treatments-xml/data/3A/6C/87/3A6C87E6FF8B4C60FF53D86D053B0731.xml
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<document id="22BD617B64E2538CF45852CBD50C52C6" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF8B4C60FF53D86D053B0731" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes hanagarthi Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="48" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="6D96E5C3C61FC7B4B31B5C28A8C1AD55">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="1DC9069062802141375329DF68049504">Irmler, Ulrich</mods:namePart>
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<mods:typeOfResource id="8C6C3AD59538ED92AD3D3659DB4E1D22">text</mods:typeOfResource>
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<mods:title id="FF231D283111DB36507723E5E9D24861">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="2C36659484B15BD04084165527D16286">
<mods:date id="E940B55161D22CBE0EE4B6B868B0C51A">2021</mods:date>
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<mods:number id="D75288C9823D4D39598B118A54A42AAC">2021-06-30</mods:number>
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<mods:identifier id="00276916C065BF4D3E246BFB93D7D5D0" type="DOI">10.21248/contrib.entomol.71.1.029-085</mods:identifier>
<mods:identifier id="DAE359906D870A568BF907A625228CA5" type="ISSN">0005-805X</mods:identifier>
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<treatment id="3A6C87E6FF8B4C60FF53D86D053B0731" ID-DOI="http://doi.org/10.5281/zenodo.5743072" ID-Zenodo-Dep="5743072" LSID="urn:lsid:plazi:treatment:3A6C87E6FF8B4C60FF53D86D053B0731" httpUri="http://treatment.plazi.org/id/3A6C87E6FF8B4C60FF53D86D053B0731" lastPageNumber="48" pageId="19" pageNumber="48">
<subSubSection id="FADF657BFF8B4C60FF53D86D02EA017F" pageId="19" pageNumber="48" type="nomenclature">
<paragraph id="B27A36F0FF8B4C60FF53D86D0215011E" blockId="19.[189,716,787,879]" box="[189,563,787,814]" pageId="19" pageNumber="48">
<heading id="E932819CFF8B4C60FF53D86D0215011E" box="[189,563,787,814]" fontSize="12" level="2" pageId="19" pageNumber="48" reason="2">
<taxonomicName id="75C54D73FF8B4C60FF53D86D01E5011E" ID-CoL="8Y63M" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[189,451,787,814]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagarthi" status="sp. nov.">
<emphasis id="80B1EAE2FF8B4C60FF53D86D01E5011E" box="[189,451,787,814]" italics="true" pageId="19" pageNumber="48">Lithocharodes hanagarthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B825799FF8B4C60FE24D86D0215011E" box="[458,563,787,814]" pageId="19" pageNumber="48" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
<uuid id="C6630C25FF8B4C60FE9ED84502EA017F" box="[368,716,827,847]" pageId="19" pageNumber="48">CB47DB88-4D42-4FC9-AF39-A5ED1D1758A3</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8B4C60FF53D8250178015F" box="[189,350,859,879]" pageId="19" pageNumber="48" type="description">
<paragraph id="B27A36F0FF8B4C60FF53D8250178015F" blockId="19.[189,716,787,879]" box="[189,350,859,879]" pageId="19" pageNumber="48">
<figureCitation id="2AFE2A75FF8B4C60FF53D82500CF015F" box="[189,233,859,879]" captionStart="Fig" captionStartId="40.[189,222,1871,1893]" captionTargetBox="[197,1420,250,1840]" captionTargetId="figure-109@40.[189,1436,232,1848]" captionTargetPageId="40" captionText="Fig. 812: L.-hanagarthi-group and L.-sordida-group:Lithocharodes boliviana (8), L. hanagarthi (9), L. obscura (10), L. bicornis (11), L. sordida (12); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A D: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743009" httpUri="https://zenodo.org/record/5743009/files/figure.png" pageId="19" pageNumber="48">Figs 9</figureCitation>
a-d, 43 B, 46 B
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8B4C60FF79D8F700F806B1" pageId="19" pageNumber="48" type="materials_examined">
<paragraph id="B27A36F0FF8B4C60FF79D8F700F806B1" blockId="19.[151,757,905,1153]" pageId="19" pageNumber="48">
<emphasis id="80B1EAE2FF8B4C60FF79D8F701130191" bold="true" box="[151,309,905,929]" pageId="19" pageNumber="48">Type material</emphasis>
:
<materialsCitation id="02AD3CADFF8B4C60FEA6D8F701060671" collectingDate="1975-05-17" collectingDateMax="1976-05-06" collectingDateMin="1975-05-17" collectionCode="UIC" collectorName="W. Hanagarth &amp; Llullapichis &amp; Panguana Biol. Stat. &amp; W. Hana" country="Peru" latitude="-9.616667" location="Llullapichis" longLatPrecision="925" longitude="-74.094666" pageId="19" pageNumber="48" specimenCount="2" stateProvince="Huanuco" typeStatus="holotype">
male,
<typeStatus id="6D7E8852FF8B4C60FE61D8F701D60191" box="[399,496,905,929]" pageId="19" pageNumber="48" type="holotype">holotype</typeStatus>
:
<collectingCountry id="CAD27660FF8B4C60FDEED8F402120191" box="[512,564,906,929]" name="Peru" pageId="19" pageNumber="48">Peru</collectingCountry>
,
<collectingRegion id="7001F812FF8B4C60FDABD8F4028F0191" box="[581,681,906,929]" country="Peru" name="Huanuco" pageId="19" pageNumber="48">Huanuco</collectingRegion>
,
<location id="B71A602BFF8B4C60FD57D8F700C501F1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF8B4C60FF53D86D053B0731:B71A602BFF8B4C60FD57D8F700C501F1" country="Peru" latitude="-9.616667" longLatPrecision="925" longitude="-74.094666" name="Llullapichis" pageId="19" pageNumber="48" stateProvince="Huanuco">Llullapichis</location>
, Panguana Biol. Stat. (
<geoCoordinate id="D7F15037FF8B4C60FE3AD8D7026401F1" box="[468,578,937,961]" degrees="74" direction="west" minutes="5.68" orientation="longitude" pageId="19" pageNumber="48" precision="9" value="-74.094666">74°5.68'W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF8B4C60FDA0D8D702B401F1" box="[590,658,937,961]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="19" pageNumber="48" precision="925" value="-9.616667">9°37'S</geoCoordinate>
), banana plantation,
<date id="C67B1030FF8B4C60FEF6D8B701A601D1" box="[280,384,969,993]" pageId="19" pageNumber="48" value="1975-05-17">
<collectingDate id="D63FE9D8FF8B4C60FEF6D8B701A601D1" box="[280,384,969,993]" pageId="19" pageNumber="48" value="1975-05-17">17.5.1975</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF8B4C60FE2AD8B4027F01D1" box="[452,601,969,993]" pageId="19" pageNumber="48">W. Hanagarth</collectorName>
(
<collectionCode id="D4D4AE35FF8B4C60FD80D8B7028701D1" box="[622,673,969,993]" pageId="19" pageNumber="48">UIC</collectionCode>
).
<typeStatus id="6D7E8852FF8B4C60FD55D8B400F40632" pageId="19" pageNumber="48" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="CAD27660FF8B4C60FF31D89401350631" box="[223,275,1002,1025]" name="Peru" pageId="19" pageNumber="48">Peru</collectingCountry>
,
<collectingRegion id="7001F812FF8B4C60FECED89401A20631" box="[288,388,1002,1025]" country="Peru" name="Huanuco" pageId="19" pageNumber="48">Huanuco</collectingRegion>
,
<collectorName id="1F305326FF8B4C60FE7CD89702360631" box="[402,528,1001,1025]" pageId="19" pageNumber="48">Llullapichis</collectorName>
,
<collectorName id="1F305326FF8B4C60FDF3D89402D30631" box="[541,757,1001,1026]" pageId="19" pageNumber="48">Panguana Biol. Stat.</collectorName>
(
<geoCoordinate id="D7F15037FF8B4C60FF70DF77012A0611" box="[158,268,1033,1057]" degrees="74" direction="west" minutes="5.68" orientation="longitude" pageId="19" pageNumber="48" precision="9" value="-74.094666">74°5.68'W</geoCoordinate>
,
<geoCoordinate id="D7F15037FF8B4C60FEF1DF7701450611" box="[287,355,1033,1057]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="19" pageNumber="48" precision="925" value="-9.616667">9°37'S</geoCoordinate>
), floodplain forest, male,
<date id="C67B1030FF8B4C60FD7BDF7702D60611" box="[661,752,1033,1057]" pageId="19" pageNumber="48" value="1976-05-06">
<collectingDate id="D63FE9D8FF8B4C60FD7BDF7702D60611" box="[661,752,1033,1057]" pageId="19" pageNumber="48" value="1976-05-06">6.5.1976</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF8B4C60FF2ADF5401060671" box="[196,288,1066,1089]" pageId="19" pageNumber="48">W. Hana</collectorName>
</materialsCitation>
<materialsCitation id="02AD3CADFF8B4C60FEC6DF5700FD06B1" collectingDate="1976-05-09" collectingDateMax="1976-05-06" collectingDateMin="1975-05-17" collectionCode="UIC" collectorName="W. Hanagarth &amp; Llullapichis &amp; Panguana Biol. Stat. &amp; W. Hana" country="Peru" latitude="-9.616667" location="Cocha" longLatPrecision="925" longitude="-74.094666" municipality="Rio Pachitea" pageId="19" pageNumber="48" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
garth; same location and collector but
<collectingMunicipality id="521EAC8AFF8B4C60FD20DF5700D50651" pageId="19" pageNumber="48">Rio Pachitea</collectingMunicipality>
,
<location id="B71A602BFF8B4C60FF10DF3701630651" LSID="urn:lsid:plazi:treatment:3A6C87E6FF8B4C60FF53D86D053B0731:B71A602BFF8B4C60FF10DF3701630651" box="[254,325,1097,1121]" municipality="Rio Pachitea" name="Cocha" pageId="19" pageNumber="48">Cocha</location>
,
<collectingMethod id="6B844EE7FF8B4C60FEBFDF3701E70651" box="[337,449,1097,1121]" pageId="19" pageNumber="48">pitfall trap</collectingMethod>
#216, BPC, female,
<date id="C67B1030FF8B4C60FD75DF3702D20651" box="[667,756,1097,1121]" pageId="19" pageNumber="48" value="1976-05-09">
<collectingDate id="D63FE9D8FF8B4C60FD75DF3702D20651" box="[667,756,1097,1121]" pageId="19" pageNumber="48" value="1976-05-09">9.5.1976</collectingDate>
</date>
(
<collectionCode id="D4D4AE35FF8B4C60FF71DF1700F406B1" box="[159,210,1129,1153]" pageId="19" pageNumber="48">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8B4C60FF79DFD7027D07F1" pageId="19" pageNumber="48" type="diagnosis">
<paragraph id="B27A36F0FF8B4C60FF79DFD7027D07F1" blockId="19.[151,756,1193,1473]" pageId="19" pageNumber="48">
<emphasis id="80B1EAE2FF8B4C60FF79DFD7012206F1" bold="true" box="[151,260,1193,1217]" pageId="19" pageNumber="48">Diagnosis</emphasis>
:
<taxonomicName id="75C54D73FF8B4C60FEF3DFD7021506F1" authorityName="Irmler" authorityYear="2021" box="[285,563,1193,1217]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagarthi">
<emphasis id="80B1EAE2FF8B4C60FEF3DFD7021506F1" box="[285,563,1193,1217]" italics="true" pageId="19" pageNumber="48">Lithocharodes hanagarthi</emphasis>
</taxonomicName>
belongs to the group of light coloured species with sparse punctation of the head. In this respect,
<taxonomicName id="75C54D73FF8B4C60FE49DF9402130731" authorityName="Irmler" authorityYear="2021" box="[423,565,1257,1281]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagarthi">
<emphasis id="80B1EAE2FF8B4C60FE49DF9402130731" box="[423,565,1257,1281]" italics="true" pageId="19" pageNumber="48">L. hanagarthi</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="75C54D73FF8B4C60FF79DE7401030711" authorityName="SHARP" authorityYear="1876" box="[151,293,1289,1313]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="80B1EAE2FF8B4C60FF79DE7401030711" box="[151,293,1289,1313]" italics="true" pageId="19" pageNumber="48">L. fuscipennis</emphasis>
</taxonomicName>
but the eyes of
<taxonomicName id="75C54D73FF8B4C60FE35DE7402780711" box="[475,606,1289,1313]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagathi">
<emphasis id="80B1EAE2FF8B4C60FE35DE7402780711" box="[475,606,1289,1313]" italics="true" pageId="19" pageNumber="48">L. hanagathi</emphasis>
</taxonomicName>
are larger. In
<taxonomicName id="75C54D73FF8B4C60FF79DE5401030771" authorityName="Irmler" authorityYear="2021" box="[151,293,1321,1345]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagarthi">
<emphasis id="80B1EAE2FF8B4C60FF79DE5401030771" box="[151,293,1321,1345]" italics="true" pageId="19" pageNumber="48">L. hanagarthi</emphasis>
</taxonomicName>
, postocular sides are 2.5 times as long as eyes, whereas in
<taxonomicName id="75C54D73FF8B4C60FEA2DE3401FC0751" authorityName="SHARP" authorityYear="1876" box="[332,474,1353,1377]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="80B1EAE2FF8B4C60FEA2DE3401FC0751" box="[332,474,1353,1377]" italics="true" pageId="19" pageNumber="48">L. fuscipennis</emphasis>
</taxonomicName>
postocular sides are three times as long as eyes. Males of
<taxonomicName id="75C54D73FF8B4C60FE19DE1402A307B1" authorityName="Irmler" authorityYear="2021" box="[503,645,1385,1409]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hanagarthi">
<emphasis id="80B1EAE2FF8B4C60FE19DE1402A307B1" box="[503,645,1385,1409]" italics="true" pageId="19" pageNumber="48">L. hanagarthi</emphasis>
</taxonomicName>
are easily to identify by the small nearly circular aedeagus with endophallus covered only by minute teeth.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8B4C60FF79DE9704CD06B1" pageId="19" pageNumber="48" type="description">
<paragraph id="B27A36F0FF8B4C60FF79DE9700C50451" blockId="19.[151,756,1513,2081]" pageId="19" pageNumber="48">
<emphasis id="80B1EAE2FF8B4C60FF79DE97013F0431" bold="true" box="[151,281,1513,1537]" pageId="19" pageNumber="48">Description</emphasis>
: Length:
<quantity id="753D9B15FF8B4C60FE90DE9701F70431" box="[382,465,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="19" pageNumber="48" unit="mm" value="4.6">4.6 mm</quantity>
. Colouration: Light brown; head and elytra dark brown to black; small stripe at posterior margin of elytra light brown; legs slightly lighter brown.
</paragraph>
<paragraph id="B27A36F0FF8B4C60FF79DD1704CD06B1" blockId="19.[151,756,1513,2081]" lastBlockId="19.[794,1399,233,1153]" pageId="19" pageNumber="48">
Head:
<quantity id="753D9B15FF8B4C60FF0DDD17016404B1" box="[227,322,1641,1665]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="19" pageNumber="48" unit="mm" value="0.78">0.78 mm</quantity>
long,
<quantity id="753D9B15FF8B4C60FE7FDD1701C904B1" box="[401,495,1641,1665]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="19" pageNumber="48" unit="mm" value="0.62">0.62 mm</quantity>
wide; eyes moderately long; postocular space 2.5 times as long as eyes; sides slightly divergent posteriad; posterior angles widely rounded; short part of posterior margin straight; interantennal furrows deep and moderately long; reaching posterior third of eyes; setiferous punctation deep and irregularly dense; on anterior vertex denser than on posterior vertex; interstices on anterior vertex shorter than diameter of punctures; interstices on posterior vertex as wide as diameter of punctures or even wider; space between interantennal furrows impunctate; vertex with wide impunctate midline; impunctate midline widened posteriad; surface without microsculpture; shiny. Antennae with first antennomere as long as half-length of head; second and third antennomere longer than wide; conical; subsequent antennomeres wider than long and increasing in width; fourth antennomere 1.5 times as wide as long; tenth antennomere nearly 2.5 times as wide as long; antennomeres four to eleven densely pubescent. Pronotum:
<quantity id="753D9B15FF8B4C60FCB9DAF703900391" box="[855,950,393,417]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="19" pageNumber="48" unit="mm" value="0.88">0.88 mm</quantity>
long,
<quantity id="753D9B15FF8B4C60FBEFDAF704460391" box="[1025,1120,393,417]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="19" pageNumber="48" unit="mm" value="0.55">0.55 mm</quantity>
wide; widest in anterior third; strongly convergent anteriad; posteriad, slightly convergent in middle third and approximately parallel in posterior third; posterior angles sub-rectangular; posterior margin slightly curved; setiferous punctation irregular and moderately deep; adjacent to wide impunctate midline with irregular line of 12 to 13 punctures; surface without microsculpture; shiny. Elytra:
<quantity id="753D9B15FF8B4C60FAF6D917055100B1" box="[1304,1399,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="19" pageNumber="48" unit="mm" value="0.79">0.79 mm</quantity>
long,
<quantity id="753D9B15FF8B4C60FC8FD9F703990091" box="[865,959,649,673]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" pageId="19" pageNumber="48" unit="mm" value="0.66">0.66 mm</quantity>
wide; sides nearly parallel; humeral angles sub-rectangular; posterior angles nearly rectangular; posterior margin slightly triangularly emarginate to suture; setiferous punctation moderately dense and deep; interstices between punctures nearly twice as wide as diameter of punctures; surface with weak microsculpture; less shiny than head and pronotum. Abdomen with weak setiferous punctation and weak microsculpture; posterior margin of tergite VII and sternite VII of male nearly straight or slightly convex; posterior margin nearly semi-circular. Mesotibia and metatibia with 3 subapical ctenidia each. Aedeagus small; oval; at anterior orifice slightly prominent; endophallus forming transparent band with central loop; covered by minute knobs; paramere slender; in apical third, nearly rectangularly flexed; along inner margin with several short setae.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF8B4C60FCF4DFD7053B0731" pageId="19" pageNumber="48" type="etymology">
<paragraph id="B27A36F0FF8B4C60FCF4DFD7053B0731" blockId="19.[794,1398,1193,1281]" pageId="19" pageNumber="48">
<emphasis id="80B1EAE2FF8B4C60FCF4DFD703B706F1" bold="true" box="[794,913,1193,1217]" pageId="19" pageNumber="48">Etymology</emphasis>
: The name honours Werner Hanagarth, who extensively collected in the Panguana region and in other parts of
<collectingCountry id="CAD27660FF8B4C60FC9FDF9403850731" box="[881,931,1258,1281]" name="Peru" pageId="19" pageNumber="48">Peru</collectingCountry>
and unfortunately passed too early.
</paragraph>
</subSubSection>
</treatment>
</document>