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<mods:title id="B7262C1A54C99529F0D118AC2108DAF8">A Study in Scarlet: Integrative Taxonomy of the Spider Genus Loureedia (Araneae: Eresidae)</mods:title>
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<treatment id="B55C7B1AED608D7CFE10A37FD525FBCF" ID-DOI="http://doi.org/10.5281/zenodo.7630111" ID-GBIF-Taxon="209286242" ID-Zenodo-Dep="7630111" LSID="urn:lsid:plazi:treatment:B55C7B1AED608D7CFE10A37FD525FBCF" httpUri="http://treatment.plazi.org/id/B55C7B1AED608D7CFE10A37FD525FBCF" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
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<taxonomicName id="FAF5B18FED608D7AFE10A37FD2B4FE95" ID-CoL="3W8D8" ID-ENA="2663092" authority="Henriques, Miaeano and Perez-Zarcos, 2018" authorityName="Henriques, Miaeano and Perez-Zarcos" authorityYear="2018" box="[500,1197,261,285]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFE10A37FD4B0FE95" box="[500,681,261,285]" italics="true" pageId="11" pageNumber="12">Loureedia colleni</emphasis>
Henriques, Miaeano and Pérez-Zarcos, 2018
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.
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<figureCitation id="A5CED689ED608D7AFE10A35DD46BFEB6" box="[500,626,295,319]" captionStart="Figure 1" captionStartId="1.[444,514,1505,1527]" captionTargetBox="[448,1360,509,1475]" captionTargetId="figure-441@1.[448,1360,509,1480]" captionTargetPageId="1" captionText="Figure 1. Males of five species of Loureedia. (A) L. annulipes from Israel, in a defensive posture, with the bifid conductor visible (arrow) (photo: Tamás Szűuts); (B) L. annulipes from Israel (photo: Tamás Szűuts); (C) L. maroccana from Morocco (photo: János Gál); (D) L. jerbae from Djerba, Tunisia (photo: Stanislav Macík); (E) L. colleni from Spain (photo: Magali Fabregat); (F) L. phoenixi from Iran (photo: Alireza Zamani). (C) reproduced from Gál et al. [2]; (F) reproduced from Zamani and Marusik [6]." figureDoi="http://doi.org/10.5281/zenodo.7630146" httpUri="https://zenodo.org/record/7630146/files/figure.png" pageId="11" pageNumber="12">Figures 1E</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD99A35DD4B8FEB7" box="[637,673,295,319]" captionStart="Figure 2" captionStartId="2.[444,514,699,721]" captionTargetBox="[232,1356,362,664]" captionTargetId="figure-569@2.[226,1362,339,683]" captionTargetPageId="2" captionText="Figure 2. Male coloration patterns of six species of Loureedia. (A) L. colleni; (B) L. maroccana; (C) L. lucasi; (D) L. jerbae; (E) L. annulipes; (F) L. phoenixi." figureDoi="http://doi.org/10.5281/zenodo.7630150" httpUri="https://zenodo.org/record/7630150/files/figure.png" pageId="11" pageNumber="12">2A</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD48A35DD4D5FEB6" box="[684,716,295,319]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="11" pageNumber="12">4E</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD33A35DD4FDFEB7" box="[727,740,295,319]" captionStart="Figure 5" captionStartId="6.[444,514,820,842]" captionTargetBox="[128,1458,259,784]" captionTargetId="figure-183@6.[128,1458,259,784]" captionTargetPageId="6" captionText="Figure 5. Distribution records of species of Loureedia. Color codes: yellow—L. colleni, blue—L. maroccana, white—L. lucasi, green—L. jerbae, khaki—L. annulipes (SAJ264 not shown), magenta—L. phoenixi, black dot—unidentified Loureedia specimen from Jordan [7]." figureDoi="http://doi.org/10.5281/zenodo.7630156" httpUri="https://zenodo.org/record/7630156/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD16A35DD517FEB6" box="[754,782,295,319]" captionStart="Figure 6" captionStartId="7.[444,514,1953,1975]" captionTargetBox="[182,1409,261,1910]" captionTargetId="figure-88@7.[177,1409,252,1929]" captionTargetPageId="7" captionText="Figure 6. Male palps of five species of Loureedia (retrolateral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630158" httpUri="https://zenodo.org/record/7630158/files/figure.png" pageId="11" pageNumber="12">6F</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFCFDA35DD52CFEB6" box="[793,821,295,318]" captionStart="Figure 7" captionStartId="9.[444,514,2002,2024]" captionTargetBox="[160,1429,277,1966]" captionTargetId="figure-87@9.[155,1429,258,1967]" captionTargetPageId="9" captionText="Figure 7. Male palps of five species of Loureedia (ventral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630160" httpUri="https://zenodo.org/record/7630160/files/figure.png" pageId="11" pageNumber="12">7F</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFCA4A35DD545FEB6" box="[832,860,295,319]" captionStart="Figure 8" captionStartId="10.[444,514,1978,2000]" captionTargetBox="[238,1350,377,1917]" captionTargetId="figure-109@10.[238,1358,377,1936]" captionTargetPageId="10" captionText="Figure 8. Male palps of five species of Loureedia (prolateral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630162" httpUri="https://zenodo.org/record/7630162/files/figure.png" pageId="11" pageNumber="12">8F</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFC82A35DD598FEB7" box="[870,897,295,319]" captionStart="Figure 10" captionStartId="13.[444,514,843,865]" captionTargetBox="[205,1382,261,807]" captionTargetId="figure-199@13.[205,1391,251,807]" captionTargetPageId="13" captionText="Figure 10. Retreats of Loureedia colleni from Spain. (A,B) retreat layout, dorsal and lateral views (illustration: Thomas Romanoff); (C) female with the open hatch; (D) hatch being closed; (E) closed hatch. Legend (A,B): 1—main lodge; 2—external access tunnels; 3—hunting canopy and radial silk lines; 4—“retreat” hatch; 5—annex “retreat” lodge." figureDoi="http://doi.org/10.5281/zenodo.7630166" httpUri="https://zenodo.org/record/7630166/files/figure.png" pageId="11" pageNumber="12">10</figureCitation>
and
<figureCitation id="A5CED689ED608D7AFC5FA35DD5C1FEB7" box="[955,984,295,319]" captionStart="Figure 15" captionStartId="20.[444,514,1518,1540]" captionTargetBox="[148,1435,483,1482]" captionTargetId="figure-407@20.[148,1435,483,1482]" captionTargetPageId="20" captionText="Figure 15. Intraspecific variation in Loureedia colleni. (AC) male color form 1, (DF) color form 2. (A) male habitus, dorsal view; (B) bulb, retrolateral view; (C) same, prolateral view; (D) male habitus, dorsal view; (E) bulb, retrolateral view; (F) same, prolateral view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630179" httpUri="https://zenodo.org/record/7630179/files/figure.png" pageId="11" pageNumber="12">15</figureCitation>
.
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<taxonomicName id="FAF5B18FED608D7AFE10A332D267FEE8" ID-CoL="3W8D8" authority="Henriques, Miaeano and Perez-Zarcos" authorityName="Henriques, Miaeano and Perez-Zarcos" authorityYear="2018" box="[500,1150,328,352]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFE10A332D4B6FEE8" box="[500,687,328,352]" italics="true" pageId="11" pageNumber="12">Loureedia colleni</emphasis>
Henriques, Miaeano and Pérez-Zarcos
</taxonomicName>
in Henriques et al., 2018: 8,
<figureCitation id="A5CED689ED608D7AFE58A310D41BFE0A" box="[444,514,362,386]" captionStart="Figure 3" captionStartId="3.[444,514,1835,1857]" captionTargetBox="[451,1257,980,1804]" captionTargetId="figure-527@3.[451,1257,980,1804]" captionTargetPageId="3" captionText="Figure 3. SEM images of the conductor and embolus of Loureedia maroccana, male paratype. (A) oblique ventral view; (B) oblique retrolateral view; (C) apico-retrolateral view; (D) apico-prolateral view; (E) retrolateral view; (F) apico-ventral view. Abbreviations: bcm: basal conductor margin, ecm: ectal conductor margin, em: embolus, fcm: frontal conductor margin, mcm: mesal conductor margin, pa: prolateral arm of the conductor, pal: lamella of prolateral arm, ra: retrolateral arm of the conductor, ral: lamella of retrolateral arm, sc: shoulder of the conductor, st: stem of the conductor. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630152" httpUri="https://zenodo.org/record/7630152/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFDE9A310D447FE0A" box="[525,606,362,386]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStart-4="Figure 8" captionStartId-0="5.[444,514,1694,1716]" captionStartId-1="6.[444,514,820,842]" captionStartId-2="7.[444,514,1953,1975]" captionStartId-3="9.[444,514,2002,2024]" captionStartId-4="10.[444,514,1978,2000]" captionTargetBox-0="[455,1412,1046,1670]" captionTargetBox-1="[128,1458,259,784]" captionTargetBox-2="[182,1409,261,1910]" captionTargetBox-3="[160,1429,277,1966]" captionTargetBox-4="[238,1350,377,1917]" captionTargetId-0="figure-394@5.[436,1428,1034,1692]" captionTargetId-1="figure-183@6.[128,1458,259,784]" captionTargetId-2="figure-88@7.[177,1409,252,1929]" captionTargetId-3="figure-87@9.[155,1429,258,1967]" captionTargetId-4="figure-109@10.[238,1358,377,1936]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." captionText-1="Figure 5. Distribution records of species of Loureedia. Color codes: yellow—L. colleni, blue—L. maroccana, white—L. lucasi, green—L. jerbae, khaki—L. annulipes (SAJ264 not shown), magenta—L. phoenixi, black dot—unidentified Loureedia specimen from Jordan [7]." captionText-2="Figure 6. Male palps of five species of Loureedia (retrolateral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." captionText-3="Figure 7. Male palps of five species of Loureedia (ventral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." captionText-4="Figure 8. Male palps of five species of Loureedia (prolateral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7630154" figureDoi-1="http://doi.org/10.5281/zenodo.7630156" figureDoi-2="http://doi.org/10.5281/zenodo.7630158" figureDoi-3="http://doi.org/10.5281/zenodo.7630160" figureDoi-4="http://doi.org/10.5281/zenodo.7630162" httpUri-0="https://zenodo.org/record/7630154/files/figure.png" httpUri-1="https://zenodo.org/record/7630156/files/figure.png" httpUri-2="https://zenodo.org/record/7630158/files/figure.png" httpUri-3="https://zenodo.org/record/7630160/files/figure.png" httpUri-4="https://zenodo.org/record/7630162/files/figure.png" pageId="11" pageNumber="12">48ac</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD8DA310D4BAFE0A" box="[617,675,362,386]" captionStart="Figure 9" captionStartId="11.[444,514,1548,1570]" captionTargetBox="[132,1451,616,1512]" captionTargetId="figure-509@11.[132,1451,616,1518]" captionTargetPageId="11" captionText="Figure 9. Habitats of three species of Loureedia. (A,B) L. annulipes from Israel (Reut Ein-Gil); (C) L. phoenixi from Iran (photo: Alireza Zamani); (D) L. colleni from Spain (photo: Magali Fabregat)." figureDoi="http://doi.org/10.5281/zenodo.7630164" httpUri="https://zenodo.org/record/7630164/files/figure.png" pageId="11" pageNumber="12">9ad</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFD4AA310D522FE0A" box="[686,827,362,386]" captionStart="Figure 13" captionStartId="18.[444,514,1907,1929]" captionTargetBox="[248,1336,439,1866]" captionTargetId="figure-126@18.[248,1336,439,1866]" captionTargetPageId="18" captionText="Figure 13. Intraspecific variation in the male palp of Loureedia maroccana Gál et al., 2017. (AC) ventral view, (DF) prolateral view. (A,D) non-type specimen; (B,E) paratype specimen; (C,F) fresh non-type specimen. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630173" httpUri="https://zenodo.org/record/7630173/files/figure.png" pageId="11" pageNumber="12">13b and SS</figureCitation>
<figureCitation id="A5CED689ED608D7AFCD8A310D54FFE0A" box="[828,854,362,386]" captionStart="Figure 12" captionStartId="16.[444,514,954,976]" captionTargetBox="[230,1337,259,916]" captionTargetId="figure-116@16.[230,1337,259,916]" captionTargetPageId="16" captionText="Figure 12. Holotype of Loureedia maroccana Gál et al., 2017 (damaged male palp). (A) Palp, before the damage, retrolateral view; (B) palp, after the damage (ra disconnected), retrolateral view; (C) palp, after the damage (ra disconnected), ventral view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630171" httpUri="https://zenodo.org/record/7630171/files/figure.png" pageId="11" pageNumber="12">12</figureCitation>
(
<specimenCount id="2BF30185ED608D7AFC81A310D591FEF7" box="[869,904,360,385]" pageId="11" pageNumber="12">♂♀</specimenCount>
) [3] (original description).
</paragraph>
<paragraph id="3D4ACA0CED608D7AFE10A3F1D2D3FE2B" blockId="11.[444,1495,261,587]" box="[500,1226,395,419]" pageId="11" pageNumber="12">
<taxonomicName id="FAF5B18FED608D7AFE10A3F1D23DFE2B" ID-CoL="3W8D8" authority=": Zamani and Marusik 2020: 242" authorityName=": Zamani and Marusik" authorityPageNumber="242" authorityYear="2020" box="[500,1060,395,419]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFE10A3F1D4B0FE2B" box="[500,681,395,419]" italics="true" pageId="11" pageNumber="12">Loureedia colleni</emphasis>
: Zamani and Marusik 2020: 242
</taxonomicName>
,
<figureCitation id="A5CED689ED608D7AFBD4A3F1D268FE2B" box="[1072,1137,395,419]" captionStart="Figure 3" captionStartId="3.[444,514,1835,1857]" captionTargetBox="[451,1257,980,1804]" captionTargetId="figure-527@3.[451,1257,980,1804]" captionTargetPageId="3" captionText="Figure 3. SEM images of the conductor and embolus of Loureedia maroccana, male paratype. (A) oblique ventral view; (B) oblique retrolateral view; (C) apico-retrolateral view; (D) apico-prolateral view; (E) retrolateral view; (F) apico-ventral view. Abbreviations: bcm: basal conductor margin, ecm: ectal conductor margin, em: embolus, fcm: frontal conductor margin, mcm: mesal conductor margin, pa: prolateral arm of the conductor, pal: lamella of prolateral arm, ra: retrolateral arm of the conductor, ral: lamella of retrolateral arm, sc: shoulder of the conductor, st: stem of the conductor. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630152" httpUri="https://zenodo.org/record/7630152/files/figure.png" pageId="11" pageNumber="12">Fig 3i</figureCitation>
(
<specimenCount id="2BF30185ED608D7AFB64A3F6D28CFE2B" box="[1152,1173,396,419]" pageId="11" pageNumber="12"></specimenCount>
) [6].
</paragraph>
</subSubSection>
<subSubSection id="75EF9987ED608D7AFE10A3D7D5C0FDC3" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="3D4ACA0CED608D7AFE10A3D7D276FE6E" blockId="11.[444,1495,261,587]" pageId="11" pageNumber="12">
Type material.
<materialsCitation id="8D9DC051ED608D7AFD45A3D7D276FE6E" ID-GBIF-Occurrence="4027595305" collectingDate="2010-10-10" collectionCode="MNCN" collectorName="Carlos Jerez del Valle" country="Spain" elevation="820" location="Granada province" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Andalucia" typeStatus="holotype">
<typeStatus id="E24E74AEED608D7AFD45A3D7D508FE4D" box="[673,785,429,453]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
<specimenCount id="2BF30185ED608D7AFCF9A3D7D54CFE4D" box="[797,853,429,453]" pageId="11" pageNumber="12" type="male">male</specimenCount>
(
<collectionCode id="5BE452C9ED608D7AFC87A3D7D5A4FE4D" box="[867,957,429,453]" collectionName="MNCN" country="Spain" lsid="urn:lsid:biocol.org:col:35275" name="Museo Nacional de Ciencias Naturales" pageId="11" pageNumber="12" type="Museum">MNCN</collectionCode>
),
<collectingCountry id="45E28A9CED608D7AFC34A3D7D238FE4D" box="[976,1057,429,453]" name="Spain" pageId="11" pageNumber="12">SPAIN</collectingCountry>
:
<collectingRegion id="FF3104EEED608D7AFBCFA3D7D2BCFE4C" box="[1067,1189,429,453]" country="Spain" name="Andalucia" pageId="11" pageNumber="12">Andalucía</collectingRegion>
:
<location id="382A9CD7ED608D7AFB56A3D7D39AFE4D" LSID="urn:lsid:plazi:treatment:B55C7B1AED608D7CFE10A37FD525FBCF:382A9CD7ED608D7AFB56A3D7D39AFE4D" box="[1202,1411,429,453]" country="Spain" name="Granada province" pageId="11" pageNumber="12" stateProvince="Andalucia">Granada province</location>
,
<quantity id="FA0D67E9ED608D7AFA6BA3D7D3CAFE4C" box="[1423,1491,429,453]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="12" unit="m" value="820.0">
<elevation id="B6D82D3FED608D7AFA6BA3D7D3CAFE4C" box="[1423,1491,429,453]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="12" unit="m" value="820.0">820 m</elevation>
</quantity>
a.s.l.,
<date id="494BECCCED608D7AFE19A3B4D475FE6E" box="[509,620,462,486]" pageId="11" pageNumber="12" value="2010-10-10">
<collectingDate id="590F1524ED608D7AFE19A3B4D475FE6E" box="[509,620,462,486]" pageId="11" pageNumber="12" value="2010-10-10">10.X.2010</collectingDate>
</date>
(leg.
<collectorName id="9000AFDAED608D7AFD4AA3B4D5B1FE6E" box="[686,936,462,486]" pageId="11" pageNumber="12">Carlos Jerez del Valle</collectorName>
) (not examined).
</materialsCitation>
</paragraph>
<paragraph id="3D4ACA0CED608D7AFE10A38AD5C0FDC3" blockId="11.[444,1495,261,587]" pageId="11" pageNumber="12">
Other examined material.
<materialsCitation id="8D9DC051ED608D7AFCC8A38AD5C0FDC3" ID-GBIF-Occurrence="4027595303" pageId="11" pageNumber="12">
Two males and
<specimenCount id="2BF30185ED608D7AFC01A38AD27CFD80" box="[997,1125,496,520]" pageId="11" pageNumber="12" type="female">one female</specimenCount>
(
<collectionCode id="5BE452C9ED608D7AFB90A38AD2C8FD80" box="[1140,1233,496,520]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="11" pageNumber="12">HNHM</collectionCode>
9207, 9209, and 9215),
<collectingCountry id="45E28A9CED608D7AFE58A06BD417FDA1" box="[444,526,529,553]" name="Spain" pageId="11" pageNumber="12">SPAIN</collectingCountry>
:
<collectingRegion id="FF3104EEED608D7AFDFCA06BD48CFDA1" box="[536,661,529,553]" country="Spain" name="Andalucia" pageId="11" pageNumber="12">Andalucía</collectingRegion>
: Almería Province, Sierra de Gádor, Vícar, 36
<emphasis id="0F81161EED608D7AFB41A071D2A9FD96" bold="true" box="[1189,1200,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFB41A071D2A9FD96" attach="none" box="[1189,1200,523,542]" fontSize="8" pageId="11" pageNumber="12"></superScript>
</emphasis>
49
<emphasis id="0F81161EED608D7AFB28A071D2CBFD96" bold="true" box="[1228,1234,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFB28A071D2CBFD96" attach="none" box="[1228,1234,523,542]" fontSize="8" pageId="11" pageNumber="12">Į</superScript>
</emphasis>
03.0
<emphasis id="0F81161EED608D7AFAE6A071D317FD96" bold="true" box="[1282,1294,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFAE6A071D317FD96" attach="left" box="[1282,1294,523,542]" fontSize="8" pageId="11" pageNumber="12">ĮĮ</superScript>
</emphasis>
N, 2
<emphasis id="0F81161EED608D7AFAA3A071D34BFD96" bold="true" box="[1351,1362,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFAA3A071D34BFD96" attach="none" box="[1351,1362,523,542]" fontSize="8" pageId="11" pageNumber="12"></superScript>
</emphasis>
39
<emphasis id="0F81161EED608D7AFA8AA071D36DFD96" bold="true" box="[1390,1396,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFA8AA071D36DFD96" attach="none" box="[1390,1396,523,542]" fontSize="8" pageId="11" pageNumber="12">Į</superScript>
</emphasis>
14.1
<emphasis id="0F81161EED608D7AFA40A071D3A9FD96" bold="true" box="[1444,1456,523,542]" pageId="11" pageNumber="12">
<superScript id="CA806744ED608D7AFA40A071D3A9FD96" attach="left" box="[1444,1456,523,542]" fontSize="8" pageId="11" pageNumber="12">ĮĮ</superScript>
</emphasis>
W,
<quantity id="FA0D67E9ED608D7AFE58A049D418FDC2" box="[444,513,563,587]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="11" pageNumber="12" unit="m" value="820.0">820 m</quantity>
a.s.l.,
<date id="494BECCCED608D7AFDAEA049D4D1FDC3" box="[586,712,563,587]" pageId="11" pageNumber="12" value="2017-09-10">10.IX. 2017</date>
(leg. Magali Fabregat).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="698A9A84ED608D7AFE58A476D385F9CC" ID-DOI="http://doi.org/10.5281/zenodo.7630164" ID-Zenodo-Dep="7630164" httpUri="https://zenodo.org/record/7630164/files/figure.png" pageId="11" pageNumber="12" startId="11.[444,514,1548,1570]" targetBox="[132,1451,616,1512]" targetPageId="11">
<paragraph id="3D4ACA0CED608D7AFE58A476D385F9CC" blockId="11.[444,1495,1548,1604]" pageId="11" pageNumber="12">
<emphasis id="0F81161EED608D7AFE58A476D402F9A9" bold="true" box="[444,539,1548,1570]" pageId="11" pageNumber="12">Figure 9.</emphasis>
Habitats of three species of
<taxonomicName id="FAF5B18FED608D7AFCA9A476D5B4F9A9" box="[845,941,1548,1569]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="0F81161EED608D7AFCA9A476D5B4F9A9" box="[845,941,1548,1569]" italics="true" pageId="11" pageNumber="12">Loureedia</emphasis>
</taxonomicName>
. (
<emphasis id="0F81161EED608D7AFC20A476D5CEF9A9" bold="true" box="[964,983,1548,1569]" pageId="11" pageNumber="12">A</emphasis>
,
<emphasis id="0F81161EED608D7AFC38A476D5F5F9A9" bold="true" box="[988,1004,1548,1569]" pageId="11" pageNumber="12">B</emphasis>
)
<taxonomicName id="FAF5B18FED608D7AFC1FA477D26CF9A9" box="[1019,1141,1548,1570]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="0F81161EED608D7AFC1FA477D26CF9A9" box="[1019,1141,1548,1570]" italics="true" pageId="11" pageNumber="12">L. annulipes</emphasis>
</taxonomicName>
from Israel (Reut Ein-Gil); (
<emphasis id="0F81161EED608D7AFA47A476D3ADF9AA" bold="true" box="[1443,1460,1548,1570]" pageId="11" pageNumber="12">C</emphasis>
)
<taxonomicName id="FAF5B18FED608D7AFA27A477D417F9CB" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="phoenixi">
<emphasis id="0F81161EED608D7AFA27A477D417F9CB" italics="true" pageId="11" pageNumber="12">L. phoenixi</emphasis>
</taxonomicName>
from Iran (photo: Alireza Zamani); (
<emphasis id="0F81161EED608D7AFC77A454D5BEF9CB" bold="true" box="[915,935,1582,1603]" pageId="11" pageNumber="12">D</emphasis>
)
<taxonomicName id="FAF5B18FED608D7AFC51A455D214F9CB" authorityName=": Zamani and Marusik" authorityYear="2020" box="[949,1037,1582,1604]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFC51A455D214F9CB" box="[949,1037,1582,1604]" italics="true" pageId="11" pageNumber="12">L. colleni</emphasis>
</taxonomicName>
from Spain (photo: Magali Fabregat).
</paragraph>
</caption>
<subSubSection id="75EF9987ED608D7AFE10A412D32EF8E3" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="3D4ACA0CED608D7AFE10A412D32EF8E3" blockId="11.[443,1497,1640,2066]" pageId="11" pageNumber="12">
Diagnosis. This species differs from all of its congeners by the black-and-white coloration pattern of the male (
<figureCitation id="A5CED689ED608D7AFD1BA4F0D563F929" box="[767,890,1674,1698]" captionStart="Figure 1" captionStartId="1.[444,514,1505,1527]" captionTargetBox="[448,1360,509,1475]" captionTargetId="figure-441@1.[448,1360,509,1480]" captionTargetPageId="1" captionText="Figure 1. Males of five species of Loureedia. (A) L. annulipes from Israel, in a defensive posture, with the bifid conductor visible (arrow) (photo: Tamás Szűuts); (B) L. annulipes from Israel (photo: Tamás Szűuts); (C) L. maroccana from Morocco (photo: János Gál); (D) L. jerbae from Djerba, Tunisia (photo: Stanislav Macík); (E) L. colleni from Spain (photo: Magali Fabregat); (F) L. phoenixi from Iran (photo: Alireza Zamani). (C) reproduced from Gál et al. [2]; (F) reproduced from Zamani and Marusik [6]." figureDoi="http://doi.org/10.5281/zenodo.7630146" httpUri="https://zenodo.org/record/7630146/files/figure.png" pageId="11" pageNumber="12">Figures 1E</figureCitation>
and
<figureCitation id="A5CED689ED608D7AFC57A4F0D5CFF92A" box="[947,982,1674,1698]" captionStart="Figure 2" captionStartId="2.[444,514,699,721]" captionTargetBox="[232,1356,362,664]" captionTargetId="figure-569@2.[226,1362,339,683]" captionTargetPageId="2" captionText="Figure 2. Male coloration patterns of six species of Loureedia. (A) L. colleni; (B) L. maroccana; (C) L. lucasi; (D) L. jerbae; (E) L. annulipes; (F) L. phoenixi." figureDoi="http://doi.org/10.5281/zenodo.7630150" httpUri="https://zenodo.org/record/7630150/files/figure.png" pageId="11" pageNumber="12">2A</figureCitation>
) vs. having yellowish to scarlet red abdominal patterns (see
<figureCitation id="A5CED689ED608D7AFD9BA4D1D504F94B" box="[639,797,1707,1731]" captionStart="Figure 1" captionStartId="1.[444,514,1505,1527]" captionTargetBox="[448,1360,509,1475]" captionTargetId="figure-441@1.[448,1360,509,1480]" captionTargetPageId="1" captionText="Figure 1. Males of five species of Loureedia. (A) L. annulipes from Israel, in a defensive posture, with the bifid conductor visible (arrow) (photo: Tamás Szűuts); (B) L. annulipes from Israel (photo: Tamás Szűuts); (C) L. maroccana from Morocco (photo: János Gál); (D) L. jerbae from Djerba, Tunisia (photo: Stanislav Macík); (E) L. colleni from Spain (photo: Magali Fabregat); (F) L. phoenixi from Iran (photo: Alireza Zamani). (C) reproduced from Gál et al. [2]; (F) reproduced from Zamani and Marusik [6]." figureDoi="http://doi.org/10.5281/zenodo.7630146" httpUri="https://zenodo.org/record/7630146/files/figure.png" pageId="11" pageNumber="12">Figures 1AD</figureCitation>
and
<figureCitation id="A5CED689ED608D7AFCB2A4D1D58AF94B" box="[854,915,1707,1731]" captionStart="Figure 2" captionStartId="2.[444,514,699,721]" captionTargetBox="[232,1356,362,664]" captionTargetId="figure-569@2.[226,1362,339,683]" captionTargetPageId="2" captionText="Figure 2. Male coloration patterns of six species of Loureedia. (A) L. colleni; (B) L. maroccana; (C) L. lucasi; (D) L. jerbae; (E) L. annulipes; (F) L. phoenixi." figureDoi="http://doi.org/10.5281/zenodo.7630150" httpUri="https://zenodo.org/record/7630150/files/figure.png" pageId="11" pageNumber="12">2BF</figureCitation>
). The male palp of
<taxonomicName id="FAF5B18FED608D7AFB89A4D6D2D4F94B" authorityName=": Zamani and Marusik" authorityYear="2020" box="[1133,1229,1707,1731]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFB89A4D6D2D4F94B" box="[1133,1229,1707,1731]" italics="true" pageId="11" pageNumber="12">L. colleni</emphasis>
</taxonomicName>
(
<figureCitation id="A5CED689ED608D7AFB38A4D1D34CF94B" box="[1244,1365,1707,1731]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="11" pageNumber="12">Figures 4F</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFABBA4D1D362F94B" box="[1375,1403,1707,1731]" captionStart="Figure 6" captionStartId="7.[444,514,1953,1975]" captionTargetBox="[182,1409,261,1910]" captionTargetId="figure-88@7.[177,1409,252,1929]" captionTargetPageId="7" captionText="Figure 6. Male palps of five species of Loureedia (retrolateral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630158" httpUri="https://zenodo.org/record/7630158/files/figure.png" pageId="11" pageNumber="12">6F</figureCitation>
,
<figureCitation id="A5CED689ED608D7AFA61A4D6D3B9F94B" box="[1413,1440,1708,1731]" captionStart="Figure 7" captionStartId="9.[444,514,2002,2024]" captionTargetBox="[160,1429,277,1966]" captionTargetId="figure-87@9.[155,1429,258,1967]" captionTargetPageId="9" captionText="Figure 7. Male palps of five species of Loureedia (ventral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630160" httpUri="https://zenodo.org/record/7630160/files/figure.png" pageId="11" pageNumber="12">7F</figureCitation>
and
<figureCitation id="A5CED689ED608D7AFE58A4B7D7C0F96D" box="[444,473,1741,1765]" captionStart="Figure 8" captionStartId="10.[444,514,1978,2000]" captionTargetBox="[238,1350,377,1917]" captionTargetId="figure-109@10.[238,1358,377,1936]" captionTargetPageId="10" captionText="Figure 8. Male palps of five species of Loureedia (prolateral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630162" httpUri="https://zenodo.org/record/7630162/files/figure.png" pageId="11" pageNumber="12">8F</figureCitation>
) is most similar to that of
<taxonomicName id="FAF5B18FED608D7AFCE2A4B7D590F96C" box="[774,905,1740,1764]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="0F81161EED608D7AFCE2A4B7D590F96C" box="[774,905,1740,1764]" italics="true" pageId="11" pageNumber="12">L. annulipes</emphasis>
</taxonomicName>
(
<figureCitation id="A5CED689ED608D7AFC73A4B6D212F96D" box="[919,1035,1740,1765]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="11" pageNumber="12">Figure 4C</figureCitation>
), as the prolateral arm of the conductor is much shorter than the retrolateral arm (
<figureCitation id="A5CED689ED608D7AFC45A494D20BF88E" box="[929,1042,1774,1798]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="11" pageNumber="12">Figure 4F</figureCitation>
), which bears a gradual curvature (7F). The male of
<taxonomicName id="FAF5B18FED608D7AFDA8A56AD4B7F8AF" authorityName=": Zamani and Marusik" authorityYear="2020" box="[588,686,1807,1831]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED608D7AFDA8A56AD4B7F8AF" box="[588,686,1807,1831]" italics="true" pageId="11" pageNumber="12">L. colleni</emphasis>
</taxonomicName>
can be diagnosed by the narrower stem of the conductor (
<figureCitation id="A5CED689ED608D7AFAB3A575D3D3F8A0" box="[1367,1482,1807,1832]" captionStart="Figure 6" captionStartId="7.[444,514,1953,1975]" captionTargetBox="[182,1409,261,1910]" captionTargetId="figure-88@7.[177,1409,252,1929]" captionTargetPageId="7" captionText="Figure 6. Male palps of five species of Loureedia (retrolateral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630158" httpUri="https://zenodo.org/record/7630158/files/figure.png" pageId="11" pageNumber="12">Figure 6F</figureCitation>
), with an almost straight mesal margin (
<figureCitation id="A5CED689ED608D7AFC76A54BD210F8C1" box="[914,1033,1841,1865]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="11" pageNumber="12">Figure 4F</figureCitation>
). The female can be recognized by an epigynal fovea that is longer than it is wide (see Henriques et al. [3]:
<figureCitation id="A5CED689ED608D7AFB05A528D333F8E3" box="[1249,1322,1874,1899]" captionStart="Figure 8" captionStartId="10.[444,514,1978,2000]" captionTargetBox="[238,1350,377,1917]" captionTargetId="figure-109@10.[238,1358,377,1936]" captionTargetPageId="10" captionText="Figure 8. Male palps of five species of Loureedia (prolateral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630162" httpUri="https://zenodo.org/record/7630162/files/figure.png" pageId="11" pageNumber="12">Fig 8a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="75EF9987ED608D7CFE10A50ED28DFB8C" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="description">
<paragraph id="3D4ACA0CED608D7DFE10A50ED469FE4D" blockId="11.[443,1497,1640,2066]" lastBlockId="12.[442,1497,261,1558]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Description. Male. Habitus as in
<figureCitation id="A5CED689ED608D7AFC68A50ED215F803" box="[908,1036,1908,1932]" captionStart="Figure 1" captionStartId="1.[444,514,1505,1527]" captionTargetBox="[448,1360,509,1475]" captionTargetId="figure-441@1.[448,1360,509,1480]" captionTargetPageId="1" captionText="Figure 1. Males of five species of Loureedia. (A) L. annulipes from Israel, in a defensive posture, with the bifid conductor visible (arrow) (photo: Tamás Szűuts); (B) L. annulipes from Israel (photo: Tamás Szűuts); (C) L. maroccana from Morocco (photo: János Gál); (D) L. jerbae from Djerba, Tunisia (photo: Stanislav Macík); (E) L. colleni from Spain (photo: Magali Fabregat); (F) L. phoenixi from Iran (photo: Alireza Zamani). (C) reproduced from Gál et al. [2]; (F) reproduced from Zamani and Marusik [6]." figureDoi="http://doi.org/10.5281/zenodo.7630146" httpUri="https://zenodo.org/record/7630146/files/figure.png" pageId="11" pageNumber="12">Figures 1E</figureCitation>
and
<figureCitation id="A5CED689ED608D7AFBAEA50ED276F804" box="[1098,1135,1908,1932]" captionStart="Figure 2" captionStartId="2.[444,514,699,721]" captionTargetBox="[232,1356,362,664]" captionTargetId="figure-569@2.[226,1362,339,683]" captionTargetPageId="2" captionText="Figure 2. Male coloration patterns of six species of Loureedia. (A) L. colleni; (B) L. maroccana; (C) L. lucasi; (D) L. jerbae; (E) L. annulipes; (F) L. phoenixi." figureDoi="http://doi.org/10.5281/zenodo.7630150" httpUri="https://zenodo.org/record/7630150/files/figure.png" pageId="11" pageNumber="12">2A</figureCitation>
. Total length: 6.43. Carapace: 3.35 long and 2.84 wide. Abdomen: 3.19 long and 2.55 wide. Eye sizes and inter-eye distances: AME 0.14, PME 0.16, ALE 0.04, PLE 0.08, AMEAME 0.30, and ALEAME 0.76. The carapace, sternum, labium, chelicerae, and maxillae black. Carapace mostly covered with long black setae and scattered shorter white setae. White setae localized densely on the pars thoracica and form a triangle on the pars cephalica (
<figureCitation id="A5CED689ED678D7DFB42A37FD306FE95" box="[1190,1311,261,285]" captionStart="Figure 1" captionStartId="1.[444,514,1505,1527]" captionTargetBox="[448,1360,509,1475]" captionTargetId="figure-441@1.[448,1360,509,1480]" captionTargetPageId="1" captionText="Figure 1. Males of five species of Loureedia. (A) L. annulipes from Israel, in a defensive posture, with the bifid conductor visible (arrow) (photo: Tamás Szűuts); (B) L. annulipes from Israel (photo: Tamás Szűuts); (C) L. maroccana from Morocco (photo: János Gál); (D) L. jerbae from Djerba, Tunisia (photo: Stanislav Macík); (E) L. colleni from Spain (photo: Magali Fabregat); (F) L. phoenixi from Iran (photo: Alireza Zamani). (C) reproduced from Gál et al. [2]; (F) reproduced from Zamani and Marusik [6]." figureDoi="http://doi.org/10.5281/zenodo.7630146" httpUri="https://zenodo.org/record/7630146/files/figure.png" pageId="12" pageNumber="13">Figure 1E</figureCitation>
). Legs covered with thick white hairs. Abdomen velvet black with a longitudinal median white foliate pattern bearing a distinct mediolateral lobe; the most anterior part of the folium merging and forming a distinct white spot. Measurements of legs: I: 7.15 (2.07, 1.16, 1.45, 1.43, 1.01); II: 6.33 (1.94, 1.19, 1.18, 1.22, 0.78); III: 5.44 (1.85, 0.80, 1.08, 1.01, 0.68); IV: 7.29 (2.32, 1.28, 1.56, 1.39, 0.71).
</paragraph>
<paragraph id="3D4ACA0CED678D7DFE10A3B4D5CFFDC3" blockId="12.[442,1497,261,1558]" pageId="12" pageNumber="13">
Palp as in
<figureCitation id="A5CED689ED678D7DFD96A3B4D4F6FE6E" box="[626,751,462,486]" captionStart="Figure 4" captionStartId="5.[444,514,1694,1716]" captionTargetBox="[455,1412,1046,1670]" captionTargetId="figure-394@5.[436,1428,1034,1692]" captionTargetPageId="5" captionText="Figure 4. Conductors of six species of Loureedia (ventral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. lucasi; (E) L. maroccana; (F) L. colleni. A, D, and F are based on Zamani and Marusik [6]. Line drawings by Mahla Pourcheraghi." figureDoi="http://doi.org/10.5281/zenodo.7630154" httpUri="https://zenodo.org/record/7630154/files/figure.png" pageId="12" pageNumber="13">Figures 4F</figureCitation>
,
<figureCitation id="A5CED689ED678D7DFD1FA3B4D501FE6E" box="[763,792,462,486]" captionStart="Figure 6" captionStartId="7.[444,514,1953,1975]" captionTargetBox="[182,1409,261,1910]" captionTargetId="figure-88@7.[177,1409,252,1929]" captionTargetPageId="7" captionText="Figure 6. Male palps of five species of Loureedia (retrolateral view). (A) L. phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630158" httpUri="https://zenodo.org/record/7630158/files/figure.png" pageId="12" pageNumber="13">6F</figureCitation>
,
<figureCitation id="A5CED689ED678D7DFCC0A3B5D559FE6E" box="[804,832,463,486]" captionStart="Figure 7" captionStartId="9.[444,514,2002,2024]" captionTargetBox="[160,1429,277,1966]" captionTargetId="figure-87@9.[155,1429,258,1967]" captionTargetPageId="9" captionText="Figure 7. Male palps of five species of Loureedia (ventral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630160" httpUri="https://zenodo.org/record/7630160/files/figure.png" pageId="12" pageNumber="13">7F</figureCitation>
and
<specimenCount id="2BF30185ED678D7DFC99A3B4D586FE6E" box="[893,927,462,486]" pageId="12" pageNumber="13" type="generic">
<figureCitation id="A5CED689ED678D7DFC99A3B4D582FE6E" box="[893,923,462,486]" captionStart="Figure 8" captionStartId="10.[444,514,1978,2000]" captionTargetBox="[238,1350,377,1917]" captionTargetId="figure-109@10.[238,1358,377,1936]" captionTargetPageId="10" captionText="Figure 8. Male palps of five species of Loureedia (prolateral view). (A) Loureedia phoenixi; (B) L. jerbae; (C) L. annulipes; (D) L. maroccana, holotype (after conductor was broken); (E) L. maroccana, holotype (conductor still intact); (F) L. colleni. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630162" httpUri="https://zenodo.org/record/7630162/files/figure.png" pageId="12" pageNumber="13">8F</figureCitation>
.
</specimenCount>
The stem of the conductor ca. 1.5 times longer than wide. The mesal margin of the conductor almost straight, and the ectal margin with an apical invagination. The retrolateral arm of the conductor ca. 2.5 times longer than the prolateral arm, and both arms with blunt tips.
</paragraph>
<paragraph id="3D4ACA0CED678D7DFE10A02ED575FDE4" blockId="12.[442,1497,261,1558]" box="[500,876,596,620]" pageId="12" pageNumber="13">Female. See Henriques et al. [3].</paragraph>
<paragraph id="3D4ACA0CED678D7DFE10A00CD2D1FCBD" blockId="12.[442,1497,261,1558]" pageId="12" pageNumber="13">
Variation. A wide array of abdominal pattern variations has already been illustrated [3]. The highest amount of variation occurs in the white foliate pattern, which may either be solid or form two large separate patches. Here, we examined two distinct color pattern forms (
<figureCitation id="A5CED689ED678D7DFDF4A0A0D4B7FD7A" box="[528,686,730,754]" captionStart="Figure 15" captionStartId="20.[444,514,1518,1540]" captionTargetBox="[148,1435,483,1482]" captionTargetId="figure-407@20.[148,1435,483,1482]" captionTargetPageId="20" captionText="Figure 15. Intraspecific variation in Loureedia colleni. (AC) male color form 1, (DF) color form 2. (A) male habitus, dorsal view; (B) bulb, retrolateral view; (C) same, prolateral view; (D) male habitus, dorsal view; (E) bulb, retrolateral view; (F) same, prolateral view. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7630179" httpUri="https://zenodo.org/record/7630179/files/figure.png" pageId="12" pageNumber="13">Figure 15A,B</figureCitation>
). Minor variations also occur on the male palp; these are considered intraspecific variations, as the COI sequences of the
<specimenCount id="2BF30185ED678D7DFBF4A086D29FFC9B" box="[1040,1158,763,787]" pageId="12" pageNumber="13" type="male">two males</specimenCount>
were identical, whereas they were slightly different (99.965% similarity) from that of the female.
</paragraph>
<paragraph id="3D4ACA0CED678D7DFE10A144D308FC75" blockId="12.[442,1497,261,1558]" pageId="12" pageNumber="13">
Natural history. The species habitat preference has already been described [3]. The examined specimens were collected on a hillside with south and south-east exposure in a semi-arid open area (
<figureCitation id="A5CED689ED678D7DFD35A1FBD552FC11" box="[721,843,897,921]" captionStart="Figure 9" captionStartId="11.[444,514,1548,1570]" captionTargetBox="[132,1451,616,1512]" captionTargetId="figure-509@11.[132,1451,616,1518]" captionTargetPageId="11" captionText="Figure 9. Habitats of three species of Loureedia. (A,B) L. annulipes from Israel (Reut Ein-Gil); (C) L. phoenixi from Iran (photo: Alireza Zamani); (D) L. colleni from Spain (photo: Magali Fabregat)." figureDoi="http://doi.org/10.5281/zenodo.7630164" httpUri="https://zenodo.org/record/7630164/files/figure.png" pageId="12" pageNumber="13">Figure 9D</figureCitation>
) with short, sparse vegetation. The vegetation in this area mainly consists of degraded bushes, tufts of thyme, thorny broom,
<taxonomicName id="FAF5B18FED678D7DFB1CA1DED3C9FC33" box="[1272,1488,931,955]" class="Magnoliopsida" family="Asteraceae" genus="Launaea" kingdom="Plantae" order="Asterales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="arborescens">
<emphasis id="0F81161EED678D7DFB1CA1DED3C9FC33" box="[1272,1488,931,955]" italics="true" pageId="12" pageNumber="13">Launaea arborescens</emphasis>
</taxonomicName>
, and
<taxonomicName id="FAF5B18FED678D7DFE14A1BFD4EFFC54" box="[496,758,964,988]" class="Magnoliopsida" family="Fabaceae" genus="Ononis" kingdom="Plantae" order="Fabales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="subSpecies" species="natrix" subSpecies="hispanica">
<emphasis id="0F81161EED678D7DFE14A1BFD4EFFC54" box="[496,758,964,988]" italics="true" pageId="12" pageNumber="13">Ononis natrix hispanica</emphasis>
</taxonomicName>
. The soil is mainly puddingstone, made up of the Alpujarride complex and Baetic/Penibaetic cordillera, covered with small flat stones.
</paragraph>
<paragraph id="3D4ACA0CED678D7DFE10A67DD564F99E" blockId="12.[442,1497,261,1558]" pageId="12" pageNumber="13">
Several webs have been observed in multiple similar biotopes in
<collectingRegion id="FF3104EEED678D7DFAEBA67DD397FB97" box="[1295,1422,1031,1055]" country="Spain" name="Andalucia" pageId="12" pageNumber="13">Andalusia</collectingRegion>
. This singular web pattern turned out to be common, and adult and juvenile specimens both constructed it (including those who were kept alive in captivity). The very discreet webs are located under small stones on the ground. This structure provides the spider with protection against the elements: mainly intense heat but also wind and rare precipitation. Hunting canopies are very short and simple compared to those woven by species of
<taxonomicName id="FAF5B18FED678D7DFA6FA6D5D3CBFB4E" authorityName="Walckenaer" authorityYear="1805" box="[1419,1490,1199,1222]" class="Arachnida" family="Eresidae" genus="Eresus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="0F81161EED678D7DFA6FA6D5D3CBFB4E" box="[1419,1490,1199,1222]" italics="true" pageId="12" pageNumber="13">Eresus</emphasis>
</taxonomicName>
. The details of a retreat are illustrated in
<figureCitation id="A5CED689ED678D7DFC71A6AAD222FB60" box="[917,1083,1232,1256]" captionStart="Figure 10" captionStartId="13.[444,514,843,865]" captionTargetBox="[205,1382,261,807]" captionTargetId="figure-199@13.[205,1391,251,807]" captionTargetPageId="13" captionText="Figure 10. Retreats of Loureedia colleni from Spain. (A,B) retreat layout, dorsal and lateral views (illustration: Thomas Romanoff); (C) female with the open hatch; (D) hatch being closed; (E) closed hatch. Legend (A,B): 1—main lodge; 2—external access tunnels; 3—hunting canopy and radial silk lines; 4—“retreat” hatch; 5—annex “retreat” lodge." figureDoi="http://doi.org/10.5281/zenodo.7630166" httpUri="https://zenodo.org/record/7630166/files/figure.png" pageId="12" pageNumber="13">Figure 10AE</figureCitation>
. Sectional views of the canvas and lodge assembly (between the stone and the ground) are depicted in
<figureCitation id="A5CED689ED678D7DFB20A688D344FA82" box="[1220,1373,1266,1290]" captionStart="Figure 10" captionStartId="13.[444,514,843,865]" captionTargetBox="[205,1382,261,807]" captionTargetId="figure-199@13.[205,1391,251,807]" captionTargetPageId="13" captionText="Figure 10. Retreats of Loureedia colleni from Spain. (A,B) retreat layout, dorsal and lateral views (illustration: Thomas Romanoff); (C) female with the open hatch; (D) hatch being closed; (E) closed hatch. Legend (A,B): 1—main lodge; 2—external access tunnels; 3—hunting canopy and radial silk lines; 4—“retreat” hatch; 5—annex “retreat” lodge." figureDoi="http://doi.org/10.5281/zenodo.7630166" httpUri="https://zenodo.org/record/7630166/files/figure.png" pageId="12" pageNumber="13">Figure 10A,B</figureCitation>
. The main lodge, located below the surface of the stone, is the main living space. Females have been observed in captivity to consume their prey and sometimes molt or copulate (sharing this lodge with the male) in this area. Hunting behavior is mainly sit-and-wait. Vibration is received from the external radial silk lines. The periphery of this lodge consists of dense cribellate silk, mixed with small pieces of agglomerated soil, anchored both to the ground and under the stone. From the main lodge, two separate exits with two capture canopies exist. The silk retreat is covered with a trapdoor-like hatch made of thick cribellate silk with soil particles in it (
<figureCitation id="A5CED689ED678D7DFD2AA787D568F99E" box="[718,881,1533,1558]" captionStart="Figure 10" captionStartId="13.[444,514,843,865]" captionTargetBox="[205,1382,261,807]" captionTargetId="figure-199@13.[205,1391,251,807]" captionTargetPageId="13" captionText="Figure 10. Retreats of Loureedia colleni from Spain. (A,B) retreat layout, dorsal and lateral views (illustration: Thomas Romanoff); (C) female with the open hatch; (D) hatch being closed; (E) closed hatch. Legend (A,B): 1—main lodge; 2—external access tunnels; 3—hunting canopy and radial silk lines; 4—“retreat” hatch; 5—annex “retreat” lodge." figureDoi="http://doi.org/10.5281/zenodo.7630166" httpUri="https://zenodo.org/record/7630166/files/figure.png" pageId="12" pageNumber="13">Figure 10CE</figureCitation>
).
</paragraph>
<caption id="698A9A84ED668D7CFE58A131D5CAFC40" ID-DOI="http://doi.org/10.5281/zenodo.7630166" ID-Zenodo-Dep="7630166" httpUri="https://zenodo.org/record/7630166/files/figure.png" pageId="13" pageNumber="14" startId="13.[444,514,843,865]" targetBox="[205,1382,261,807]" targetPageId="13">
<paragraph id="3D4ACA0CED668D7CFE58A131D5CAFC40" blockId="13.[443,1492,843,968]" pageId="13" pageNumber="14">
<emphasis id="0F81161EED668D7CFE58A131D431FCE8" bold="true" box="[444,552,843,865]" pageId="13" pageNumber="14">Figure 10.</emphasis>
Retreats of
<taxonomicName id="FAF5B18FED668D7CFD54A131D541FCE8" authorityName=": Zamani and Marusik" authorityYear="2020" box="[688,856,843,864]" class="Arachnida" family="Eresidae" genus="Loureedia" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="colleni">
<emphasis id="0F81161EED668D7CFD54A131D541FCE8" box="[688,856,843,864]" italics="true" pageId="13" pageNumber="14">Loureedia colleni</emphasis>
</taxonomicName>
from Spain. (
<emphasis id="0F81161EED668D7CFC17A131D21FFCE8" bold="true" box="[1011,1030,843,864]" pageId="13" pageNumber="14">A</emphasis>
,
<emphasis id="0F81161EED668D7CFBE8A131D205FCE8" bold="true" box="[1036,1052,843,864]" pageId="13" pageNumber="14">B</emphasis>
) retreat layout, dorsal and lateral views (illustration: Thomas Romanoff); (
<emphasis id="0F81161EED668D7CFCC1A117D52FFC0B" bold="true" box="[805,822,877,899]" pageId="13" pageNumber="14">C</emphasis>
) female with the open hatch; (
<emphasis id="0F81161EED668D7CFB9DA117D294FC0A" bold="true" box="[1145,1165,877,898]" pageId="13" pageNumber="14">D</emphasis>
) hatch being closed; (
<emphasis id="0F81161EED668D7CFA90A117D39AFC0A" bold="true" box="[1396,1411,877,898]" pageId="13" pageNumber="14">E</emphasis>
) closed hatch. Legend (
<emphasis id="0F81161EED668D7CFD86A1F5D46CFC2C" bold="true" box="[610,629,911,932]" pageId="13" pageNumber="14">A</emphasis>
,
<emphasis id="0F81161EED668D7CFD9EA1EAD493FC2D" bold="true" box="[634,650,912,933]" pageId="13" pageNumber="14">B</emphasis>
):
<emphasis id="0F81161EED668D7CFD7AA1F5D4B3FC2C" bold="true" box="[670,682,911,932]" pageId="13" pageNumber="14">1</emphasis>
—main lodge;
<emphasis id="0F81161EED668D7CFCA2A1F5D54BFC2C" bold="true" box="[838,850,911,932]" pageId="13" pageNumber="14">2</emphasis>
—external access tunnels;
<emphasis id="0F81161EED668D7CFB8DA1F5D26CFC2C" bold="true" box="[1129,1141,911,932]" pageId="13" pageNumber="14">3</emphasis>
—hunting canopy and radial silk lines;
<emphasis id="0F81161EED668D7CFE1DA1C8D41CFC4F" bold="true" box="[505,517,946,967]" pageId="13" pageNumber="14">4</emphasis>
—“retreat” hatch;
<emphasis id="0F81161EED668D7CFD21A1C8D4C8FC4F" bold="true" box="[709,721,946,967]" pageId="13" pageNumber="14">5</emphasis>
—annex “retreat” lodge.
</paragraph>
</caption>
<paragraph id="3D4ACA0CED668D7CFE10A196D28DFB8C" blockId="13.[443,1497,1004,2065]" box="[500,1172,1004,1028]" pageId="13" pageNumber="14">Phenology. Males are active during FebruaryNovember.</paragraph>
</subSubSection>
<subSubSection id="75EF9987ED668D7CFE10A677D525FBCF" pageId="13" pageNumber="14" type="distribution">
<paragraph id="3D4ACA0CED668D7CFE10A677D525FBCF" blockId="13.[443,1497,1004,2065]" pageId="13" pageNumber="14">
Distribution.
<collectingCountry id="45E28A9CED668D7CFD76A677D4CDFBAD" box="[658,724,1037,1061]" name="Spain" pageId="13" pageNumber="14">Spain</collectingCountry>
(Albacete, Alicante, Almería, Ciudad Real, Granada,
<collectingRegion id="FF3104EEED668D7CFAA5A677D385FBAD" box="[1345,1436,1037,1061]" country="Spain" name="Madrid" pageId="13" pageNumber="14">Madrid</collectingRegion>
, and
<collectingRegion id="FF3104EEED668D7CFE58A654D409FBCE" box="[444,528,1070,1094]" country="Spain" name="Murcia" pageId="13" pageNumber="14">Murcia</collectingRegion>
provinces) (see
<figureCitation id="A5CED689ED668D7CFD29A654D535FBCE" box="[717,812,1070,1094]" captionStart="Figure 5" captionStartId="6.[444,514,820,842]" captionTargetBox="[128,1458,259,784]" captionTargetId="figure-183@6.[128,1458,259,784]" captionTargetPageId="6" captionText="Figure 5. Distribution records of species of Loureedia. Color codes: yellow—L. colleni, blue—L. maroccana, white—L. lucasi, green—L. jerbae, khaki—L. annulipes (SAJ264 not shown), magenta—L. phoenixi, black dot—unidentified Loureedia specimen from Jordan [7]." figureDoi="http://doi.org/10.5281/zenodo.7630156" httpUri="https://zenodo.org/record/7630156/files/figure.png" pageId="13" pageNumber="14">Figure 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>