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<document id="14C4C6815268641E4B9772A0126571D1" ID-CLB-Dataset="21992" ID-DOI="10.5852/ejt.2017.355" ID-GBIF-Dataset="d8be473a-53c9-4ce6-8bd4-ec818d87adc0" ID-ISSN="2118-9773" ID-Zenodo-Dep="3836313" ID-ZooBank="80D35410-38CD-42FB-AFAF-605B24FC0A8B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589997846232" checkinUser="carolina" docAuthor="Jocque, Merlijn, Wellens, Siel, Andrianarivosoa, J. D., Rakotondraparany, F., Seing, Sam The &amp; Jocqué, Rudy" docDate="2017" docId="0A076352FF98FFC46D442D7B6D03FD97" docLanguage="en" docName="ejt-2017-355.pdf.imf" docOrigin="European Journal of Taxonomy 355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ocyale ghost Jocque M. &amp; Jocque R. 2017, sp. nov." docType="treatment" docUuid="37AAF237-A0C7-40D1-9BA8-6FF3D4864D79" docUuidSource="ZooBank" docVersion="7" lastPageNumber="11" masterDocId="F63E1B2AFF9CFFCF6F4A2F006E45FFF4" masterDocTitle="A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus" masterLastPageNumber="13" masterPageNumber="1" pageNumber="5" updateTime="1698840500717" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A49B875C954EC80547A016DD926B154D">A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus</mods:title>
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<mods:namePart id="FEC16589208D8E174FB5B095F1E45E5B">Jocque, Merlijn</mods:namePart>
<mods:nameIdentifier id="431BE80442E6A06E6851B0AE5CBA69EC" type="LSID">6F4C1B5C-BDD9-4217-8F29-95ADA5E9F4D3</mods:nameIdentifier>
<mods:affiliation id="CEE1E9E1410B006D6A177850564990BC">Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium. &amp; Operation Wallacea, Wallacea House, Old Bolingbroke, Lincolnshire, PE 23 4 EX United Kingdom. &amp; Aquatic and Terrestrial Ecology (ATECO), Royal Belgian Institute of Natural Sciences (RBINS), Vautierstraat 29, 1000 Brussels, Belgium. &amp; Email: merlijnjocque @ gmail. com &amp; urn: lsid: zoobank. org: author: 6 F 4 C 1 B 5 C-BDD 9 - 4217 - 8 F 29 - 95 ADA 5 E 9 F 4 D 3</mods:affiliation>
<mods:nameIdentifier id="CDEE70B47FB10B38894834E8D33591C6" type="email">merlijnjocque@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5720C39B391863301A4004240F0A2955">Wellens, Siel</mods:namePart>
<mods:nameIdentifier id="DF073D2F344C7DA50C5F712EF36232E1" type="LSID">B549CCD8-9A34-4FCD-996F-C49633331D33</mods:nameIdentifier>
<mods:affiliation id="D488608B196F8A5C8A8B842FF9D5E21F">Biodiversity Inventory for Conservation (BINCO), Walmersumstraat 44, 3380 Glabbeek, Belgium. &amp; Operation Wallacea, Wallacea House, Old Bolingbroke, Lincolnshire, PE 23 4 EX United Kingdom. &amp; Email: siel. wellens @ gmail. com &amp; urn: lsid: zoobank. org: author: B 549 CCD 8 - 9 A 34 - 4 FCD- 996 F-C 49633331 D 33</mods:affiliation>
<mods:nameIdentifier id="31B3A82098BD63C745D54ABA4A9C3355" type="email">siel.wellens@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EFAC13E8746FCE382AC11BA82FF01913">Andrianarivosoa, J. D.</mods:namePart>
<mods:nameIdentifier id="EF38407E601AB9BF623CC1CE560AAC9F" type="LSID">8F3B36C3-D80B-455D-B48E-50F27491167C</mods:nameIdentifier>
<mods:affiliation id="4F248ACA294CA236CD7648B3D1841689">Mention Zoologie et Biologie Animale (MZBA), Université d’Antananarivo, Madagascar. &amp; Email: davsandrianarivosoa @ gmail. com &amp; urn: lsid: zoobank. org: author: 8 F 3 B 36 C 3 - D 80 B- 455 D-B 48 E- 50 F 27491167 C</mods:affiliation>
<mods:nameIdentifier id="49CCF194251B0BDFD23DD8453261ABF9" type="email">davsandrianarivosoa@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D828127156B02E3242245D45B158974B">Rakotondraparany, F.</mods:namePart>
<mods:nameIdentifier id="64FBD66D291E36FEC30A8D7058A19C33" type="LSID">487CF2F0-E5AC-4DF0-A2B2-4BCE6C7F8CB3</mods:nameIdentifier>
<mods:affiliation id="7F7A233281052AB4FEC1D9021404DE7B">Mention Zoologie et Biologie Animale (MZBA), Université d’Antananarivo, Madagascar. &amp; Email: felitenrecs @ gmail. com &amp; urn: lsid: zoobank. org: author: 487 CF 2 F 0 - E 5 AC- 4 DF 0 - A 2 B 2 - 4 BCE 6 C 7 F 8 CB 3</mods:affiliation>
<mods:nameIdentifier id="098EEF2E297EE61EE14EDD127EAE8A44" type="email">felitenrecs@gmail.com</mods:nameIdentifier>
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<mods:namePart id="31C0DEEBCF1D5CD66CB745B3FF45ACE6">Seing, Sam The</mods:namePart>
<mods:affiliation id="ECA0679FE5C81F0A3213A5B96F82A969">Development and Biodiversity Conservation Action for Madagascar (DBCAM). Lot II A 93 L, Anjanahary, Antananarivo, Madagascar. &amp; Email: sam _ theseing @ yahoo. fr &amp; urn: lsid: zoobank. org: author: 28 F 7 A 019 - 1581 - 4 C 8 B-B 0 CE- 1 C 4948287 F 65</mods:affiliation>
<mods:nameIdentifier id="0ADF6A2D23D6189FCD95680ED279A672" type="email">sam_theseing@yahoo.fr</mods:nameIdentifier>
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<mods:namePart id="2789A71CB60BA45D6CC364A7BA04E8E2">Jocqué, Rudy</mods:namePart>
<mods:affiliation id="00C9947F646DBEBA4D9F8C21C9ED7A45">Royal Museum for Central Africa, Leuvense Steenweg 13, 3080 Tervuren, Belgium. &amp; urn: lsid: zoobank. org: author: CF 15016 C- 8 CD 1 - 4 C 9 D- 9021 - 44 CA 7 DC 7 A 5 D 5 &amp; Corresponding author: rudy. jocque @ africamuseum. be</mods:affiliation>
<mods:nameIdentifier id="00F6EF1418E56358B1A0033983F25F36" type="email">rudy.jocque@africamuseum.be</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="CF6C243DA268CAA1D3F56972E353FC8A">text</mods:typeOfResource>
<mods:relatedItem id="9621F59F4BAF75CFAB402A338B381EDD" type="host">
<mods:titleInfo id="8E70CD17963860C2B55BF432D2F468EF">
<mods:title id="16D67A1988188557F843BE436C348EAB">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="10F346F7E61F979862226E2BF80D2742">
<mods:date id="263EE2D7C4CDBA4B45946858C530FD0C">2017</mods:date>
<mods:detail id="195BB353D3F78BC96A9E899E3EB02627" type="pubDate">
<mods:number id="78FCBF13AA3F530E0FF84E63481CDCFD">2017-10-03</mods:number>
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<mods:detail id="E0AB88CC1EB24573439534D0E0E80431" type="issue">
<mods:number id="7300E07F7029D5932C97B9F8EE597D8F">355</mods:number>
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<mods:start id="1A093D7B80FEF42EBD74EDD5FB9DE4EF">1</mods:start>
<mods:end id="518EA31BC9AA32EDE9E632C82E0678A4">13</mods:end>
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<mods:classification id="F1DE0CED001C7EA176DAAB51BCE8B7EB">journal article</mods:classification>
<mods:identifier id="1D169F9CCBB26472911C15C7133B064B" type="CLB-Dataset">21992</mods:identifier>
<mods:identifier id="973BCCC38FDAF8715ABC28A4308E5705" type="DOI">10.5852/ejt.2017.355</mods:identifier>
<mods:identifier id="26893DCAA18F39C24C82D5D7F3F675F0" type="GBIF-Dataset">d8be473a-53c9-4ce6-8bd4-ec818d87adc0</mods:identifier>
<mods:identifier id="2E702412F46CAD7F4F0F7920045699FE" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="3224FC6202270330FB369D852C762F4C" type="Zenodo-Dep">3836313</mods:identifier>
<mods:identifier id="FAB5D1DC70945DE5340EA84796A9D2F4" type="ZooBank">80D35410-38CD-42FB-AFAF-605B24FC0A8B</mods:identifier>
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<treatment id="0A076352FF98FFC46D442D7B6D03FD97" ID-DOI="http://doi.org/10.5281/zenodo.3851682" ID-GBIF-Taxon="164243581" ID-Zenodo-Dep="3851682" LSID="urn:lsid:zoobank.org:act:37AAF237-A0C7-40D1-9BA8-6FF3D4864D79" httpUri="http://treatment.plazi.org/id/0A076352FF98FFC46D442D7B6D03FD97" lastPageId="11" lastPageNumber="11" pageId="4" pageNumber="5">
<subSubSection id="CAB481CFFF98FFCB6D442D7B6A63FD62" box="[526,1062,635,662]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8211D244FF98FFCB6D442D7B6A63FD62" blockId="4.[363,1224,635,733]" box="[526,1062,635,662]" pageId="4" pageNumber="5">
<heading id="D9596528FF98FFCB6D442D7B6A63FD62" box="[526,1062,635,662]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="45AEA9C7FF98FFCB6D442D7B6D81FD62" ID-CoL="48L73" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[526,964,635,662]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF98FFCB6D442D7B6CECFD62" bold="true" box="[526,681,635,662]" italics="true" pageId="4" pageNumber="5">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF98FFCB6C812D7C6A63FD62" box="[971,1062,636,662]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF98FFCB6E212D9F6A59FD28" pageId="4" pageNumber="5" type="description">
<paragraph id="8211D244FF98FFCB6E212D9F6A8DFD4D" blockId="4.[363,1224,635,733]" box="[363,1224,671,697]" pageId="4" pageNumber="5">
<uri id="F63FDE46FF98FFCB6E212D9F6A8DFD4D" box="[363,1224,671,697]" pageId="4" pageNumber="5">
urn:lsid:zoobank.org:act:
<uuid id="F608E891FF98FFCB6DC72D9F6A8DFD4D" box="[653,1224,671,697]" pageId="4" pageNumber="5">37AAF237-A0C7-40D1-9BA8-6FF3D4864D79</uuid>
</uri>
</paragraph>
<paragraph id="8211D244FF98FFCB6D522DC26A59FD28" blockId="4.[363,1224,635,733]" box="[536,1052,706,733]" pageId="4" pageNumber="5">
<figureCitation id="1A95CEC1FF98FFCB6D522DC26C24FD28" box="[536,609,706,732]" captionStart="Fig" captionStartId="1.[189,232,2000,2026]" captionTargetBox="[189,1398,1008,1961]" captionTargetId="figure@1.[187,1399,1006,1962]" captionTargetPageId="1" captionText="Fig. 1. Type locality of Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3836315" httpUri="https://zenodo.org/record/3836315/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
,
<figureCitation id="1A95CEC1FF98FFCB6D252DC26CC5FD28" box="[623,640,706,732]" captionStart="Fig" captionStartId="5.[190,233,1965,1991]" captionTargetBox="[190,1398,1023,1927]" captionTargetId="figure@5.[187,1401,1021,1930]" captionTargetPageId="5" captionText="Fig. 2. Habitat on type locality of Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. (photo by MJ, July 2012)." figureDoi="http://doi.org/10.5281/zenodo.3836317" httpUri="https://zenodo.org/record/3836317/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
,
<figureCitation id="1A95CEC1FF98FFCB6DC62DC26CDDFD28" box="[652,664,706,732]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
A–F, 4A–D, 5A–F, 6A–C, 7A–C
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF98FFCB6FF72C046F17FC28" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8211D244FF98FFCB6FF72C046F73FCEA" blockId="4.[189,310,772,798]" box="[189,310,772,798]" pageId="4" pageNumber="5">
<heading id="D9596528FF98FFCB6FF72C046F73FCEA" bold="true" box="[189,310,772,798]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B0DA0E56FF98FFCB6FF72C046F73FCEA" bold="true" box="[189,310,772,798]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF98FFCB6FF72C336F17FC28" blockId="4.[189,1399,819,988]" pageId="4" pageNumber="5">
Males of
<taxonomicName id="45AEA9C7FF98FFCB6E612C346FCAFCB9" authority="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué, 2017" authorityName="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué" authorityYear="2017" box="[299,399,819,846]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF98FFCB6E612C346FCAFCB9" box="[299,399,819,846]" italics="true" pageId="4" pageNumber="5">O. ghost</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF98FFCB6EDF2C346FB5FCBA" box="[405,496,820,846]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
can be recognized by details of the male palp: the tegulum is restricted to the prolateral side of the bulbus, the distal prong of the palea appendage is much narrower than the proximal one and the MA with the perpendicular prong is rectangular. Females are characterized by the epigyne in which the T-shaped posterior sclerite is fully exposed and not covered with setae as in other species in the genus.
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF98FFCB6FF72B036C8EFB67" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8211D244FF98FFCB6FF72B036F01FBE9" blockId="4.[189,324,1027,1053]" box="[189,324,1027,1053]" pageId="4" pageNumber="5">
<heading id="D9596528FF98FFCB6FF72B036F01FBE9" bold="true" box="[189,324,1027,1053]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B0DA0E56FF98FFCB6FF72B036F01FBE9" bold="true" box="[189,324,1027,1053]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF98FFCB6FF72B326C8EFB67" blockId="4.[189,1399,1074,1172]" pageId="4" pageNumber="5">
The species name ‘ghost’ refers to the fully white appearance of this spider. Additional reference is made to the large white direwolf ‘Ghost’ in
<emphasis id="B0DA0E56FF98FFCB6D372B566D0CFB84" box="[637,841,1110,1136]" italics="true" pageId="4" pageNumber="5">Game of Thrones</emphasis>
, the first book in the series of fantasy novels
<emphasis id="B0DA0E56FF98FFCB6A2F2B566FE8FB67" italics="true" pageId="4" pageNumber="5">A Song of Ice and Fire</emphasis>
by George R.R. Martin.
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF98FFCB6FF72BBB6C0BF972" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8211D244FF98FFCB6FF72BBB6F29FB21" blockId="4.[189,364,1211,1237]" box="[189,364,1211,1237]" pageId="4" pageNumber="5">
<heading id="D9596528FF98FFCB6FF72BBB6F29FB21" bold="true" box="[189,364,1211,1237]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B0DA0E56FF98FFCB6FF72BBB6F29FB21" bold="true" box="[189,364,1211,1237]" pageId="4" pageNumber="5">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="32C6D819FF98FFCB6FA02BEA6AFEFAB8" ID-GBIF-Occurrence="2621113301" collectingDate="2016-07-05" collectionCode="MRAC" collectorName="S. Wellens" country="Madagascar" latitude="-15.49275" location="Lake" longLatPrecision="1" longitude="46.651142" municipality="Matsedroy" pageId="4" pageNumber="5" specimenCode="MRAC 245337" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga Prov." typeStatus="holotype">
<paragraph id="8211D244FF98FFCB6FA02BEA6F19FAF0" blockId="4.[189,1398,1258,1356]" box="[234,348,1258,1284]" pageId="4" pageNumber="5">
<emphasis id="B0DA0E56FF98FFCB6FA02BEA6F19FAF0" bold="true" box="[234,348,1258,1284]" pageId="4" pageNumber="5">
<typeStatus id="5D156CE6FF98FFCB6FA02BEA6F19FAF0" box="[234,348,1258,1284]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8211D244FF98FFCB6FF72A0E6AFEFAB8" blockId="4.[189,1398,1258,1356]" pageId="4" pageNumber="5">
<collectingCountry id="FAB992D4FF98FFCB6FF72A0E6FD7FADC" box="[189,402,1294,1320]" name="Madagascar" pageId="4" pageNumber="5">MADAGASCAR</collectingCountry>
:
<specimenCount id="94A819CDFF98FFCB6ED62A0E6FF1FADC" box="[412,436,1294,1320]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
,
<collectingRegion id="406A1CA6FF98FFCB6E8A2A0E6CCFFADC" box="[448,650,1294,1320]" country="Madagascar" name="Mahajanga" pageId="4" pageNumber="5">Mahajanga Prov.</collectingRegion>
,
<collectingMunicipality id="6275483EFF98FFCB6DDE2A0E6D56FADC" box="[660,787,1294,1320]" pageId="4" pageNumber="5">Matsedroy</collectingMunicipality>
,
<location id="8771849FFF98FFCB6C542A0E6D1CFADC" LSID="urn:lsid:plazi:treatment:0A076352FF98FFC46D442D7B6D03FD97:8771849FFF98FFCB6C542A0E6D1CFADC" box="[798,857,1294,1320]" country="Madagascar" latitude="-15.49275" longLatPrecision="1" longitude="46.651142" municipality="Matsedroy" name="Lake" pageId="4" pageNumber="5" stateProvince="Mahajanga Prov.">Lake</location>
2,
<geoCoordinate id="E79AB483FF98FFCB6C362A0C6A64FADC" box="[892,1057,1292,1320]" degrees="15" direction="south" minutes="29" orientation="latitude" pageId="4" pageNumber="5" precision="1" seconds="33.9" value="-15.49275">15°29′33.9″ S</geoCoordinate>
,
<geoCoordinate id="E79AB483FF98FFCB6B672A0C6AA6FADC" box="[1069,1251,1292,1320]" degrees="046" direction="east" minutes="39" orientation="longitude" pageId="4" pageNumber="5" precision="1" seconds="04.1" value="46.651142">046°39′04.1″ E</geoCoordinate>
,
<date id="F610F484FF98FFCB6BA52A0D6B37FADC" box="[1263,1394,1293,1320]" pageId="4" pageNumber="5" value="2016-07-05">
<collectingDate id="E6540D6CFF98FFCB6BA52A0D6B37FADC" box="[1263,1394,1293,1320]" pageId="4" pageNumber="5" value="2016-07-05">5 Jul. 2016</collectingDate>
</date>
, 19:55–20:45, hand collected on sandy beach (
<figureCitation id="1A95CEC1FF98FFCB6D9C2A316D5BFABF" box="[726,798,1329,1355]" captionStart="Fig" captionStartId="5.[190,233,1965,1991]" captionTargetBox="[190,1398,1023,1927]" captionTargetId="figure@5.[187,1401,1021,1930]" captionTargetPageId="5" captionText="Fig. 2. Habitat on type locality of Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. (photo by MJ, July 2012)." figureDoi="http://doi.org/10.5281/zenodo.3836317" httpUri="https://zenodo.org/record/3836317/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
),
<collectorName id="2F5BB792FF98FFCB6C7E2A316DF4FABF" box="[820,945,1329,1355]" pageId="4" pageNumber="5">S. Wellens</collectorName>
leg. (
<specimenCode id="D2087A3FFF98FFCB6CB82A316AEAFAB8" box="[1010,1199,1329,1356]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC 245337</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8211D244FF98FFCB6FA02A736F22FA79" blockId="4.[189,1399,1395,1671]" box="[234,359,1395,1421]" pageId="4" pageNumber="5">
<emphasis id="B0DA0E56FF98FFCB6FA02A736F22FA79" bold="true" box="[234,359,1395,1421]" pageId="4" pageNumber="5">
<typeStatus id="5D156CE6FF98FFCB6FA02A736F22FA79" box="[234,359,1395,1421]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8211D244FF98FFCB6FF72A976C0BF972" blockId="4.[189,1399,1395,1671]" pageId="4" pageNumber="5">
<collectingCountry id="FAB992D4FF98FFCB6FF72A976FD7FA45" box="[189,402,1431,1457]" name="Madagascar" pageId="4" pageNumber="5">MADAGASCAR</collectingCountry>
:
<specimenCount id="94A819CDFF98FFCB6EEA2A976F8BFA45" box="[416,462,1431,1457]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
, 2 juvs, same data as
<typeStatus id="5D156CE6FF98FFCB6D942A976D00FA45" box="[734,837,1431,1457]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(
<collectionCode id="E4BF4A81FF98FFCB6C1C2A976DEBFA45" box="[854,942,1431,1457]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245338);
<specimenCount id="94A819CDFF98FFCB6B662A976A37FA45" box="[1068,1138,1431,1457]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
, 2 juvs, same data as preceding (
<collectionCode id="E4BF4A81FF98FFCB6E0B2ABA6FDCFA20" box="[321,409,1466,1492]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245340);
<specimenCount id="94A819CDFF98FFCB6D5E2ABA6C1DFA21" box="[532,600,1466,1493]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
, 2 juvs, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6C2E2ABA6DF9FA20" box="[868,956,1466,1492]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245341);
<specimenCount id="94A819CDFF98FFCB6B7D2ABA6A3EFA21" box="[1079,1147,1466,1493]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
, 2 juvs, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6F8F2ADE6F58FA0C" box="[197,285,1502,1528]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245342);
<specimenCount id="94A819CDFF98FFCB6EDF2ADE6F84FA0C" box="[405,449,1502,1528]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6D3B2ADE6C8CFA0C" box="[625,713,1502,1528]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245347);
<specimenCount id="94A819CDFF98FFCB6C082ADE6D35FA0C" box="[834,880,1502,1528]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
, 6 juvs, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6B392ADE6A8EFA0C" box="[1139,1227,1502,1528]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245348);
<specimenCount id="94A819CDFF98FFCB6A092ADE6B34FA0C" box="[1347,1393,1502,1528]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
, 3 juvs, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6EF529026C52F9E8" box="[447,535,1538,1564]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245361);
<specimenCount id="94A819CDFF98FFCB6DDF29026C9EF9E8" box="[661,731,1538,1564]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
, 2 juvs, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6CA729026A00F9E8" box="[1005,1093,1538,1564]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245350);
<specimenCount id="94A819CDFF98FFCB6B8E29026AB7F9E8" box="[1220,1266,1538,1564]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
, 5 juvs, as preceding except
<date id="F610F484FF98FFCB6EC629256C58F9CB" box="[396,541,1573,1599]" pageId="4" pageNumber="5" value="2016-07-12">12 Jul. 2016</date>
(
<collectionCode id="E4BF4A81FF98FFCB6D6629256CC1F9CB" box="[556,644,1573,1599]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245339);
<specimenCount id="94A819CDFF98FFCB6DB429256D6FF9B4" box="[766,810,1573,1600]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
, 5 juvs, as preceding except
<date id="F610F484FF98FFCB6B3429256B4AF9CB" box="[1150,1295,1573,1599]" pageId="4" pageNumber="5" value="2016-07-22">22 Jul. 2016</date>
(
<collectionCode id="E4BF4A81FF98FFCB6A5429256B33F9CB" box="[1310,1398,1573,1599]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="4" pageNumber="5" type="Museum">MRAC</collectionCode>
245351);
<specimenCount id="94A819CDFF98FFCB6E6529496F19F997" box="[303,348,1609,1635]" pageId="4" pageNumber="5" type="female">1 ♀</specimenCount>
, 4 juvs, as preceding except
<date id="F610F484FF98FFCB6DFC29496D0AF996" box="[694,847,1608,1635]" pageId="4" pageNumber="5" value="2016-06-24">24 Jun. 2016</date>
(
<collectionCode id="E4BF4A81FF98FFCB6C2A29496DE4F997" box="[864,929,1609,1635]" pageId="4" pageNumber="5">ZBA</collectionCode>
);
<specimenCount id="94A819CDFF98FFCB6CFC29496DA0F997" box="[950,997,1609,1635]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
,
<specimenCount id="94A819CDFF98FFCB6CB929496A72F997" box="[1011,1079,1609,1635]" pageId="4" pageNumber="5" type="female">4 ♀♀</specimenCount>
, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6BA629496B68F997" box="[1260,1325,1609,1635]" pageId="4" pageNumber="5">ZBA</collectionCode>
);
<specimenCount id="94A819CDFF98FFCB6A0829496B34F997" box="[1346,1393,1609,1635]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
,
<specimenCount id="94A819CDFF98FFCB6FF7296C6F45F973" box="[189,256,1644,1671]" pageId="4" pageNumber="5" type="female">2 ♀♀</specimenCount>
,
<specimenCount id="94A819CDFF98FFCB6E44296C6F17F972" box="[270,338,1644,1670]" pageId="4" pageNumber="5" type="generic">1 juv.</specimenCount>
, as preceding (
<collectionCode id="E4BF4A81FF98FFCB6D4B296C6C07F972" box="[513,578,1644,1670]" pageId="4" pageNumber="5">ZBA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF98FFC66FF729AE6D8EFC71" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="description">
<paragraph id="8211D244FF98FFCB6FF729AE6F0BF93C" blockId="4.[189,334,1710,1736]" box="[189,334,1710,1736]" pageId="4" pageNumber="5">
<heading id="D9596528FF98FFCB6FF729AE6F0BF93C" bold="true" box="[189,334,1710,1736]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B0DA0E56FF98FFCB6FF729AE6F0BF93C" bold="true" box="[189,334,1710,1736]" pageId="4" pageNumber="5">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF98FFCB6FA029DD6FEEF90C" blockId="4.[189,1399,1757,1855]" box="[234,427,1757,1784]" pageId="4" pageNumber="5">
<emphasis id="B0DA0E56FF98FFCB6FA029DD6F6FF903" bold="true" box="[234,298,1757,1783]" pageId="4" pageNumber="5">Male</emphasis>
(
<typeStatus id="5D156CE6FF98FFCB6E7329DD6FE6F90C" box="[313,419,1757,1784]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8211D244FF98FFCB6FF728016D95F8CA" blockId="4.[189,1399,1757,1855]" pageId="4" pageNumber="5">
MEASUREMENTS. Total length 15.35; carapace 8.77 long, 2.27 high, 6.58 wide, narrowed to
<quantity id="45567FA1FF98FFCB6BB828006B00F8EE" box="[1266,1349,1792,1819]" metricMagnitude="-1" metricUnit="m" metricValue="1.0033" pageId="4" pageNumber="5" unit="in" value="3.95">3.95 in</quantity>
eye region; labium 1.14 wide, 0.79 long; sternum 3.68 long, 3.07 wide.
</paragraph>
<paragraph id="8211D244FF98FFCA6FF728666DC2FDE8" blockId="4.[189,1399,1894,2027]" lastBlockId="5.[189,1399,267,540]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
COLOUR. Alive (
<figureCitation id="1A95CEC1FF98FFCB6E3028666F85F874" box="[378,448,1894,1920]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
A–E): almost entirely white with slightly contrasting darker spots as in ethanol specimens; eyes surrounded by yellow rings; cheliceral condyle reddish brown. In ethanol (
<figureCitation id="1A95CEC1FF98FFCB6BA228896B68F850" box="[1256,1325,1929,1956]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[194,1393,262,1925]" captionTargetId="figure@7.[191,1396,259,1928]" captionTargetPageId="7" captionText="Fig. 4. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–B. Holotype, ♂. A. Dorsal habitus. B. Ventral habitus. C–D. Paratype, ♀ (MRAC 245338). C. Dorsal habitus. D. Ventral habitus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836321" httpUri="https://zenodo.org/record/3836321/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
A–B): fairly different from colouration of live specimens; carapace brownish cream with interrupted black radiating striae, black fovea, two pairs of spots in front of fovea and dispersed irregular spots along margins; PEQ covered with white setae, tegument dark in its anterior half; chelicerae brown with dark setae as seen in frontal view, dark brown with black setae in ventral view; labium dark brown with cream crescent shape along anterior concave margin; sternum with strongly sinuous lateral margins, ending in long, tapered point; with dispersed short dark setae; legs formula IV-III-I-II; uniform cream with dispersed short, dark setae and dark spines; pedipalp: femur, patella and tibia cream, cymbium and bulbus contrasting dark brown; abdomen: dorsum cream with dispersed small dark spots, two reddish apodemes in anterior half and dispersed dark setae, sides and venter uniform cream; spinnerets: ALS dark, PLS and PMS pale on dorsal side, dark on ventral side.
</paragraph>
<paragraph id="8211D244FF99FFCA6FF72D446B06FDA9" blockId="5.[189,1347,579,606]" box="[189,1347,579,606]" pageId="5" pageNumber="6">EYES. AME: 0.35; ALE: 0.15; PME: 0.67: PLE: 0.61; eye rows: ALE: 1.51, PME: 1.51, PLE: 2.18.</paragraph>
<paragraph id="8211D244FF99FFCA6FF72D856F7AFD36" blockId="5.[189,1398,645,706]" pageId="5" pageNumber="6">CHELICERAE. With three teeth on retromargin, one small proximal tooth and one larger distal tooth on promargin.</paragraph>
<paragraph id="8211D244FF99FFCA6FF72DEA6DBAFCD3" blockId="5.[189,1399,745,807]" pageId="5" pageNumber="6">
LEGS. Spination of leg I (identical on both sides): femur pl4, d3, rl2; patella pl1, rl1; tibia pl2, d2, rl2, v2-2-2; metatarsus pl2, rl2, v2-2, dw5. Leg measurements: see
<tableCitation id="CF2CE7FFFF99FFCA6CEB2C0C6DBAFCD3" box="[929,1023,780,807]" captionStart="Table 1" captionStartId="8.[189,258,268,294]" captionTargetPageId="8" captionText="Table 1. Leg measurements of male holotype and female paratype (MRAC 245338)." httpUri="http://table.plazi.org/id/D6D182CCFF94FFC76FF72E0C6ADAFED2" pageId="5" pageNumber="6" tableUuid="D6D182CCFF94FFC76FF72E0C6ADAFED2">Table 1.</tableCitation>
</paragraph>
<paragraph id="8211D244FF99FFCA6FF72C4E6D9DFC25" blockId="5.[189,1399,846,978]" pageId="5" pageNumber="6">
PALP (
<figureCitation id="1A95CEC1FF99FFCA6E4D2C4E6F14FC9D" box="[263,337,846,873]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[192,1398,786,1853]" captionTargetId="figure@8.[186,1401,785,1858]" captionTargetPageId="8" captionText="Fig. 5. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–D. Palp, ♂, holotype. A. Ventral view. B. Same, detail. C. Lateral view. D. Same, detail. E–F. Epigyne, ♀, paratype (MRAC 245338). E. Ventral view. F. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="5" pageNumber="6">Figs 5</figureCitation>
A–D, 6A–C, 7A–B). Tegulum ribbed, developed on prolateral basal part of bulbus; palea with two prongs, proximal one broad and slightly curved, distal one thin and strongly curved; embolus originating on retrolateral part of palea and curved ventrally around it; MA large, with prolateral part a short hook, ventral part subrectangular, perpendicular to the former.
</paragraph>
<caption id="D6D182CCFF99FFCA6FF428AD6F4EF81F" ID-DOI="http://doi.org/10.5281/zenodo.3836317" ID-Zenodo-Dep="3836317" httpUri="https://zenodo.org/record/3836317/files/figure.png" pageId="5" pageNumber="6" startId="5.[190,233,1965,1991]" targetBox="[190,1398,1023,1927]" targetPageId="5">
<paragraph id="8211D244FF99FFCA6FF428AD6F4EF81F" blockId="5.[190,1399,1965,2027]" pageId="5" pageNumber="6">
<emphasis id="B0DA0E56FF99FFCA6FF428AD6F4BF833" bold="true" box="[190,270,1965,1991]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
Habitat on type locality of
<taxonomicName id="45AEA9C7FF99FFCA6D1528AD6A67F83C" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[607,1058,1965,1992]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF99FFCA6D1528AD6CB9F833" box="[607,764,1965,1991]" italics="true" pageId="5" pageNumber="6">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF99FFCA6B6128AE6ACDF83C" box="[1067,1160,1966,1992]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(photo by MJ, July 2012).
</paragraph>
</caption>
<caption id="D6D182CCFF9AFFC96FF729736F56F90C" ID-DOI="http://doi.org/10.5281/zenodo.3836319" ID-Zenodo-Dep="3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1651,1677]" targetBox="[189,1396,265,1611]" targetPageId="6">
<paragraph id="8211D244FF9AFFC96FF729736F56F90C" blockId="6.[189,1399,1651,1784]" pageId="6" pageNumber="7">
<emphasis id="B0DA0E56FF9AFFC96FF729736F4EF979" bold="true" box="[189,267,1651,1677]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
<taxonomicName id="45AEA9C7FF9AFFC96E5829736C8CF97A" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[274,713,1651,1678]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF9AFFC96E5829736FE8F979" box="[274,429,1651,1677]" italics="true" pageId="6" pageNumber="7">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF9AFFC96D9B29746D69F97A" box="[721,812,1652,1678]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
photographed at type locality.
<emphasis id="B0DA0E56FF9AFFC96BD229736AE8F979" bold="true" box="[1176,1197,1651,1677]" pageId="6" pageNumber="7">A</emphasis>
. Female habitus.
<emphasis id="B0DA0E56FF9AFFC96FF729976E94F945" bold="true" box="[189,209,1687,1713]" pageId="6" pageNumber="7">B</emphasis>
. Same, detail.
<emphasis id="B0DA0E56FF9AFFC96E3529966FD1F944" bold="true" box="[383,404,1686,1712]" pageId="6" pageNumber="7">C</emphasis>
. Female in sand retreat.
<emphasis id="B0DA0E56FF9AFFC96DFD29976C89F945" bold="true" box="[695,716,1687,1713]" pageId="6" pageNumber="7">D</emphasis>
. Female with spiderlings on abdomen.
<emphasis id="B0DA0E56FF9AFFC96BD629976AF5F945" bold="true" box="[1180,1200,1687,1713]" pageId="6" pageNumber="7">E</emphasis>
. Two males, one being eaten by the other.
<emphasis id="B0DA0E56FF9AFFC96EBA29BA6C47F920" bold="true" box="[496,514,1722,1748]" pageId="6" pageNumber="7">F</emphasis>
. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016).
</paragraph>
</caption>
<caption id="D6D182CCFF9BFFC86FF728AD6B32F81F" ID-DOI="http://doi.org/10.5281/zenodo.3836321" ID-Zenodo-Dep="3836321" httpUri="https://zenodo.org/record/3836321/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1965,1991]" targetBox="[194,1393,262,1925]" targetPageId="7">
<paragraph id="8211D244FF9BFFC86FF728AD6B32F81F" blockId="7.[189,1399,1965,2027]" pageId="7" pageNumber="8">
<emphasis id="B0DA0E56FF9BFFC86FF728AD6F4EF833" bold="true" box="[189,267,1965,1991]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="45AEA9C7FF9BFFC86E5928AD6C89F83C" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[275,716,1965,1992]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF9BFFC86E5928AD6FEBF833" box="[275,430,1965,1991]" italics="true" pageId="7" pageNumber="8">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF9BFFC86D9E28AE6D75F83C" box="[724,816,1966,1992]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B0DA0E56FF9BFFC86C7D28AD6D2BF833" bold="true" box="[823,878,1965,1991]" pageId="7" pageNumber="8">A–B</emphasis>
. Holotype, ♂.
<emphasis id="B0DA0E56FF9BFFC86B6A28AD6A70F833" bold="true" box="[1056,1077,1965,1991]" pageId="7" pageNumber="8">A</emphasis>
. Dorsal habitus.
<emphasis id="B0DA0E56FF9BFFC86BB528AE6B56F83C" bold="true" box="[1279,1299,1966,1992]" pageId="7" pageNumber="8">B</emphasis>
. Ventral habitus.
<emphasis id="B0DA0E56FF9BFFC86E5528D06F1DF81E" bold="true" box="[287,344,2000,2026]" pageId="7" pageNumber="8">C–D</emphasis>
. Paratype, ♀ (MRAC 245338).
<emphasis id="B0DA0E56FF9BFFC86D8128D06CA5F81E" bold="true" box="[715,736,2000,2026]" pageId="7" pageNumber="8">C</emphasis>
. Dorsal habitus.
<emphasis id="B0DA0E56FF9BFFC86CEE28D16DFCF81F" bold="true" box="[932,953,2001,2027]" pageId="7" pageNumber="8">D</emphasis>
. Ventral habitus. Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="D6D182CCFF94FFC76FF72E0C6ADAFED2" ID-Table-UUID="D6D182CCFF94FFC76FF72E0C6ADAFED2" box="[189,1183,267,294]" httpUri="http://table.plazi.org/id/D6D182CCFF94FFC76FF72E0C6ADAFED2" pageId="8" pageNumber="9" targetBox="[200,1346,337,739]" targetIsTable="true" targetPageId="8">
<paragraph id="8211D244FF94FFC76FF72E0C6ADAFED2" blockId="8.[189,1183,267,294]" box="[189,1183,267,294]" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76FF72E0C6F65FED1" bold="true" box="[189,288,267,294]" pageId="8" pageNumber="9">Table 1.</emphasis>
Leg measurements of male holotype and female paratype (MRAC 245338).
</paragraph>
</caption>
<paragraph id="8211D244FF94FFC76F822E516B07FD16" pageId="8" pageNumber="9">
<table id="F0AE20E4FF9400306F822E516B07FD17" box="[200,1346,337,739]" gridcols="7" gridrows="10" pageId="8" pageNumber="9">
<tr id="3C9ED006FF9400306F822E516B07FE98" box="[200,1346,337,364]" gridrow="0" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822E516C2EFE98" box="[200,619,337,364]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76F822E516F3EFE9F" bold="true" box="[200,379,337,363]" pageId="8" pageNumber="9">Male holotype</emphasis>
(MRAC 245337)
</th>
<th id="7F4FB97AFF9400306DCD2E516CFFFE98" box="[647,698,337,364]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76DCD2E516CE3FE9F" bold="true" box="[647,678,337,363]" pageId="8" pageNumber="9">Fe</emphasis>
</th>
<th id="7F4FB97AFF9400306C4C2E516D7CFE98" box="[774,825,337,364]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76C4C2E516D5DFE9F" bold="true" box="[774,792,337,363]" pageId="8" pageNumber="9">P</emphasis>
</th>
<th id="7F4FB97AFF9400306CCE2E516DF2FE98" box="[900,951,337,364]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76CCE2E516DDAFE9F" bold="true" box="[900,927,337,363]" pageId="8" pageNumber="9">Ti</emphasis>
</th>
<th id="7F4FB97AFF9400306B492E516A73FE98" box="[1027,1078,337,364]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76B492E516A6DFE9F" bold="true" box="[1027,1064,337,363]" pageId="8" pageNumber="9">Mt</emphasis>
</th>
<th id="7F4FB97AFF9400306BCB2E516AF1FE98" box="[1153,1204,337,364]" gridcol="5" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76BCB2E516ACEFE9F" bold="true" box="[1153,1163,337,363]" pageId="8" pageNumber="9">t</emphasis>
</th>
<th id="7F4FB97AFF9400306A4A2E516B07FE98" box="[1280,1346,337,364]" gridcol="6" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76A4A2E516B05FE9F" bold="true" box="[1280,1344,337,363]" pageId="8" pageNumber="9">Total</emphasis>
</th>
</tr>
<tr id="3C9ED006FF9400306F822E7B6B07FE62" box="[200,1346,379,406]" gridrow="1" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822E7B6C2EFE62" box="[200,619,379,406]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">I</th>
<td id="7F4FB97AFF9400306DCD2E7B6CFFFE62" box="[647,698,379,406]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">6.82</td>
<td id="7F4FB97AFF9400306C4C2E7B6D7CFE62" box="[774,825,379,406]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">3.83</td>
<td id="7F4FB97AFF9400306CCE2E7B6DF2FE62" box="[900,951,379,406]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">6.25</td>
<td id="7F4FB97AFF9400306B492E7B6A73FE62" box="[1027,1078,379,406]" gridcol="4" gridrow="1" pageId="8" pageNumber="9">5.68</td>
<td id="7F4FB97AFF9400306BCB2E7B6AF1FE62" box="[1153,1204,379,406]" gridcol="5" gridrow="1" pageId="8" pageNumber="9">4.83</td>
<td id="7F4FB97AFF9400306A4A2E7B6B07FE62" box="[1280,1346,379,406]" gridcol="6" gridrow="1" pageId="8" pageNumber="9">27.41</td>
</tr>
<tr id="3C9ED006FF9400306F822EA46B07FE4B" box="[200,1346,420,447]" gridrow="2" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822EA46C2EFE4B" box="[200,619,420,447]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">II</th>
<td id="7F4FB97AFF9400306DCD2EA46CFFFE4B" box="[647,698,420,447]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">6.82</td>
<td id="7F4FB97AFF9400306C4C2EA46D7CFE4B" box="[774,825,420,447]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">3.41</td>
<td id="7F4FB97AFF9400306CCE2EA46DF2FE4B" box="[900,951,420,447]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">5.96</td>
<td id="7F4FB97AFF9400306B492EA46A73FE4B" box="[1027,1078,420,447]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">6.39</td>
<td id="7F4FB97AFF9400306BCB2EA46AF1FE4B" box="[1153,1204,420,447]" gridcol="5" gridrow="2" pageId="8" pageNumber="9">4.54</td>
<td id="7F4FB97AFF9400306A4A2EA46B07FE4B" box="[1280,1346,420,447]" gridcol="6" gridrow="2" pageId="8" pageNumber="9">27.12</td>
</tr>
<tr id="3C9ED006FF9400306F822ECE6B07FE1D" box="[200,1346,462,489]" gridrow="3" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822ECE6C2EFE1D" box="[200,619,462,489]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">III</th>
<td id="7F4FB97AFF9400306DCD2ECE6CFFFE1D" box="[647,698,462,489]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">7.24</td>
<td id="7F4FB97AFF9400306C4C2ECE6D7CFE1D" box="[774,825,462,489]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">3.12</td>
<td id="7F4FB97AFF9400306CCE2ECE6DF2FE1D" box="[900,951,462,489]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">5.96</td>
<td id="7F4FB97AFF9400306B492ECE6A73FE1D" box="[1027,1078,462,489]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">7.67</td>
<td id="7F4FB97AFF9400306BCB2ECE6AF1FE1D" box="[1153,1204,462,489]" gridcol="5" gridrow="3" pageId="8" pageNumber="9">4.97</td>
<td id="7F4FB97AFF9400306A4A2ECE6B07FE1D" box="[1280,1346,462,489]" gridcol="6" gridrow="3" pageId="8" pageNumber="9">28.97</td>
</tr>
<tr id="3C9ED006FF9400306F822EF86B07FDE7" box="[200,1346,504,531]" gridrow="4" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822EF86C2EFDE7" box="[200,619,504,531]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">IV</th>
<td id="7F4FB97AFF9400306DCD2EF86CFFFDE7" box="[647,698,504,531]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">8.52</td>
<td id="7F4FB97AFF9400306C4C2EF86D7CFDE7" box="[774,825,504,531]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">3.12</td>
<td id="7F4FB97AFF9400306CCE2EF86DF2FDE7" box="[900,951,504,531]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">7.53</td>
<td id="7F4FB97AFF9400306B492EF86A73FDE7" box="[1027,1078,504,531]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">9.09</td>
<td id="7F4FB97AFF9400306BCB2EF86AF1FDE7" box="[1153,1204,504,531]" gridcol="5" gridrow="4" pageId="8" pageNumber="9">5.25</td>
<td id="7F4FB97AFF9400306A4A2EF86B07FDE7" box="[1280,1346,504,531]" gridcol="6" gridrow="4" pageId="8" pageNumber="9">33.51</td>
</tr>
<tr id="3C9ED006FF9400306F822D216B07FDC8" box="[200,1346,545,572]" gridrow="5" pageId="8" pageNumber="9" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="7F4FB97AFF9400306F822D216C2EFDC8" box="[200,619,545,572]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76F822D226FDFFDC8" bold="true" box="[200,410,546,572]" pageId="8" pageNumber="9">Female paratype</emphasis>
(MRAC 245338)
</th>
</tr>
<tr id="3C9ED006FF9400306F822D4B6B07FD92" box="[200,1346,587,614]" gridrow="6" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822D4B6C2EFD92" box="[200,619,587,614]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">I</th>
<td id="7F4FB97AFF9400306DCD2D4B6CFFFD92" box="[647,698,587,614]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">7.24</td>
<td id="7F4FB97AFF9400306C4C2D4B6D7CFD92" box="[774,825,587,614]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">4.12</td>
<td id="7F4FB97AFF9400306CCE2D4B6DF2FD92" box="[900,951,587,614]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">5.40</td>
<td id="7F4FB97AFF9400306B492D4B6A73FD92" box="[1027,1078,587,614]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">5.82</td>
<td id="7F4FB97AFF9400306BCB2D4B6AF1FD92" box="[1153,1204,587,614]" gridcol="5" gridrow="6" pageId="8" pageNumber="9">3.98</td>
<td id="7F4FB97AFF9400306A4A2D4B6B07FD92" box="[1280,1346,587,614]" gridcol="6" gridrow="6" pageId="8" pageNumber="9">26.55</td>
</tr>
<tr id="3C9ED006FF9400306F822D756B07FD64" box="[200,1346,629,656]" gridrow="7" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822D756C2EFD64" box="[200,619,629,656]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">II</th>
<td id="7F4FB97AFF9400306DCD2D756CFFFD64" box="[647,698,629,656]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">6.67</td>
<td id="7F4FB97AFF9400306C4C2D756D7CFD64" box="[774,825,629,656]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">3.41</td>
<td id="7F4FB97AFF9400306CCE2D756DF2FD64" box="[900,951,629,656]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">4.97</td>
<td id="7F4FB97AFF9400306B492D756A73FD64" box="[1027,1078,629,656]" gridcol="4" gridrow="7" pageId="8" pageNumber="9">5.40</td>
<td id="7F4FB97AFF9400306BCB2D756AF1FD64" box="[1153,1204,629,656]" gridcol="5" gridrow="7" pageId="8" pageNumber="9">3.69</td>
<td id="7F4FB97AFF9400306A4A2D756B07FD64" box="[1280,1346,629,656]" gridcol="6" gridrow="7" pageId="8" pageNumber="9">24.14</td>
</tr>
<tr id="3C9ED006FF9400306F822D9E6B07FD4D" box="[200,1346,670,697]" gridrow="8" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822D9E6C2EFD4D" box="[200,619,670,697]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">III</th>
<td id="7F4FB97AFF9400306DCD2D9E6CFFFD4D" box="[647,698,670,697]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">7.81</td>
<td id="7F4FB97AFF9400306C4C2D9E6D7CFD4D" box="[774,825,670,697]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">3.83</td>
<td id="7F4FB97AFF9400306CCE2D9E6DF2FD4D" box="[900,951,670,697]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">5.40</td>
<td id="7F4FB97AFF9400306B492D9E6A73FD4D" box="[1027,1078,670,697]" gridcol="4" gridrow="8" pageId="8" pageNumber="9">6.53</td>
<td id="7F4FB97AFF9400306BCB2D9E6AF1FD4D" box="[1153,1204,670,697]" gridcol="5" gridrow="8" pageId="8" pageNumber="9">4.69</td>
<td id="7F4FB97AFF9400306A4A2D9E6B07FD4D" box="[1280,1346,670,697]" gridcol="6" gridrow="8" pageId="8" pageNumber="9">28.26</td>
</tr>
<tr id="3C9ED006FF9400306F822DC86B07FD17" box="[200,1346,712,739]" gridrow="9" pageId="8" pageNumber="9">
<th id="7F4FB97AFF9400306F822DC86C2EFD17" box="[200,619,712,739]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">IV</th>
<td id="7F4FB97AFF9400306DCD2DC86CFFFD17" box="[647,698,712,739]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">7.81</td>
<td id="7F4FB97AFF9400306C4C2DC86D7CFD17" box="[774,825,712,739]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">4.54</td>
<td id="7F4FB97AFF9400306CCE2DC86DF2FD17" box="[900,951,712,739]" gridcol="3" gridrow="9" pageId="8" pageNumber="9">6.67</td>
<td id="7F4FB97AFF9400306B492DC86A73FD17" box="[1027,1078,712,739]" gridcol="4" gridrow="9" pageId="8" pageNumber="9">7.67</td>
<td id="7F4FB97AFF9400306BCB2DC86AF1FD17" box="[1153,1204,712,739]" gridcol="5" gridrow="9" pageId="8" pageNumber="9">5.68</td>
<td id="7F4FB97AFF9400306A4A2DC86B07FD17" box="[1280,1346,712,739]" gridcol="6" gridrow="9" pageId="8" pageNumber="9">32.38</td>
</tr>
</table>
</paragraph>
<caption id="D6D182CCFF94FFC76FF728676FF0F81F" ID-DOI="http://doi.org/10.5281/zenodo.3836323" ID-Zenodo-Dep="3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1895,1921]" targetBox="[192,1398,786,1853]" targetPageId="8">
<paragraph id="8211D244FF94FFC76FF728676FF0F81F" blockId="8.[189,1399,1895,2027]" pageId="8" pageNumber="9">
<emphasis id="B0DA0E56FF94FFC76FF728676F4FF875" bold="true" box="[189,266,1895,1921]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
<taxonomicName id="45AEA9C7FF94FFC76E5A28676C84F876" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[272,705,1895,1922]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF94FFC76E5A28676FEFF875" box="[272,426,1895,1921]" italics="true" pageId="8" pageNumber="9">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF94FFC76D8D28686D64F876" box="[711,801,1896,1922]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B0DA0E56FF94FFC76C6228676D24F875" bold="true" box="[808,865,1895,1921]" pageId="8" pageNumber="9">A–D</emphasis>
. Palp, ♂, holotype.
<emphasis id="B0DA0E56FF94FFC76B0328676A1BF875" bold="true" box="[1097,1118,1895,1921]" pageId="8" pageNumber="9">A</emphasis>
. Ventral view.
<emphasis id="B0DA0E56FF94FFC76A4628676B65F875" bold="true" box="[1292,1312,1895,1921]" pageId="8" pageNumber="9">B</emphasis>
. Same, detail.
<emphasis id="B0DA0E56FF94FFC76E41288A6F65F850" bold="true" box="[267,288,1930,1956]" pageId="8" pageNumber="9">C</emphasis>
. Lateral view.
<emphasis id="B0DA0E56FF94FFC76E81288A6FA5F850" bold="true" box="[459,480,1930,1956]" pageId="8" pageNumber="9">D</emphasis>
. Same, detail.
<emphasis id="B0DA0E56FF94FFC76DC0288A6CFBF850" bold="true" box="[650,702,1930,1956]" pageId="8" pageNumber="9">E–F</emphasis>
. Epigyne, ♀, paratype (MRAC 245338).
<emphasis id="B0DA0E56FF94FFC76BD4288A6AF7F850" bold="true" box="[1182,1202,1930,1956]" pageId="8" pageNumber="9">E</emphasis>
. Ventral view.
<emphasis id="B0DA0E56FF94FFC76A17288A6B2AF850" bold="true" box="[1373,1391,1930,1956]" pageId="8" pageNumber="9">F</emphasis>
. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8211D244FF95FFC66FA02E0C6CC6FED2" blockId="9.[189,1398,267,368]" box="[234,643,267,294]" pageId="9" pageNumber="10">
<emphasis id="B0DA0E56FF95FFC66FA02E0C6F00FED2" bold="true" box="[234,325,268,294]" pageId="9" pageNumber="10">Female</emphasis>
(
<typeStatus id="5D156CE6FF95FFC66E1F2E0B6FF9FED1" box="[341,444,267,293]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
MRAC 245338)
</paragraph>
<paragraph id="8211D244FF95FFC66FF72E316D30FE9B" blockId="9.[189,1398,267,368]" pageId="9" pageNumber="10">
MEASUREMENTS. TL 16.24; carapace 7.95 long, 3.69 high, 6.31 wide, narrowed to
<quantity id="45567FA1FF95FFC66BD92E306AA2FEBE" box="[1171,1255,304,330]" metricMagnitude="-2" metricUnit="m" metricValue="9.3726" pageId="9" pageNumber="10" unit="in" value="3.69">3.69 in</quantity>
eye region; labium 0.92 long, 1.08 wide; sternum 3.33 long, 2.83 wide.
</paragraph>
<paragraph id="8211D244FF95FFC66FF72E996B10FE2C" blockId="9.[189,1398,409,472]" pageId="9" pageNumber="10">
COLOUR. In ethanol (
<figureCitation id="1A95CEC1FF95FFC66EE72E996FB6FE47" box="[429,499,409,435]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[194,1393,262,1925]" captionTargetId="figure@7.[191,1396,259,1928]" captionTargetPageId="7" captionText="Fig. 4. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–B. Holotype, ♂. A. Dorsal habitus. B. Ventral habitus. C–D. Paratype, ♀ (MRAC 245338). C. Dorsal habitus. D. Ventral habitus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836321" httpUri="https://zenodo.org/record/3836321/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
C–D): dorsal surface of abdomen more uniform cream than in male but with similar dark spots and apodemes; pedipalp as in the male except for unmodified tarsus with dark tip.
</paragraph>
<paragraph id="8211D244FF95FFC66FF72D026B06FDEF" blockId="9.[189,1347,513,540]" box="[189,1347,513,540]" pageId="9" pageNumber="10">EYES. AME: 0.42; ALE: 0.21; PME: 0.67: PLE: 0.60; eye rows: ALE: 1.44, PME: 1.44, PLE: 2.14.</paragraph>
<paragraph id="8211D244FF95FFC66FF72D456B33FD70" blockId="9.[189,1399,580,644]" pageId="9" pageNumber="10">
LEGS. Spination of leg I, right (left): femur pl2(5), d3(4), rl3(3); patella pl1(1), rl1(1); tibia pl2(2), d1(1), rl2(2), v2-2-2 (3-2-2); metatarsus pl2(2), rl2(2), v2-2-2(2-2-2), dw5(5). Leg measurements: see
<tableCitation id="CF2CE7FFFF95FFC66A532D696B33FD70" box="[1305,1398,617,644]" captionStart="Table 1" captionStartId="8.[189,258,268,294]" captionTargetPageId="8" captionText="Table 1. Leg measurements of male holotype and female paratype (MRAC 245338)." httpUri="http://table.plazi.org/id/D6D182CCFF94FFC76FF72E0C6ADAFED2" pageId="9" pageNumber="10" tableUuid="D6D182CCFF94FFC76FF72E0C6ADAFED2">Table 1.</tableCitation>
</paragraph>
<paragraph id="8211D244FF95FFC66FF72DAD6C88FCE5" blockId="9.[189,1399,685,785]" pageId="9" pageNumber="10">
EPIGYNE (
<figureCitation id="1A95CEC1FF95FFC66E672DAD6F3CFD3C" box="[301,377,685,712]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[192,1398,786,1853]" captionTargetId="figure@8.[186,1401,785,1858]" captionTargetPageId="8" captionText="Fig. 5. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–D. Palp, ♂, holotype. A. Ventral view. B. Same, detail. C. Lateral view. D. Same, detail. E–F. Epigyne, ♀, paratype (MRAC 245338). E. Ventral view. F. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="9" pageNumber="10">Figs 5</figureCitation>
E–F, 7C). Roughly triangular area surrounded by dense mat of white setae; large and broad inverted T-shaped sclerite, 1.6 times wider than long; spermathecae large, globular; entrance ducts Z-shaped with basal portion slightly sinuous.
</paragraph>
<paragraph id="8211D244FF95FFC66FF72C3A6F76FCA0" blockId="9.[189,307,826,852]" box="[189,307,826,852]" pageId="9" pageNumber="10">
<heading id="D9596528FF95FFC66FF72C3A6F76FCA0" bold="true" box="[189,307,826,852]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B0DA0E56FF95FFC66FF72C3A6F76FCA0" bold="true" box="[189,307,826,852]" pageId="9" pageNumber="10">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF95FFC66FF72C6B6D8EFC71" blockId="9.[189,971,874,901]" box="[189,971,874,901]" pageId="9" pageNumber="10">Males: TL 16.76–19.45 (n = 5); females: TL 16.47–22.01 (n = 19).</paragraph>
</subSubSection>
<caption id="D6D182CCFF95FFC66FF628676D71F81F" ID-DOI="http://doi.org/10.5281/zenodo.3836325" ID-Zenodo-Dep="3836325" httpUri="https://zenodo.org/record/3836325/files/figure.png" pageId="9" pageNumber="10" startId="9.[188,231,1895,1921]" targetBox="[188,1398,946,1854]" targetPageId="9">
<paragraph id="8211D244FF95FFC66FF628676D71F81F" blockId="9.[188,1398,1895,2027]" pageId="9" pageNumber="10">
<emphasis id="B0DA0E56FF95FFC66FF628676F49F875" bold="true" box="[188,268,1895,1921]" pageId="9" pageNumber="10">Fig. 6.</emphasis>
<taxonomicName id="45AEA9C7FF95FFC66E5D28676C99F876" authority="Jocque M. &amp; Jocque R." authorityName="Jocque M. &amp; Jocque R." authorityYear="2017" box="[279,732,1895,1922]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF95FFC66E5D28676FF1F875" box="[279,436,1895,1921]" italics="true" pageId="9" pageNumber="10">Ocyale ghost</emphasis>
Jocque M. &amp; Jocqué R.
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF95FFC66DAC28686D03F876" box="[742,838,1896,1922]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, palp, ♂, paratype (MRAC 245361), scanning electron micrographs.
<emphasis id="B0DA0E56FF95FFC66E8C288A6F9EF850" bold="true" box="[454,475,1930,1956]" pageId="9" pageNumber="10">A</emphasis>
. Right palp, ventral view.
<emphasis id="B0DA0E56FF95FFC66C52288A6D69F850" bold="true" box="[792,812,1930,1956]" pageId="9" pageNumber="10">B</emphasis>
. Distal end of bulbus, ventral view.
<emphasis id="B0DA0E56FF95FFC66B96288A6AB4F850" bold="true" box="[1244,1265,1930,1956]" pageId="9" pageNumber="10">C</emphasis>
. Main part of bulbus ventro-prolateral view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; * = embolus. Scale bars: A = 0.5 mm; B–C = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="CAB481CFFF96FFC56FF72E0B6CFFFEA0" box="[189,698,267,340]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8211D244FF96FFC56FF72E0B6F13FED1" blockId="10.[189,342,267,293]" box="[189,342,267,293]" pageId="10" pageNumber="11">
<heading id="D9596528FF96FFC56FF72E0B6F13FED1" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B0DA0E56FF96FFC56FF72E0B6F13FED1" bold="true" box="[189,342,267,293]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF96FFC56FF72E3A6CFFFEA0" blockId="10.[189,698,313,340]" box="[189,698,313,340]" pageId="10" pageNumber="11">
Known only from the
<typeStatus id="5D156CE6FF96FFC56E882E3A6FB0FEA0" box="[450,501,314,340]" pageId="10" pageNumber="11">type</typeStatus>
locality (
<figureCitation id="1A95CEC1FF96FFC56D2E2E396CEDFEA0" box="[612,680,313,340]" captionStart="Fig" captionStartId="1.[189,232,2000,2026]" captionTargetBox="[189,1398,1008,1961]" captionTargetId="figure@1.[187,1399,1006,1962]" captionTargetPageId="1" captionText="Fig. 1. Type locality of Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3836315" httpUri="https://zenodo.org/record/3836315/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF96FFC56FF72E7B6FADFCF5" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8211D244FF96FFC56FF72E7B6F6AFE61" blockId="10.[189,303,379,405]" box="[189,303,379,405]" pageId="10" pageNumber="11">
<heading id="D9596528FF96FFC56FF72E7B6F6AFE61" bold="true" box="[189,303,379,405]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B0DA0E56FF96FFC56FF72E7B6F6AFE61" bold="true" box="[189,303,379,405]" pageId="10" pageNumber="11">Affinities</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF96FFC56FF72EAA6FADFCF5" blockId="10.[189,1399,425,769]" pageId="10" pageNumber="11">
We placed this species in the genus
<taxonomicName id="45AEA9C7FF96FFC56D3A2EA96C86FE37" authorityName="Audouin" authorityYear="1823" box="[624,707,425,451]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B0DA0E56FF96FFC56D3A2EA96C86FE37" box="[624,707,425,451]" italics="true" pageId="10" pageNumber="11">Ocyale</emphasis>
</taxonomicName>
based on the presence of the two elongate curved prongs on the palea (
<figureCitation id="1A95CEC1FF96FFC56E2A2ECC6F87FE13" box="[352,450,460,487]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[194,1393,262,1925]" captionTargetId="figure@7.[191,1396,259,1928]" captionTargetPageId="7" captionText="Fig. 4. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–B. Holotype, ♂. A. Dorsal habitus. B. Ventral habitus. C–D. Paratype, ♀ (MRAC 245338). C. Dorsal habitus. D. Ventral habitus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836321" httpUri="https://zenodo.org/record/3836321/files/figure.png" pageId="10" pageNumber="11">Figs 4B</figureCitation>
,
<figureCitation id="1A95CEC1FF96FFC56E9A2ECC6F98FE13" box="[464,477,460,487]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[192,1398,786,1853]" captionTargetId="figure@8.[186,1401,785,1858]" captionTargetPageId="8" captionText="Fig. 5. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–D. Palp, ♂, holotype. A. Ventral view. B. Same, detail. C. Lateral view. D. Same, detail. E–F. Epigyne, ♀, paratype (MRAC 245338). E. Ventral view. F. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="10" pageNumber="11">5</figureCitation>
B–C, 6B), the epigyne surrounded with white hairs and with a wide inverted T-shaped sclerite (
<figureCitation id="1A95CEC1FF96FFC56ED12EF06FBBFDFF" box="[411,510,496,523]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[192,1398,786,1853]" captionTargetId="figure@8.[186,1401,785,1858]" captionTargetPageId="8" captionText="Fig. 5. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–D. Palp, ♂, holotype. A. Ventral view. B. Same, detail. C. Lateral view. D. Same, detail. E–F. Epigyne, ♀, paratype (MRAC 245338). E. Ventral view. F. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="10" pageNumber="11">Figs 5E</figureCitation>
,
<figureCitation id="1A95CEC1FF96FFC56D472EF06C76FDFE" box="[525,563,496,522]" captionStart="Fig" captionStartId="10.[189,232,1930,1956]" captionTargetBox="[197,1391,1038,1875]" captionTargetId="figure@10.[186,1402,961,1893]" captionTargetPageId="10" captionText="Fig. 7. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–B. Palp (holotype, ♂). A. Retrolateral view. B. Ventral view. C. Epigyne (paratype, ♀, MRAC 245338), ventral view. Scale bar = 1mm.Abbreviations: MA = median apophysis; P = palea; T = tegulum; * = embolus." figureDoi="http://doi.org/10.5281/zenodo.3836327" httpUri="https://zenodo.org/record/3836327/files/figure.png" pageId="10" pageNumber="11">7C</figureCitation>
), the large globular spermathecae (
<figureCitation id="1A95CEC1FF96FFC56CAB2EF06A79FDFF" box="[993,1084,496,523]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[192,1398,786,1853]" captionTargetId="figure@8.[186,1401,785,1858]" captionTargetPageId="8" captionText="Fig. 5. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. A–D. Palp, ♂, holotype. A. Ventral view. B. Same, detail. C. Lateral view. D. Same, detail. E–F. Epigyne, ♀, paratype (MRAC 245338). E. Ventral view. F. Digested, dorsal view. Abbreviations: MA = median apophysis; P = palea; T = tegulum; E, * = embolus. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3836323" httpUri="https://zenodo.org/record/3836323/files/figure.png" pageId="10" pageNumber="11">Fig. 5F</figureCitation>
) and the conformation of the copulatory ducts (
<bibRefCitation id="E63FAFB5FF96FFC56E822D136CF4FDD9" author="Alderweireldt M." box="[456,689,531,557]" pageId="10" pageNumber="11" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt 1996</bibRefCitation>
: fig. 27). The specimen illustrated by
<bibRefCitation id="E63FAFB5FF96FFC56BC02D136B2BFDDA" author="Siyam M. &amp; Dunlop J. A. &amp; El-Hennawy H. K." box="[1162,1390,531,558]" pageId="10" pageNumber="11" pagination="264 - 272" refId="ref4834" refString="Siyam M., Dunlop J. A. &amp; El-Hennawy H. K. 2015. New spider records from the Republic of the Sudan. Arachnology 16 (7): 264 - 272. https: // doi. org / 10.13156 / arac. 2015.16.7.264" type="journal article" year="2015">
Siyam
<emphasis id="B0DA0E56FF96FFC56BAA2D146B5AFDD9" box="[1248,1311,531,557]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
(2015
</bibRefCitation>
: figs 13–15) from
<collectingCountry id="FAB992D4FF96FFC56EC42D366F92FDA4" box="[398,471,566,592]" name="Sudan" pageId="10" pageNumber="11">Sudan</collectingCountry>
is probably not an
<taxonomicName id="45AEA9C7FF96FFC56D8A2D366D56FDA4" authorityName="Audouin" authorityYear="1823" box="[704,787,566,592]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B0DA0E56FF96FFC56D8A2D366D56FDA4" box="[704,787,566,592]" italics="true" pageId="10" pageNumber="11">Ocyale</emphasis>
</taxonomicName>
, at least not
<taxonomicName id="45AEA9C7FF96FFC56CE42D376A5EFDA4" authorityName="Roewer" authorityYear="1960" box="[942,1051,566,593]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B0DA0E56FF96FFC56CE42D376A5EFDA4" box="[942,1051,566,593]" italics="true" pageId="10" pageNumber="11">O. pilosa</emphasis>
</taxonomicName>
because the palp does not fit the illustrations of
<bibRefCitation id="E63FAFB5FF96FFC56ED32D596CC2FD87" author="Alderweireldt M." box="[409,647,601,627]" pageId="10" pageNumber="11" pagination="103 - 111" refId="ref4420" refString="Alderweireldt M. 1996. A taxonomic revision of the genus Ocyale Audouin, 1826 in Africa (Araneae: Lycosidae). Journal of Natural History 30: 1349 - 1365. https: // doi. org / 10.1080 / 00222939600771261 Alderweireldt M. &amp; Jocque R. 1991. A remarkable new genus of wolf spiders from southwestern Spain (Araneae, Lycosidae). Bulletin van het Koninklijk Belgisch Instituut voor Natuurwetenschappen, Entomologie 61: 103 - 111." type="journal article" year="1996">Alderweireldt (1996</bibRefCitation>
: figs 12–13). The colour pattern described for the genus in both these papers may be inaccurate based on the observed differences between specimens in ethanol and photos of the living spiders for this new species. This indicates the value of including images of living animals in descriptions of new species. It is not clear to which species the representative of
<collectingCountry id="FAB992D4FF96FFC56BA32DC36B32FD29" box="[1257,1399,707,733]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
is most closely related to.
</paragraph>
</subSubSection>
<subSubSection id="CAB481CFFF96FFC46FF72C276D03FD97" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="8211D244FF96FFC56FF72C276F59FCB5" blockId="10.[189,284,807,833]" box="[189,284,807,833]" pageId="10" pageNumber="11">
<heading id="D9596528FF96FFC56FF72C276F59FCB5" bold="true" box="[189,284,807,833]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B0DA0E56FF96FFC56FF72C276F59FCB5" bold="true" box="[189,284,807,833]" pageId="10" pageNumber="11">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8211D244FF96FFC46FF72C566D03FD97" blockId="10.[189,1399,854,916]" lastBlockId="11.[189,1399,267,612]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<taxonomicName id="45AEA9C7FF96FFC56FF72C566F1DFC84" authority="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué, 2017" authorityName="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué" authorityYear="2017" box="[189,344,854,880]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF96FFC56FF72C566F1DFC84" box="[189,344,854,880]" italics="true" pageId="10" pageNumber="11">Ocyale ghost</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF96FFC56E2A2C576FFEFC85" box="[352,443,855,881]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
was only found on the white sandy beaches (
<figureCitation id="1A95CEC1FF96FFC56C9C2C566A5FFC84" box="[982,1050,854,880]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
A–B) of an inland lake in the study region. The surveys also included grassland and dry forest, but the species seems restricted to a white-sand habitat, as reflected in its habitus.
<taxonomicName id="45AEA9C7FF97FFC46D9D2E0B6D37FED1" authority="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué, 2017" authorityName="Jocque &amp; Wellens &amp; Andrianarivosoa &amp; Rakotondraparany &amp; Seing &amp; Jocqué" authorityYear="2017" box="[727,882,267,293]" class="Arachnida" family="Lycosidae" genus="Ocyale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ghost" status="sp. nov.">
<emphasis id="B0DA0E56FF97FFC46D9D2E0B6D37FED1" box="[727,882,267,293]" italics="true" pageId="11" pageNumber="12">Ocyale ghost</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABE9B32DFF97FFC46C332E0C6D91FED2" box="[889,980,268,294]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
is active at night and all specimens were caught with headlamps after sunset. Captured animals that were kept alive in large ziplock bags overnight constructed retreats in the sand, lined with silk (
<figureCitation id="1A95CEC1FF97FFC46C272E526D8FFE98" box="[877,970,338,364]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="11" pageNumber="12">Fig. 3C</figureCitation>
). Possible prey include large insects such as grasshoppers (
<figureCitation id="1A95CEC1FF97FFC46E8E2E756C5FFE7B" box="[452,538,373,399]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="11" pageNumber="12">Fig. 3F</figureCitation>
) that also exhibit camouflage colours as an adaptation to the white beach they live on. Intraspecific predation is also likely to occur (
<figureCitation id="1A95CEC1FF97FFC46C2D2E986D85FE47" box="[871,960,408,435]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="11" pageNumber="12">Fig. 3E</figureCitation>
), a phenomenon which is not unusual among lycosids (
<bibRefCitation id="E63FAFB5FF97FFC46EC02EBC6C51FE22" author="Edgar W. D." box="[394,532,444,470]" pageId="11" pageNumber="12" pagination="487 - 491" refId="ref4579" refString="Edgar W. D. 1969. Prey and feeding behaviour of adult females of the wolf spider Pardosa amentata (Clerck). Netherlands Journal of Zoology 20: 487 - 491. https: // doi. org / 10.1163 / 002829670 X 00259" type="journal article" year="1969">Edgar 1969</bibRefCitation>
;
<bibRefCitation id="E63FAFB5FF97FFC46D6F2EBC6C9BFE22" author="Hallander H." box="[549,734,444,470]" pageId="11" pageNumber="12" pagination="337 - 340" refId="ref4667" refString="Hallander H. 1970. Prey, cannibalism and microhabitat selection in the wolf spiders Pardosa chelata O. F. Muller and P. pullata Clerck. Oikos 21: 337 - 340. https: // doi. org / 10.2307 / 3543691" type="journal article" year="1970">Hallander 1970</bibRefCitation>
). We observed copulation and females with spiderlings (
<figureCitation id="1A95CEC1FF97FFC46F8C2EDF6F67FE0D" box="[198,290,479,505]" captionStart="Fig" captionStartId="6.[189,232,1651,1677]" captionTargetBox="[189,1396,265,1611]" captionTargetId="figure@6.[186,1399,262,1614]" captionTargetPageId="6" captionText="Fig. 3. Ocyale ghost Jocque M. &amp; Jocqué R. sp. nov. photographed at type locality. A. Female habitus. B. Same, detail. C. Female in sand retreat. D. Female with spiderlings on abdomen. E. Two males, one being eaten by the other. F. Female with white grasshopper prey. Photos A–B: MJ (2012), C–F: SW (2016)." figureDoi="http://doi.org/10.5281/zenodo.3836319" httpUri="https://zenodo.org/record/3836319/files/figure.png" pageId="11" pageNumber="12">Fig. 3D</figureCitation>
) in the midst of the dry season (June–July). Juveniles of a complete range of size, from very small ones (
<quantity id="45567FA1FF97FFC46E072D026FD6FDE9" box="[333,403,514,541]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="mm" value="6.0">6 mm</quantity>
TL) to subadults, were observed, indicating that this species might reproduce yearround. The permanent presence of water in its habitat might explain why this species is also active in the dry season when spider activity is on average very low.
</paragraph>
</subSubSection>
</treatment>
</document>
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