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<taxonomicName id="B4954D64F22DAC5AFF67F9B5FD00AD43" authority="(Matsumura, 1912)" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[151,687,1574,1601]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sapporensis">
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<emphasis id="41E1EAF5F22DAC5AFF67F9B5FE18AD43" bold="true" box="[151,439,1574,1601]" italics="true" pageId="93" pageNumber="94">Pamphilius sapporensis</emphasis>
(
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)
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(
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,
<figureCitation id="EBAE2A62F22DAC5AFEFFF9D9FE96AD67" box="[271,313,1610,1636]" captionStart="FIGURE 116" captionStartId="205.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-19@205.[151,1435,181,1845]" captionTargetPageId="205" captionText="FIGURE 116. Pamphilius sapporensis, male, Japan (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) fore wing, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876783" httpUri="https://zenodo.org/record/6876783/files/figure.png" pageId="93" pageNumber="94">116</figureCitation>
) (https://doi.org/10.6084/m9.figshare.11405283)
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<emphasis id="41E1EAF5F22DAC5AFF67F91CFEE8ADA5" box="[151,327,1679,1703]" italics="true" pageId="93" pageNumber="94">Lyda sapporensis</emphasis>
<bibRefCitation id="17044B16F22DAC5AFEBEF91CFD85ADA4" author="Matsumura, S." box="[334,554,1679,1703]" pageId="93" pageNumber="94" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura, 1912: 80</bibRefCitation>
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;
<treatmentCitation id="F23410F6F22DAC5AFDC4F91CFD45ADA4" author="Takeuchi, K." box="[564,746,1679,1703]" page="9" pageId="93" pageNumber="94" year="1930">
<bibRefCitation id="17044B16F22DAC5AFDC4F91CFD45ADA4" author="Takeuchi, K." box="[564,746,1679,1703]" pageId="93" pageNumber="94" pagination="3 - 16" refId="ref80368" refString="Takeuchi, K. (1930) A revisional list of the Japanese Pamphiliidae, with description of nine new species. Transactions of the Kansai Entomological Society, 1, 3 - 16." type="journal article" year="1930">Takeuchi, 1930: 9</bibRefCitation>
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(syn. of
<taxonomicName id="B4954D64F22DAC5AFCB4F91CFBFCADA4" authority="(Smith, 1874))" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[836,1107,1679,1703]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22DAC5AFCB4F91CFC1BADA5" box="[836,948,1679,1702]" italics="true" pageId="93" pageNumber="94">P. venustus</emphasis>
(
<bibRefCitation id="17044B16F22DAC5AFC32F91CFBEDADA4" author="Smith, F." box="[962,1090,1679,1703]" pageId="93" pageNumber="94" pagination="373 - 409" refId="ref79508" refString="Smith, F. (1874) Descriptions of new species of Tenthredinidae, Ichneumonidae, Chrysididae, Formicidae &amp; c. of Japan. Transactions of the Entomological Society of London for the Year, 1874, 373 - 409. https: // doi. org / 10.1111 / j. 1365 - 2311.1874. tb 00867. x" type="journal article" year="1874">Smith, 1874</bibRefCitation>
))
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<treatmentCitation id="F23410F6F22DAC5AFF67F93EFDB4ADC6" author="Benes, K." box="[151,539,1709,1733]" page="165" pageId="93" pageNumber="94" year="1976">
<taxonomicName id="B4954D64F22DAC5AFF67F93EFDB4ADC6" authority="Benes, 1976: 165" authorityName="Benes" authorityPageNumber="165" authorityYear="1976" box="[151,539,1709,1733]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="41E1EAF5F22DAC5AFF67F93EFECEADC6" box="[151,353,1709,1733]" italics="true" pageId="93" pageNumber="94">Pamphilius rugosus</emphasis>
<bibRefCitation id="17044B16F22DAC5AFE98F93EFDB4ADC6" author="Benes, K." box="[360,539,1709,1733]" pageId="93" pageNumber="94" pagination="159 - 173" refId="ref72892" refString="Benes, K. (1976) The Siberian species of the genus Pamphilius related to Pamphilius vafer (L.) (Hymenoptera, Pamphiliidae). Acta Entomologica Bohemoslovaca, 73, 159 - 173." type="journal article" year="1976">Beneš, 1976: 165</bibRefCitation>
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;
<treatmentCitation id="F23410F6F22DAC5AFDD6F93EFCDEADC6" author="Shinohara, A. &amp; Okutani, T." box="[550,881,1709,1733]" page="278" pageId="93" pageNumber="94" year="1983">
<bibRefCitation id="17044B16F22DAC5AFDD6F93EFCDEADC6" author="Shinohara, A. &amp; Okutani, T." box="[550,881,1709,1733]" pageId="93" pageNumber="94" pagination="276 - 281" refId="ref78722" refString="Shinohara, A. &amp; Okutani, T. (1983) Host-plants of Japanese Pamphiliidae (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 51, 276 - 281." type="journal article" year="1983">Shinohara &amp; Okutani, 1983: 278</bibRefCitation>
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(syn. of
<taxonomicName id="B4954D64F22DAC5AFC3BF93DFBF3ADC6" authorityName="Matsumura" authorityYear="1912" box="[971,1116,1710,1733]" class="Insecta" family="Pamphiliidae" genus="Lyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22DAC5AFC3BF93DFBF3ADC6" box="[971,1116,1710,1733]" italics="true" pageId="93" pageNumber="94">P. sapporensis</emphasis>
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).
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<treatmentCitation id="F23410F6F22DAC5AFF67F95FFD4AADE7" author="Shinohara, A. &amp; Okutani, T." box="[151,741,1740,1764]" page="278" pageId="93" pageNumber="94" year="1983">
<taxonomicName id="B4954D64F22DAC5AFF67F95FFD4AADE7" authority=": Shinohara &amp; Okutani, 1983: 278" authorityName=": Shinohara &amp; Okutani" authorityPageNumber="278" authorityYear="1983" box="[151,741,1740,1764]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22DAC5AFF67F95FFE26ADE7" box="[151,393,1740,1764]" italics="true" pageId="93" pageNumber="94">Pamphilius sapporensis</emphasis>
:
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;
<treatmentCitation id="F23410F6F22DAC5AFD1FF95FFBDBADE7" author="Zhelochovtsev, A. N. &amp; Zinovjev, A. G." box="[751,1140,1740,1764]" page="398" pageId="93" pageNumber="94" year="1995">
<bibRefCitation id="17044B16F22DAC5AFD1FF95FFBDBADE7" author="Zhelochovtsev, A. N. &amp; Zinovjev, A. G." box="[751,1140,1740,1764]" pageId="93" pageNumber="94" pagination="395 - 415" refId="ref81182" refString="Zhelochovtsev, A. N. &amp; Zinovjev, A. G. (1995) A list of the sawflies and horntails (Hymenoptera, Symphyta) of the fauna of Russia and adjacent territories. I. Entomologicheskoe Obozrenie, 74, 395 - 415. [in Russian]" type="journal article" year="1995">Zhelochovtsev &amp; Zinovjev, 1995: 398</bibRefCitation>
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;
<treatmentCitation id="F23410F6F22DAC5AFB8FF95FFAC7ADE7" author="Shinohara, A." box="[1151,1384,1740,1764]" page="427" pageId="93" pageNumber="94" year="2002">
<bibRefCitation id="17044B16F22DAC5AFB8FF95FFAC7ADE7" author="Shinohara, A." box="[1151,1384,1740,1764]" pageId="93" pageNumber="94" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara, 2002b: 427</bibRefCitation>
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; Shinohara, 2004: 264;
<treatmentCitation id="F23410F6F22DAC5AFE7AF978FD14AC00" author="Shinohara, A. &amp; Taeger, A." box="[394,699,1771,1795]" page="38" pageId="93" pageNumber="94" year="2007">
<bibRefCitation id="17044B16F22DAC5AFE7AF978FD14AC00" author="Shinohara, A. &amp; Taeger, A." box="[394,699,1771,1795]" pageId="93" pageNumber="94" pagination="31 - 40" refId="ref78859" refString="Shinohara, A. &amp; Taeger, A. (2007) Description of a new species and collection data of pamphiliid sawflies (Insecta, Hymenoptera, Symphyta) mainly from the Russian Far East. Bulletin of the National Museum of Nature and Science, Tokyo, Series A, 33, 31 - 40." type="journal article" year="2007">Shinohara &amp; Taeger, 2007: 38</bibRefCitation>
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;
<treatmentCitation id="F23410F6F22DAC5AFD35F978FC5DAC00" author="Shinohara, A. &amp; Lelej, A. S." box="[709,1010,1771,1795]" page="931" pageId="93" pageNumber="94" year="2007">
<bibRefCitation id="17044B16F22DAC5AFD35F978FC5DAC00" author="Shinohara, A. &amp; Lelej, A. S." box="[709,1010,1771,1795]" pageId="93" pageNumber="94" pagination="922 - 942" refId="ref78590" refString="Shinohara, A. &amp; Lelej, A. S. (2007) 2. Sem. Pamphiliidae - Pautinnye Pilil'shchiki, ili Pilil'shchiki-tkachi. In: Lelej, A. S. (Ed.), Setchatokryloobraznye, Skorpionnitsy, Pereponchatokrylye. Ch. 5. Opredelitel' Nasekomykh Dal'nego Vostoka Rossii. Tom IV. Dal'nauka, Vladivostok, pp. 922 - 942." type="book chapter" year="2007">Shinohara &amp; Lelej, 2007: 931</bibRefCitation>
</treatmentCitation>
, 940;
<treatmentCitation id="F23410F6F22DAC5AFBDEF978FABDAC00" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1070,1298,1771,1795]" page="90" pageId="93" pageNumber="94" year="2010">
<bibRefCitation id="17044B16F22DAC5AFBDEF978FABDAC00" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[1070,1298,1771,1795]" pageId="93" pageNumber="94" pagination="1 - 1064" refId="ref80219" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
Taeger
<emphasis id="41E1EAF5F22DAC5AFB89F978FB04AC00" box="[1145,1195,1771,1795]" italics="true" pageId="93" pageNumber="94">et al.</emphasis>
, 2010: 90
</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="17044B16F22DAC5AFAECF978FA32AC00" author="Sundukov, Yu. N. &amp; Lelej, A. S." box="[1308,1437,1771,1795]" pageId="93" pageNumber="94" pagination="62 - 119" refId="ref79983" refString="Sundukov, Yu. N. &amp; Lelej, A. S. (2012) Podotryad Symphyta - Sidyachebryukhie. In: Lelej, A. S. (Ed), Annotated Catalogue of the Insects of Russian Far East. Vol. I. Hymenoptera. Dalnauka, Vladivostok, pp. 62 - 119. [in Russian]" type="book chapter" year="2012">Sundukov &amp;</bibRefCitation>
Lelej, 2012: 109;
<bibRefCitation id="17044B16F22DAC5AFE8BF89AFDF9AC22" author="Sundukov, Yu. N." box="[379,598,1801,1825]" pageId="93" pageNumber="94" pagination="241 - 258" refId="ref79811" refString="Sundukov, Yu. N. (2015) To the sawfly fauna (Hymenoptera, Symphyta) of southern Kuriles. A. I. Kurentsov's Annual Memorial Meetings, Issue XXVI, pp. 241 - 258. [in Russian with English abstract]" type="journal article" year="2015">Sundukov, 2015: 250</bibRefCitation>
;
<bibRefCitation id="17044B16F22DAC5AFD90F89AFC94AC22" author="Sundukov, Yu. N." box="[608,827,1801,1825]" pageId="93" pageNumber="94" pagination="20 - 117" refId="ref79858" refString="Sundukov, Yu. N. (2017) Suborder Symphyta - Sawflies and wood wasps. In: Lelej, A. S., Proshchalykin, M. Yu. &amp; Loktionov, V. M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supplement 6, pp. 20 - 117." type="journal article" year="2017">Sundukov, 2017: 105</bibRefCitation>
; Shinohara, 2019: 11;
<bibRefCitation id="17044B16F22DAC5AFBEFF89AFB42AC22" author="Shinohara, A." box="[1055,1261,1801,1825]" pageId="93" pageNumber="94" pagination="222 - 254" refId="ref77165" refString="Shinohara, A. (2020) Family Pamphiliidae. In: Naito, T., Shinohara, A., Hara, H. &amp; Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 3 - 23 + 222 - 254. [in Japanese]" type="book chapter" year="2020">Shinohara, 2020: 13</bibRefCitation>
, 243.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF22DAC5AFF67F8C3FAF4ACB2" pageId="93" pageNumber="94" type="description">
<paragraph id="732A36E7F22DAC5AFF67F8C3FAF4ACB2" blockId="93.[151,1437,1871,2042]" pageId="93" pageNumber="94">
<materialsCitation id="C3FD3CBAF22DAC5AFF67F8C3FAF4ACB2" ID-GBIF-Occurrence="3866537372" collectionCode="HU" collectorName="Here &amp; Matsumura's" determinerName="Matsumura" pageId="93" pageNumber="94" specimenCount="2" typeStatus="lectotype">
<emphasis id="41E1EAF5F22DAC5AFF67F8C3FE0CAC6A" bold="true" box="[151,419,1871,1898]" pageId="93" pageNumber="94">
<typeStatus id="AC2E8845F22DAC5AFF67F8C3FEA2AC69" box="[151,269,1872,1898]" pageId="93" pageNumber="94" type="lectotype">Lectotype</typeStatus>
designation.
</emphasis>
<bibRefCitation id="17044B16F22DAC5AFE59F8C3FDD7AC69" author="Matsumura, S." box="[425,632,1872,1898]" pageId="93" pageNumber="94" refId="ref74972" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha, Tokyo, 1 p. + 247 pp., 14 pls., 2 + 2 pp. [in Japanese and English]" type="book" year="1912">Matsumura (1912)</bibRefCitation>
described
<taxonomicName id="B4954D64F22DAC5AFD1FF8DCFC1EAC6A" authorityName="Matsumura" authorityYear="1912" box="[751,945,1871,1897]" class="Insecta" family="Pamphiliidae" genus="Lyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22DAC5AFD1FF8DCFC1EAC6A" box="[751,945,1871,1897]" italics="true" pageId="93" pageNumber="94">Lyda sapporensis</emphasis>
</taxonomicName>
without giving number of the specimens examined.
<collectorName id="DE605331F22DAC5AFF06F8E7FE83AC8D" box="[246,300,1908,1934]" pageId="93" pageNumber="94">Here</collectorName>
we designate the male specimen labeled “Sahoro, 6/5” “29” “
<taxonomicName id="B4954D64F22DAC5AFC29F8E0FB34AC8E" authority="Shinohara &amp; Kramp &amp; Taeger, 2022" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[985,1179,1907,1933]" class="Insecta" family="Pamphiliidae" genus="Lyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="sapporensis" status="sp. nov.">
<emphasis id="41E1EAF5F22DAC5AFC29F8E0FB34AC8E" box="[985,1179,1907,1933]" italics="true" pageId="93" pageNumber="94">Lyda sapporensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AD2578EF22DAC5AFB51F8E7FB74AC8D" box="[1185,1243,1908,1934]" pageId="93" pageNumber="94" rank="species">n. sp.</taxonomicNameLabel>
det.
<determinerName id="86F6DA23F22DAC5AFAE0F8E7FA3DAC8D" box="[1296,1426,1908,1934]" pageId="93" pageNumber="94">Matsumura</determinerName>
” in
<collectorName id="DE605331F22DAC5AFF44F80BFEE9ACB1" box="[180,326,1944,1970]" pageId="93" pageNumber="94">Matsumuras</collectorName>
collection (
<collectionCode id="1584AE22F22DAC5AFE3AF80BFE5AACB1" box="[458,501,1944,1970]" country="China" lsid="urn:lsid:biocol.org:col:14677" name="University of Zhejiang" pageId="93" pageNumber="94" type="Herbarium">HU</collectionCode>
) as a
<typeStatus id="AC2E8845F22DAC5AFDC6F804FD0FACB2" box="[566,672,1943,1969]" pageId="93" pageNumber="94" type="lectotype">lectotype</typeStatus>
. It is in poor condition and has the head and genitalia missing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF22DAC5AFF37F828FD7AACF9" pageId="93" pageNumber="94" type="materials_examined">
<paragraph id="732A36E7F22DAC5AFF37F828FD7AACF9" blockId="93.[151,1437,1871,2042]" pageId="93" pageNumber="94">
<emphasis id="41E1EAF5F22DAC5AFF37F828FE1DACD6" bold="true" box="[199,434,1979,2005]" pageId="93" pageNumber="94">Material examined.</emphasis>
About
<specimenCount id="6593FD6EF22DAC5AFDF7F828FD1EACD6" box="[519,689,1979,2006]" pageId="93" pageNumber="94" type="generic">545 specimens</specimenCount>
, including the
<typeStatus id="AC2E8845F22DAC5AFCA9F828FC6CACD6" box="[857,963,1979,2005]" pageId="93" pageNumber="94" type="lectotype">lectotype</typeStatus>
, and
<specimenCount id="6593FD6EF22DAC5AFC0FF82FFB09ACD6" box="[1023,1190,1979,2006]" pageId="93" pageNumber="94" type="generic">two specimens</specimenCount>
from the Russian Far East (
<bibRefCitation id="17044B16F22DAC5AFF26F84CFE35ACFA" author="Shinohara, A." box="[214,410,2015,2041]" pageId="93" pageNumber="94" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara 2002b</bibRefCitation>
;
<bibRefCitation id="17044B16F22DAC5AFE55F84CFD65ACF9" author="Shinohara, A. &amp; Taeger, A." box="[421,714,2015,2042]" pageId="93" pageNumber="94" pagination="31 - 40" refId="ref78859" refString="Shinohara, A. &amp; Taeger, A. (2007) Description of a new species and collection data of pamphiliid sawflies (Insecta, Hymenoptera, Symphyta) mainly from the Russian Far East. Bulletin of the National Museum of Nature and Science, Tokyo, Series A, 33, 31 - 40." type="journal article" year="2007">Shinohara &amp; Taeger 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF22EAC59FF37FF05FB77ABB3" box="[199,1240,150,177]" pageId="94" pageNumber="95" type="distribution">
<paragraph id="732A36E7F22EAC59FF37FF05FB77ABB3" blockId="94.[151,1437,150,645]" box="[199,1240,150,177]" pageId="94" pageNumber="95">
<emphasis id="41E1EAF5F22EAC59FF37FF05FEF6ABB3" bold="true" box="[199,345,150,176]" pageId="94" pageNumber="95">Distribution</emphasis>
.
<collectingCountry id="0B827677F22EAC59FE97FF05FE1DABB3" box="[359,434,150,176]" name="Russia" pageId="94" pageNumber="95">Russia</collectingCountry>
(
<collectingRegion id="B151F805F22EAC59FE31FF05FD86ABB3" box="[449,553,150,176]" country="Russia" name="Sakhalin" pageId="94" pageNumber="95">Sakhalin</collectingRegion>
),
<collectingCountry id="0B827677F22EAC59FDCCFF04FDD3ABB2" box="[572,636,151,177]" name="Japan" pageId="94" pageNumber="95">Japan</collectingCountry>
(
<collectingRegion id="B151F805F22EAC59FD7AFF05FD52ABB3" box="[650,765,150,176]" country="Japan" name="Hokkaido" pageId="94" pageNumber="95">Hokkaido</collectingRegion>
, Shikotan Is.) (
<bibRefCitation id="17044B16F22EAC59FC56FF05FB63ABB3" author="Shinohara, A. &amp; Taeger, A." box="[934,1228,150,177]" pageId="94" pageNumber="95" pagination="31 - 40" refId="ref78859" refString="Shinohara, A. &amp; Taeger, A. (2007) Description of a new species and collection data of pamphiliid sawflies (Insecta, Hymenoptera, Symphyta) mainly from the Russian Far East. Bulletin of the National Museum of Nature and Science, Tokyo, Series A, 33, 31 - 40." type="journal article" year="2007">Shinohara &amp; Taeger 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF22EAC59FF37FF29FB4CABD7" box="[199,1251,186,213]" pageId="94" pageNumber="95" type="biology_ecology">
<paragraph id="732A36E7F22EAC59FF37FF29FB4CABD7" blockId="94.[151,1437,150,645]" box="[199,1251,186,213]" pageId="94" pageNumber="95">
<emphasis id="41E1EAF5F22EAC59FF37FF29FEEBABD7" bold="true" box="[199,324,186,212]" pageId="94" pageNumber="95">Host plant</emphasis>
.
<taxonomicName id="B4954D64F22EAC59FEA2FF28FE6FABD6" box="[338,448,187,213]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
:
<taxonomicName id="B4954D64F22EAC59FE3BFF29FC3BABD7" authority="(Pall.) Maxim." authorityName="Maxim." baseAuthorityName="Pall." box="[459,916,186,212]" class="Magnoliopsida" family="Rosaceae" genus="Filipendula" kingdom="Plantae" order="Rosales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="species" species="camtschatica">
<emphasis id="41E1EAF5F22EAC59FE3BFF29FD46ABD7" box="[459,745,186,212]" italics="true" pageId="94" pageNumber="95">Filipendula camtschatica</emphasis>
(Pall.) Maxim.
</taxonomicName>
(
<bibRefCitation id="17044B16F22EAC59FC52FF29FB79ABD7" author="Shinohara, A. &amp; Okutani, T." box="[930,1238,186,212]" pageId="94" pageNumber="95" pagination="276 - 281" refId="ref78722" refString="Shinohara, A. &amp; Okutani, T. (1983) Host-plants of Japanese Pamphiliidae (Hymenoptera, Pamphiliidae). Kontyu, Tokyo, 51, 276 - 281." type="journal article" year="1983">Shinohara &amp; Okutani 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF22EAC59FF37FF4DFE67A987" pageId="94" pageNumber="95" type="discussion">
<paragraph id="732A36E7F22EAC59FF37FF4DFC20AAF6" blockId="94.[151,1437,150,645]" pageId="94" pageNumber="95">
<emphasis id="41E1EAF5F22EAC59FF37FF4DFE9BABFB" bold="true" box="[199,308,222,248]" pageId="94" pageNumber="95">Remarks</emphasis>
.
<bibRefCitation id="17044B16F22EAC59FEB2FF4DFDBCABFB" author="Shinohara, A." box="[322,531,222,248]" pageId="94" pageNumber="95" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara (2002b)</bibRefCitation>
placed this species in his
<taxonomicName id="B4954D64F22EAC59FCC4FF4CFC00ABFB" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[820,943,223,248]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22EAC59FCC4FF4CFC00ABFB" box="[820,943,223,248]" italics="true" pageId="94" pageNumber="95">P. venustus</emphasis>
</taxonomicName>
complex of the
<taxonomicName id="B4954D64F22EAC59FB95FF4CFAA0ABFB" box="[1125,1295,223,248]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="stramineipes">
<emphasis id="41E1EAF5F22EAC59FB95FF4CFAA0ABFB" box="[1125,1295,223,248]" italics="true" pageId="94" pageNumber="95">P. stramineipes</emphasis>
</taxonomicName>
subgroup of the
<taxonomicName id="B4954D64F22EAC59FF32FE90FEB7AA1F" authorityName=": Malaise" authorityYear="1931" box="[194,280,258,284]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="vafer">
<emphasis id="41E1EAF5F22EAC59FF32FE90FEB7AA1F" box="[194,280,258,284]" italics="true" pageId="94" pageNumber="95">P. vafer</emphasis>
</taxonomicName>
group.
<taxonomicName id="B4954D64F22EAC59FE81FE91FDD2AA1F" authorityName=": Shinohara &amp; Okutani" authorityYear="1983" box="[369,637,258,284]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22EAC59FE81FE91FDD2AA1F" box="[369,637,258,284]" italics="true" pageId="94" pageNumber="95">Pamphilius sapporensis</emphasis>
</taxonomicName>
has much in common with
<taxonomicName id="B4954D64F22EAC59FC4CFE90FB97AA1F" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[956,1080,259,284]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22EAC59FC4CFE90FB97AA1F" box="[956,1080,259,284]" italics="true" pageId="94" pageNumber="95">P. venustus</emphasis>
</taxonomicName>
in morphology, molecular data (see below) and host plants, and overlapping distributions of the two species in
<collectingRegion id="B151F805F22EAC59FBF9FEB5FBD8AA43" box="[1033,1143,294,320]" country="Japan" name="Hokkaido" pageId="94" pageNumber="95">Hokkaido</collectingRegion>
is interesting in clarifying their speciation history. The maximum intraspecific
<emphasis id="41E1EAF5F22EAC59FD1CFED8FD55AA67" box="[748,762,331,356]" italics="true" pageId="94" pageNumber="95">p</emphasis>
-distance among the three Japanese specimens in
<taxonomicName id="B4954D64F22EAC59FAC2FED8FF76AA8B" authorityName=": Shinohara &amp; Okutani" authorityYear="1983" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22EAC59FAC2FED8FF76AA8B" italics="true" pageId="94" pageNumber="95">P. sapporensis</emphasis>
</taxonomicName>
was 0.3% in COI and 0.1% in NaK and the nearest neighbour was
<taxonomicName id="B4954D64F22EAC59FC25FEFCFBF7AA8B" authorityName=": Konow" authorityYear="1897" box="[981,1112,366,392]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="balteatus">
<emphasis id="41E1EAF5F22EAC59FC25FEFCFBF7AA8B" box="[981,1112,366,392]" italics="true" pageId="94" pageNumber="95">P. balteatus</emphasis>
</taxonomicName>
, diverging by a minimum of 2.5% in the COI analysis, and
<taxonomicName id="B4954D64F22EAC59FE17FE00FDCDAAAF" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[487,610,403,428]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22EAC59FE17FE00FDCDAAAF" box="[487,610,403,428]" italics="true" pageId="94" pageNumber="95">P. venustus</emphasis>
</taxonomicName>
by a minimum of 0.8% in the NaK analysis. In the COI tree (
<figureCitation id="EBAE2A62F22EAC59FAF6FE01FAC8AAAF" box="[1286,1383,402,428]" captionStart="FIGURE 149" captionStartId="236.[151,250,1943,1967]" captionTargetBox="[169,1416,198,1910]" captionTargetId="figure-26@236.[163,1425,181,1919]" captionTargetPageId="236" captionText="FIGURE 149. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius vafer group, part 2." figureDoi="http://doi.org/10.5281/zenodo.6876861" httpUri="https://zenodo.org/record/6876861/files/figure.png" pageId="94" pageNumber="95">Fig. 149</figureCitation>
), the relationship of
<taxonomicName id="B4954D64F22EAC59FECEFE24FE72AAD3" authorityName=": Shinohara &amp; Okutani" authorityYear="1983" box="[318,477,439,464]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22EAC59FECEFE24FE72AAD3" box="[318,477,439,464]" italics="true" pageId="94" pageNumber="95">P. sapporensis</emphasis>
</taxonomicName>
with other species was not well resolved, but in the NaK tree (
<figureCitation id="EBAE2A62F22EAC59FB7AFE25FB45AAD3" box="[1162,1258,438,464]" captionStart="FIGURE 161" captionStartId="246.[151,250,1943,1967]" captionTargetBox="[177,1403,189,1910]" captionTargetId="figure-26@246.[171,1417,181,1919]" captionTargetPageId="246" captionText="FIGURE 161. Condensed ML tree based on NaK gene sequences, with details of Pamphilius vafer group, part 2." figureDoi="http://doi.org/10.5281/zenodo.6876886" httpUri="https://zenodo.org/record/6876886/files/figure.png" pageId="94" pageNumber="95">Fig. 161</figureCitation>
),
<taxonomicName id="B4954D64F22EAC59FB0DFE24FA33AAD3" authorityName=": Shinohara &amp; Okutani" authorityYear="1983" box="[1277,1436,439,464]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22EAC59FB0DFE24FA33AAD3" box="[1277,1436,439,464]" italics="true" pageId="94" pageNumber="95">P. sapporensis</emphasis>
</taxonomicName>
was retrieved as the sister of
<taxonomicName id="B4954D64F22EAC59FE2CFE48FDF7AAF7" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[476,600,475,500]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22EAC59FE2CFE48FDF7AAF7" box="[476,600,475,500]" italics="true" pageId="94" pageNumber="95">P. venustus</emphasis>
</taxonomicName>
with 99% UFBoot support.
</paragraph>
<paragraph id="732A36E7F22EAC59FF37FE6CFE67A987" blockId="94.[151,1437,150,645]" pageId="94" pageNumber="95">
In
<collectingRegion id="B151F805F22EAC59FF15FE6DFEF9A91B" box="[229,342,510,536]" country="Japan" name="Hokkaido" pageId="94" pageNumber="95">Hokkaido</collectingRegion>
,
<collectingCountry id="0B827677F22EAC59FE90FE6CFE0CA91A" box="[352,419,511,537]" name="Japan" pageId="94" pageNumber="95">Japan</collectingCountry>
, this species occurs together with two other
<taxonomicName id="B4954D64F22EAC59FC7BFE6DFBA1A91B" baseAuthorityName="Kangas &amp; Syrjanen" baseAuthorityYear="1962" box="[907,1038,510,536]" class="Magnoliopsida" family="Rosaceae" genus="Filipendula" kingdom="Plantae" order="Rosales" pageId="94" pageNumber="95" phylum="Tracheophyta" rank="genus">
<emphasis id="41E1EAF5F22EAC59FC7BFE6DFBA1A91B" box="[907,1038,510,536]" italics="true" pageId="94" pageNumber="95">Filipendula</emphasis>
</taxonomicName>
-associated pamphiliids,
<taxonomicName id="B4954D64F22EAC59FAD1FE6CFA33A91B" authorityName="NSMT" baseAuthorityName="Smith" baseAuthorityYear="1874" box="[1313,1436,511,536]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="41E1EAF5F22EAC59FAD1FE6CFA33A91B" box="[1313,1436,511,536]" italics="true" pageId="94" pageNumber="95">P. venustus</emphasis>
</taxonomicName>
(known from
<collectingRegion id="B151F805F22EAC59FEC5FDB1FE09A93F" box="[309,422,546,572]" country="Japan" name="Hokkaido" pageId="94" pageNumber="95">Hokkaido</collectingRegion>
, Honshu and Shikoku) and
<taxonomicName id="B4954D64F22EAC59FD17FDB1FBA2A93F" authorityName=": Benes" authorityYear="1972" box="[743,1037,546,572]" class="Insecta" family="Pamphiliidae" genus="Onycholyda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="kumamotonis">
<emphasis id="41E1EAF5F22EAC59FD17FDB1FBA2A93F" box="[743,1037,546,572]" italics="true" pageId="94" pageNumber="95">Onycholyda kumamotonis</emphasis>
</taxonomicName>
(known from Sakhalin,
<collectingRegion id="B151F805F22EAC59FAD7FDB1FA37A93F" box="[1319,1432,546,572]" country="Japan" name="Hokkaido" pageId="94" pageNumber="95">Hokkaido</collectingRegion>
, Kunashiri Is., Honshu and?Kyushu). The largely orange upper part of head will easily distinguish
<taxonomicName id="B4954D64F22EAC59FB0BFDD4FA33A963" authorityName=": Shinohara &amp; Okutani" authorityYear="1983" box="[1275,1436,583,608]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="sapporensis">
<emphasis id="41E1EAF5F22EAC59FB0BFDD4FA33A963" box="[1275,1436,583,608]" italics="true" pageId="94" pageNumber="95">P. sapporensis</emphasis>
</taxonomicName>
from the latter two species.
</paragraph>
</subSubSection>
</treatment>
</document>