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<mods:title id="5D374D927C8A8B64BF481A5FC227DB10">Revision of the genera Paramacroxiphus C. Willemse 1961 and Pseudomacroxiphus C. Willemse 1961 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini)</mods:title>
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<subSubSection id="C68B6511FFDAFFF419ECFF45FDDEACB2" box="[151,613,151,177]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8E2E369AFFDAFFF419ECFF45FDDEACB2" blockId="29.[151,613,151,177]" box="[151,613,151,177]" pageId="29" pageNumber="30">
<heading id="D56681F6FFDAFFF419ECFF45FDDEACB2" bold="true" box="[151,613,151,177]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<taxonomicName id="49914D19FFDAFFF419ECFF45FDDEACB2" ID-CoL="8NY74" authority="C. Willemse 1961" authorityName="C. Willemse" authorityYear="1961" box="[151,613,151,177]" class="Insecta" family="Tettigoniidae" genus="Pseudomacroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF419ECFF45FDDEACB2" bold="true" box="[151,613,151,177]" pageId="29" pageNumber="30">
<emphasis id="BCE5EA88FFDAFFF419ECFF45FE36ACB2" bold="true" box="[151,397,151,177]" italics="true" pageId="29" pageNumber="30">Pseudomacroxiphus</emphasis>
C.Willemse 1961
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C68B6511FFDAFFF419ECFF30FC77AD1A" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8E2E369AFFDAFFF419ECFF30FC77AD1A" blockId="29.[151,1268,226,281]" pageId="29" pageNumber="30">
<typeStatus id="512A8838FFDAFFF419ECFF30FF77ACFA" box="[151,204,226,249]" pageId="29" pageNumber="30">Type</typeStatus>
species:
<taxonomicName id="49914D19FFDAFFF41856FF30FCA9ACFA" authority="Willemse 1961" authorityName="C. Willemse" authorityYear="1961" box="[301,786,226,249]" class="Insecta" family="Tettigoniidae" genus="Pseudomacroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="atrifrons">
<emphasis id="BCE5EA88FFDAFFF41856FF30FDDCACFA" box="[301,615,226,249]" italics="true" pageId="29" pageNumber="30">Pseudomacroxiphus atrifrons</emphasis>
Willemse 1961
</taxonomicName>
, by original designation (= syn of.
<taxonomicName id="49914D19FFDAFFF41DFCFF30FB50ACFA" authorityName="Ingrisch" authorityYear="2008" baseAuthorityName="C. Willemse" baseAuthorityYear="1958" box="[1159,1259,226,249]" class="Insecta" family="Tettigoniidae" genus="Pseudomacroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="szentia">
<emphasis id="BCE5EA88FFDAFFF41DFCFF30FB50ACFA" box="[1159,1259,226,249]" italics="true" pageId="29" pageNumber="30">P. szentia</emphasis>
</taxonomicName>
)
<taxonomicName id="49914D19FFDAFFF419ECFED0FD86AD1A" authority="C. Willemse 1961 a" authorityName="C. Willemse" authorityYear="1961" box="[151,573,258,281]" class="Insecta" family="Tettigoniidae" genus="Pseudomacroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF419ECFED0FED5AD1A" box="[151,366,258,281]" italics="true" pageId="29" pageNumber="30">Pseudomacroxiphus</emphasis>
C.
<bibRefCitation id="EA004B6BFFDAFFF418F3FED0FD86AD1A" author="Willemse, C." box="[392,573,258,281]" pageId="29" pageNumber="30" pagination="1960 - 1961" refId="ref17659" refString="Willemse, C. (1961 a) Description of some new Orthoptera from the Indo-Malayan region (Tettigonioidea). Publicaties van het Natuurhistorisch Genootschap in Limburg 12 [1960 - 1961], 28 - 35; Maastricht." type="journal article" year="1961">Willemse 1961a</bibRefCitation>
</taxonomicName>
, Publ. natuurh. Gen.
<collectingRegion id="4C55F878FFDAFFF41A65FED0FCC0AD1A" box="[798,891,258,281]" country="Netherlands" name="Limburg" pageId="29" pageNumber="30">Limburg</collectingRegion>
12: 29.
</paragraph>
</subSubSection>
<subSubSection id="C68B6511FFDAFFF419ECFE98FDA7A8AF" pageId="29" pageNumber="30" type="description">
<paragraph id="8E2E369AFFDAFFF419ECFE98FE54AF1F" blockId="29.[151,1437,330,1556]" pageId="29" pageNumber="30">
<emphasis id="BCE5EA88FFDAFFF419ECFE98FE94AD67" bold="true" box="[151,303,330,356]" pageId="29" pageNumber="30">Description.</emphasis>
Fastigium verticis narrow, compressed, conical, shorter than scapus; dorsal surface with or without a faint swelling at base; ventral margin separated by a shallow but distinct sinuosity from fastigium frontis. Frons matt or shining, with some shallowly impressed dots; genae subsmooth (
<figureCitation id="16AA2A1FFFDAFFF41DD5FE48FAB6ADB7" box="[1198,1293,410,436]" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Fig. 82</figureCitation>
). Pronotum subsmooth, rather broad and low, disc broadly rounded into paranota, very apical area slightly raised, flat and shouldered; anterior margin broadly rounded but concave in middle; posterior margin truncate; transverse sulcus faint, broadly interrupted in middle; second transverse sulcus distinct on paranota, obsolete on disc (
<figureCitation id="16AA2A1FFFDAFFF41C17FDC0FF03AE57" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Fig. 80</figureCitation>
). Paranota with ventral margin almost straight, descending posteriorly (
<figureCitation id="16AA2A1FFFDAFFF41D64FDE8FBC1AE57" box="[1055,1146,570,596]" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Fig. 81</figureCitation>
). Fully winged; tegmen with apex rounded; hind wings of same length as tegmina (
<figureCitation id="16AA2A1FFFDAFFF41A0DFDB0FC69AE7F" box="[886,978,610,636]" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Fig. 81</figureCitation>
). Prosternal spines of medium length (shorter than coxa), but very stout. Mesosternal lobes angular or rounded, metasternal lobes rounded; medial plate at both posterior angles with an obtuse cone. Knee lobes of pro- and mesofemora spinose on both sides (spine on external side of profemur often short, mesofemur on external side sometimes bi-spinose); of postfemur bi-spinose on both sides.
</paragraph>
<paragraph id="8E2E369AFFDAFFF419BDFCF8FB79A837" blockId="29.[151,1437,330,1556]" pageId="29" pageNumber="30">
Male. Stridulatory area of tegmen with mirror almost quadrangular and behind mirror with a series of additional fields without nervature, separated from each other by subparallel transverse veins (
<figureCitation id="16AA2A1FFFDAFFF41D86FC80FA34AF6F" box="[1277,1423,850,876]" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Figs. 8384</figureCitation>
). Tenth abdominal tergite transverse but slightly descending posteriorly; apical margin roundly excised in middle and with a rounded projecting lobe at both sides of excision (
<figureCitation id="16AA2A1FFFDAFFF41AD2FC70FB87AFBF" box="[937,1084,930,956]" captionStart="FIGURES 8694" captionStartId="25.[151,269,1853,1877]" captionTargetBox="[159,1428,182,1825]" captionTargetId="figure-109@25.[151,1436,170,1830]" captionTargetPageId="25" captionText="FIGURES 8694. Pseudomacroxiphus szentia details: 8687, Titillators in ventro-distal view; 88, do. in latero-distal view; 89, do. in dorso-proximal view; 9091, male abdominal apex in dorsal view; 92 female subgenital plate in ventral view; 93 do. in lateral view; 94, male stridulatory file on underside of left tegmen.—8687, 91, 94, holotype male of P. szentia; 8889, holotype male of P. atrifrons; 90, paratype male of P. atrifrons; 9293, paratype female of P. atrifrons. Abbreviations: 10t tenth abdominal tergite; ce cercus; e epiproct; s stylus; sg subgenital plate. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5123664" httpUri="https://zenodo.org/record/5123664/files/figure.png" pageId="29" pageNumber="30">Figs. 9091</figureCitation>
). Epiproct rounded triangular with a faint medial sulcus in basal half. Projections of paraprocts rather long, stout, conical and curved mediad. Subgenital plate with apex roundly excised in middle, on both sides of excision with short and stout projections which carry the styli; styli as long as or little longer than excised area (
<figureCitation id="16AA2A1FFFDAFFF41D26FBC8FB0FA837" box="[1117,1204,1050,1076]" captionStart="FIGURES 8694" captionStartId="25.[151,269,1853,1877]" captionTargetBox="[159,1428,182,1825]" captionTargetId="figure-109@25.[151,1436,170,1830]" captionTargetPageId="25" captionText="FIGURES 8694. Pseudomacroxiphus szentia details: 8687, Titillators in ventro-distal view; 88, do. in latero-distal view; 89, do. in dorso-proximal view; 9091, male abdominal apex in dorsal view; 92 female subgenital plate in ventral view; 93 do. in lateral view; 94, male stridulatory file on underside of left tegmen.—8687, 91, 94, holotype male of P. szentia; 8889, holotype male of P. atrifrons; 90, paratype male of P. atrifrons; 9293, paratype female of P. atrifrons. Abbreviations: 10t tenth abdominal tergite; ce cercus; e epiproct; s stylus; sg subgenital plate. Scales = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5123664" httpUri="https://zenodo.org/record/5123664/files/figure.png" pageId="29" pageNumber="30">Fig. 91</figureCitation>
).
</paragraph>
<paragraph id="8E2E369AFFDAFFF419BDFB90FDA7A8AF" blockId="29.[151,1437,330,1556]" pageId="29" pageNumber="30">
Female. Tenth abdominal tergite furrowed in midline. Epiproct rounded, surface shallowly depressed. Cerci long, conical, slightly curved, apex acute. Ovipositor long, compressed, blade-shaped, substraight, highest at end of basal third (
<figureCitation id="16AA2A1FFFDAFFF418CCFB40FDB4A8AF" box="[439,527,1170,1196]" captionStart="FIGURES 8085" captionStartId="24.[151,269,1891,1915]" captionTargetBox="[244,1344,181,1863]" captionTargetId="figure-83@24.[236,1350,176,1867]" captionTargetPageId="24" captionText="FIGURES 8085. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (8284) or 10 mm (81, 85)." figureDoi="http://doi.org/10.5281/zenodo.5123660" httpUri="https://zenodo.org/record/5123660/files/figure.png" pageId="29" pageNumber="30">Fig. 85</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C68B6511FFDAFFF419BDFB68FAB4AA17" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8E2E369AFFDAFFF419BDFB68FAB4AA17" blockId="29.[151,1437,330,1556]" pageId="29" pageNumber="30">
<emphasis id="BCE5EA88FFDAFFF419BDFB68FEE9A8D7" bold="true" box="[198,338,1210,1236]" pageId="29" pageNumber="30">Discussion.</emphasis>
<taxonomicName id="49914D19FFDAFFF41827FB69FDF3A8D7" authorityName="C. Willemse" authorityYear="1961" box="[348,584,1211,1236]" class="Insecta" family="Tettigoniidae" genus="Pseudomacroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41827FB69FDF3A8D7" box="[348,584,1211,1236]" italics="true" pageId="29" pageNumber="30">Pseudomacroxiphus</emphasis>
</taxonomicName>
has the male pronotum not prolonged with the apex truncate and the paranota not deep. A character that it shares with
<taxonomicName id="49914D19FFDAFFF41BA6FB31FC30A8FF" authorityName="Karny" authorityYear="1912" box="[733,907,1251,1276]" class="Insecta" family="Tettigoniidae" genus="Pseudonicsara" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41BA6FB31FC30A8FF" box="[733,907,1251,1276]" italics="true" pageId="29" pageNumber="30">Pseudonicsara</emphasis>
</taxonomicName>
and
<taxonomicName id="49914D19FFDAFFF41ABEFB31FB98A8FF" authorityName="Kirby" authorityYear="1906" box="[965,1059,1251,1276]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41ABEFB31FB98A8FF" box="[965,1059,1251,1276]" italics="true" pageId="29" pageNumber="30">Habetia</emphasis>
</taxonomicName>
. It differs from both by the prolonged stridulatory area of the male tegmina and peculiar modifications of the phallic complex. From
<taxonomicName id="49914D19FFDAFFF41C30FAD9FF45A94F" authorityName="Karny" authorityYear="1912" class="Insecta" family="Tettigoniidae" genus="Pseudonicsara" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41C30FAD9FF45A94F" italics="true" pageId="29" pageNumber="30">Pseudonicsara</emphasis>
</taxonomicName>
it also differs by the fore tibia which is in cross-section quadrangular and by the long, compressed ovipositor. With
<taxonomicName id="49914D19FFDAFFF41821FA89FE03A977" authorityName="Kirby" authorityYear="1906" box="[346,440,1371,1396]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41821FA89FE03A977" box="[346,440,1371,1396]" italics="true" pageId="29" pageNumber="30">Habetia</emphasis>
</taxonomicName>
, it also shares the rather flat, not conical meso- and metasternal lobes. It differs, apart from the above characters, by much longer wings, the possession of a baso-internal projection of the male cercus and the male phallic complex. The genus is not related to
<taxonomicName id="49914D19FFDAFFF41A05FA79FBACA9C7" authorityName="Haan" authorityYear="1842" box="[894,1047,1451,1476]" class="Insecta" family="Tettigoniidae" genus="Macroxiphus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="BCE5EA88FFDAFFF41A05FA79FBACA9C7" box="[894,1047,1451,1476]" italics="true" pageId="29" pageNumber="30">Macroxiphus</emphasis>
</taxonomicName>
. It differs by the morphology of pronotum and sternites, the modified stridulatory apparatus, male abdominal terminalia and titillators, female subgenital plate and even details of the ovipositor, although the latter is prolonged in both genera.
</paragraph>
</subSubSection>
</treatment>
</document>