1737,1761]" italics="true" pageId="12" pageNumber="41">L. lescheni</emphasis>
</taxonomicName>
from the same region.
</paragraph>
</subSubSection>
</treatment>
</document>