187 lines
24 KiB
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187 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1160.98022" ID-Pensoft-Pub="1313-2970-1160-1" ID-Pensoft-UUID="B0708A590ABD58FEB4A1A474CA268AA4" ID-ZooBank="B7E449240D3D4529987F19A51FAA115E" ModsDocID="1313-2970-1160-1" checkinTime="1683131562219" checkinUser="pensoft" docAuthor="Man, Nem Sian, Lwin, Ngwe, Sutcharit, Chirasak & Panha, Somsak" docDate="2023" docId="39C456FE0B055B3FBE229C90A1930B69" docLanguage="en" docName="ZooKeys 1160: 1-59" docOrigin="ZooKeys 1160" docPubDate="2023-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1160.98022" docTitle="Phaedusa burmanica" docType="treatment" docVersion="1" id="B0708A590ABD58FEB4A1A474CA268AA4" lastPageNumber="1" masterDocId="B0708A590ABD58FEB4A1A474CA268AA4" masterDocTitle="Further notes on the taxonomy of the land snail family Clausiliidae Gray, 1855 (Stylommatophora, Helicina) from Myanmar with description of two new species" masterLastPageNumber="59" masterPageNumber="1" pageNumber="1" updateTime="1683131562219" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Further notes on the taxonomy of the land snail family Clausiliidae Gray, 1855 (Stylommatophora, Helicina) from Myanmar with description of two new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Man, Nem Sian</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lwin, Ngwe</mods:namePart>
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<mods:affiliation>Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-03</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1160</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>59</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1160.98022</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1160.98022</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1160-1</mods:identifier>
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<mods:identifier type="ZooBank">B7E449240D3D4529987F19A51FAA115E</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B0708A590ABD58FEB4A1A474CA268AA4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:39C456FE0B055B3FBE229C90A1930B69" httpUri="http://treatment.plazi.org/id/39C456FE0B055B3FBE229C90A1930B69" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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2
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<taxonomicName ID-CoL="7SNYF" LSID="39C456FE-0B05-5B3F-BE22-9C90A1930B69" authority="(Gude, 1914)" baseAuthorityName="Gude" baseAuthorityYear="1914" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa burmanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burmanica">Phaedusa burmanica (Gude, 1914)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell and clausilial apparatus of A, B Phaedusa shanica, specimen CUMZ 13032 from Pyinyaung Village, Mandalay Region A shell B clausilial apparatus C, D Phaedusa burmanica, specimen CUMZ 13034 from Myin Ma Hti Cave, Shan State C shell D clausilial apparatus and E, F Phaedusa bocki menglunanensis, specimen CUMZ 13036 from Aik Kham Cave, Shan State E shell F clausilial apparatus. White arrows indicate principalis and yellow arrows indicate clausilium hook." figureDoi="10.3897/zookeys.1160.98022.figure4" httpUri="https://binary.pensoft.net/fig/845466" pageId="0" pageNumber="1">Figs 4C, D</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genital anatomy of A-D Phaedusa shanica, specimen CUMZ 13033 A whole reproductive system B internal sculpture of penis C transition from penis to epiphallus D internal sculpture of epiphallus and E-G Phaedusa burmanica, specimen CUMZ 13035 E whole reproductive system F overview internal sculpture of penis and epiphallus G internal sculpture of proximal epiphallus. Yellow arrows indicate approximate transitional position from penis to epiphallus." figureDoi="10.3897/zookeys.1160.98022.figure5" httpUri="https://binary.pensoft.net/fig/845467" pageId="0" pageNumber="1">, 5E-G</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Radula morphology A Phaedusa shanica, specimen CUMZ 13033 and B Phaedusa burmanica, specimen CUMZ 13035. Blue color indicates the central teeth row; yellow color indicates the transition from lateral to marginal teeth. ' C' indicates central tooth." figureDoi="10.3897/zookeys.1160.98022.figure6" httpUri="https://binary.pensoft.net/fig/845468" pageId="0" pageNumber="1">, 6B</figureCitation>
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<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Synoptic view of the clausiliid species recorded from Myanmar A Indonenia admirabilis, holotype NHMUK 20200185 B Oospira andersoniana, specimen CUMZ 13054 C Oospira arakana, lectotype NHMUK 1888.12.4.1037 D Cylindrophaedusa bacillum, specimen NHMUK 1906.2.2.345 E Oospira bulbus, possible syntype NHMUK 1903.7.1.1278 F Phaedusa burmanica, holotype NHMUK 1888.12.4.1058 G Oospira decollata, paratype ZISP H Indonenia excellens, specimen NHMUK 20200189 I Oospira fusiformis, holotype NHMUK 1906.2.2.380 J Oospira gouldiana, lectotype NHMUK 196548 K Oospira gracilior, lectotype NHMUK 1907.12.30.243 L Oospira insignis, lectotype USNM 117120 M Phaedusa kazueae, holotype NHMUK 20200187 N Phaedusa lemani (after Gude, 1914: fig. 107) O Oospira limborgi, holotype NHMUK 1903.7.1.1266, and P Oospira luneainopsis sp. nov., holotype CUMZ 13062." figureDoi="10.3897/zookeys.1160.98022.figure18" httpUri="https://binary.pensoft.net/fig/845480" pageId="0" pageNumber="1">, 18F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Draparnaud" authorityYear="1805" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Clausilia</taxonomicName>
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[
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<taxonomicName authority="" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa (Euphaedusa)" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Euphaedusa">Phaedusa (Euphaedusa)</taxonomicName>
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]
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<taxonomicName baseAuthorityName="Gude" baseAuthorityYear="1914" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa burmanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burmanica">Phaedusa burmanica</taxonomicName>
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Gude, 1914: 311, fig. 106. Type locality: "Burma, Tonghu" [Taungoo District, Bago Region, Myanmar].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Gude" baseAuthorityYear="1914" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa (Phaedusa) burmanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burmanica" subGenus="Phaedusa">Phaedusa (Phaedusa) burmanica</taxonomicName>
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-
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Basteria" pageId="0" pageNumber="1" pagination="85 - 100" refId="B51" refString="Nordsieck, H, 2002b. Annotated checklist of the Southeast Asian Phaedusinae, with the description of new taxa (Gastropoda, Pulmonata, Clausiliidae). Basteria 66: 85 - 100" title="Annotated checklist of the Southeast Asian Phaedusinae, with the description of new taxa (Gastropoda, Pulmonata, Clausiliidae)." volume="66" year="2002 b">Nordsieck 2002b</bibRefCitation>
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: 88.
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="ConchBooks, Hackenheim" pageId="0" pageNumber="1" refId="B52" refString="Nordsieck, H, 2007. Worldwide Door Snails. ConchBooks, Hackenheim" title="Worldwide Door Snails." year="2007">Nordsieck 2007</bibRefCitation>
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: 32.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Gude" baseAuthorityYear="1914" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa burmanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="burmanica">Phaedusa burmanica</taxonomicName>
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-
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<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 49" refId="B49" refString="Nordsieck, H, 1974. Zur Anatomie und Systematik der Clausilien, 14. Phaedusinae, 2: Phaedusa bhutanensis n. sp. und ihre Beziehungen zu benachbarten Arten. Archiv fuer Molluskenkunde 104: 41 - 49" title="Zur Anatomie und Systematik der Clausilien, 14. Phaedusinae, 2: Phaedusa bhutanensis n. sp. und ihre Beziehungen zu benachbarten Arten." volume="104" year="1974">Nordsieck 1974</bibRefCitation>
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: 46, fig. 5, pl. 2, figs 3, 4.
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<bibRefCitation author="Grego, J" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="21 - 29" refId="B25" refString="Grego, J, Hunyadi, A, Szekeres, M, 2021. New and little-known species of Southeast Asian Clausiliidae (Gastropoda: Pulmonata). Journal of Conchology 44: 21 - 29" title="New and little-known species of Southeast Asian Clausiliidae (Gastropoda: Pulmonata)." volume="44" year="2021">Grego et al. 2021</bibRefCitation>
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: 27.
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<bibRefCitation author="Szekeres, M" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="155 - 187" refId="B82" refString="Szekeres, M, Grego, J, Pall-Gergely, B, Ablett, JD, 2021a. Clausiliidae (Gastropoda; Pulmonata) from India, Myanmar, Pakistan, and Sri Lanka in the collection of the Natural History Museum, London. Journal of Conchology 44: 155 - 187" title="Clausiliidae (Gastropoda; Pulmonata) from India, Myanmar, Pakistan, and Sri Lanka in the collection of the Natural History Museum, London." volume="44" year="2021 a">Szekeres et al. 2021a</bibRefCitation>
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: 181, fig. 10a.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Myanmar" latitude="20.590584" location="Myin Ma Hti Cave" longLatPrecision="1" longitude="96.61183" municipality="Kalaw City" specimenCount="1" stateProvince="Shan State">
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<location LSID="urn:lsid:plazi:treatment:39C456FE0B055B3FBE229C90A1930B69:C89B55C02E1DAFB14CE53F4FDDEDDA0F" country="Myanmar" latitude="20.590584" longLatPrecision="1" longitude="96.61183" municipality="Kalaw City" name="Myin Ma Hti Cave" stateProvince="Shan State">Myin Ma Hti Cave</location>
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,
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<collectingMunicipality>Kalaw City</collectingMunicipality>
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,
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
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,
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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(
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<geoCoordinate degrees="20" direction="north" minutes="35" orientation="latitude" precision="1" seconds="26.1" value="20.590584">20°35'26.1"N</geoCoordinate>
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,
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<geoCoordinate degrees="96" direction="east" minutes="36" orientation="longitude" precision="1" seconds="42.6" value="96.61183">96°36'42.6"E</geoCoordinate>
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): CUMZ 13034 (2 shells + 1 incomplete shell; Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell and clausilial apparatus of A, B Phaedusa shanica, specimen CUMZ 13032 from Pyinyaung Village, Mandalay Region A shell B clausilial apparatus C, D Phaedusa burmanica, specimen CUMZ 13034 from Myin Ma Hti Cave, Shan State C shell D clausilial apparatus and E, F Phaedusa bocki menglunanensis, specimen CUMZ 13036 from Aik Kham Cave, Shan State E shell F clausilial apparatus. White arrows indicate principalis and yellow arrows indicate clausilium hook." figureDoi="10.3897/zookeys.1160.98022.figure4" httpUri="https://binary.pensoft.net/fig/845466" pageId="0" pageNumber="1">4C, D</figureCitation>
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), CUMZ 13035 (
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell fusiform, turreted, translucent, and pale chestnut-brown; spire acute. Shell surface glossy with thin and dense striations; suture distinct. Whorls 12-13, little convex, regularly growing and attenuated to apex; last whorl compressed. Aperture vertically subquadrate, basis rounded and narrowing towards parietal sinus; peristome slightly protruded, thickened, and expanded. Superior lamella low, continuous with spiralis, and very low at transition to spiralis. Inferior lamella spirally ascending, close to superior lamella and ending at peristome. Subcolumellaris emerged, and visible in oblique view. Principalis running along lateral-dorsal side and anterior end visible through oblique apertural view. Palatal plicae lateral, distinctly five: first plica strong, longest, and nearly connected to principalis anteriorly; second and fifth plicae almost equal and slightly strong; third and fourth plicae weakest and almost not visible. Palatal plicae visible through translucent shell. Clausilium plate lateral side, broad, rounded and with hook.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
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(
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 1). Atrium (at) short and slender; penis (p) muscular, cylindrical, broadest at middle part and gradually narrower towards epiphallus. Epiphallus (ep) muscular, cylindrical, ca. same length and size as penis, gradually enlarging to proximal end. Penial retractor muscle (pr) attached proximally to epiphallus. Vas deferens (vd) slender, and shortly bounded at atrium and penis junction (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genital anatomy of A-D Phaedusa shanica, specimen CUMZ 13033 A whole reproductive system B internal sculpture of penis C transition from penis to epiphallus D internal sculpture of epiphallus and E-G Phaedusa burmanica, specimen CUMZ 13035 E whole reproductive system F overview internal sculpture of penis and epiphallus G internal sculpture of proximal epiphallus. Yellow arrows indicate approximate transitional position from penis to epiphallus." figureDoi="10.3897/zookeys.1160.98022.figure5" httpUri="https://binary.pensoft.net/fig/845467" pageId="0" pageNumber="1">5E</figureCitation>
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). Internal sculpture of penis smooth-surfaced and slightly wavy with ca. four longitudinal folds (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genital anatomy of A-D Phaedusa shanica, specimen CUMZ 13033 A whole reproductive system B internal sculpture of penis C transition from penis to epiphallus D internal sculpture of epiphallus and E-G Phaedusa burmanica, specimen CUMZ 13035 E whole reproductive system F overview internal sculpture of penis and epiphallus G internal sculpture of proximal epiphallus. Yellow arrows indicate approximate transitional position from penis to epiphallus." figureDoi="10.3897/zookeys.1160.98022.figure5" httpUri="https://binary.pensoft.net/fig/845467" pageId="0" pageNumber="1">5F</figureCitation>
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). Internal sculpture of epiphallus generally defined as two parts: region near penis with wavy longitudinal folds, then gradually becoming more densely crowded towards proximal epiphallus; region near vas deferens with strong folds and papillae arranged in oblique rows (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genital anatomy of A-D Phaedusa shanica, specimen CUMZ 13033 A whole reproductive system B internal sculpture of penis C transition from penis to epiphallus D internal sculpture of epiphallus and E-G Phaedusa burmanica, specimen CUMZ 13035 E whole reproductive system F overview internal sculpture of penis and epiphallus G internal sculpture of proximal epiphallus. Yellow arrows indicate approximate transitional position from penis to epiphallus." figureDoi="10.3897/zookeys.1160.98022.figure5" httpUri="https://binary.pensoft.net/fig/845467" pageId="0" pageNumber="1">5G</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Vagina (v) slender at junction of atrium, then bulging at connection of free oviduct (fo) and distal part of pedunculus (dpb), and almost as long as free oviduct. Distal part of pedunculus large, long, basally stouter, and broadest near entrance of diverticulum; diverticulum (db) large, as long as dpb; proximal part of pedunculus (ppb) large, measuring ~ 1/2 diverticulum length, and bursa copulatrix (bb) ovoid (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Genital anatomy of A-D Phaedusa shanica, specimen CUMZ 13033 A whole reproductive system B internal sculpture of penis C transition from penis to epiphallus D internal sculpture of epiphallus and E-G Phaedusa burmanica, specimen CUMZ 13035 E whole reproductive system F overview internal sculpture of penis and epiphallus G internal sculpture of proximal epiphallus. Yellow arrows indicate approximate transitional position from penis to epiphallus." figureDoi="10.3897/zookeys.1160.98022.figure5" httpUri="https://binary.pensoft.net/fig/845467" pageId="0" pageNumber="1">5E</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Radula</emphasis>
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.
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</emphasis>
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Each row contains ~ 51 teeth with half-row formula: central-(lateral)-marginal teeth (1-(13-14)-25). Central tooth small, symmetrically tricuspid; mesocone large with dull tip; ectocones very small. Lateral teeth asymmetrically bicuspid: endocone large and with dull tip; ectocone very small, pointed tip and located near the base. Marginal teeth tricuspid starting at approximately tooth number 13 or 14; mesocone long and dull tip; endocone very small and located near tip of mesocone; ectocone small with pointed tip. Marginal teeth gradually becoming polycuspid towards radula margin (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Radula morphology A Phaedusa shanica, specimen CUMZ 13033 and B Phaedusa burmanica, specimen CUMZ 13035. Blue color indicates the central teeth row; yellow color indicates the transition from lateral to marginal teeth. ' C' indicates central tooth." figureDoi="10.3897/zookeys.1160.98022.figure6" httpUri="https://binary.pensoft.net/fig/845468" pageId="0" pageNumber="1">6B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Apart from the type locality, this species is widely known from Bhamo, Kachin State (
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B26" refString="Gude, GK, 1914. The Fauna of British India, Including Ceylon and Burma. Mollusca, II. Taylor and Francis, London" title="The Fauna of British India, Including Ceylon and Burma. Mollusca, II." year="1914">Gude 1914</bibRefCitation>
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) to Shan State and the Mandalay Region in Myanmar (
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<bibRefCitation author="Grego, J" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="21 - 29" refId="B25" refString="Grego, J, Hunyadi, A, Szekeres, M, 2021. New and little-known species of Southeast Asian Clausiliidae (Gastropoda: Pulmonata). Journal of Conchology 44: 21 - 29" title="New and little-known species of Southeast Asian Clausiliidae (Gastropoda: Pulmonata)." volume="44" year="2021">Grego et al. 2021</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Luo, Chen & Zhang" baseAuthorityYear="1998" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa bocki subsp. menglunanensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="bocki" subSpecies="menglunanensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Phaedusa bocki menglunanensis</emphasis>
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</taxonomicName>
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from Shan State can be distinguished from this species by its inferior lamella less spirally ascending, palatal plicae longer, obliquely arranged and more spaced between them, shell surface finer and more curved striations, aperture vertically quadrate, and subcolumellaris distinctly visible in frontal view.
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<taxonomicName authorityName="Grego & Szekeres" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Phaedusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phaedusa bocki subsp. thompsoni" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="bocki" subSpecies="thompsoni">
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<emphasis italics="true" pageId="0" pageNumber="1">Phaedusa bocki thompsoni</emphasis>
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</taxonomicName>
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Grego & Szekeres, 2021 also differs from this species in having a more vertical aperture, acute spire, and palatal plicae more on dorsal side (see
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<bibRefCitation author="Szekeres, M" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="155 - 187" refId="B82" refString="Szekeres, M, Grego, J, Pall-Gergely, B, Ablett, JD, 2021a. Clausiliidae (Gastropoda; Pulmonata) from India, Myanmar, Pakistan, and Sri Lanka in the collection of the Natural History Museum, London. Journal of Conchology 44: 155 - 187" title="Clausiliidae (Gastropoda; Pulmonata) from India, Myanmar, Pakistan, and Sri Lanka in the collection of the Natural History Museum, London." volume="44" year="2021 a">Szekeres et al. 2021a</bibRefCitation>
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: fig. 9f).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Variation was observed among the examined specimens in having the superior lamella and spiralis either separated or connected.</paragraph>
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</subSubSection>
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</treatment>
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</document> |