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<taxonomicName id="B4954D64F221AC56FF67F938FD2BADC5" authority="Shinohara, 1988" authorityName="Shinohara" authorityYear="1988" box="[151,644,1707,1734]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="ussuriensis">
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<emphasis id="41E1EAF5F221AC56FF67F938FE1FADC5" bold="true" box="[151,432,1707,1734]" italics="true" pageId="81" pageNumber="82">Pamphilius ussuriensis</emphasis>
Shinohara, 1988
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(
<figureCitation id="EBAE2A62F221AC56FF6FF943FE94ADE9" box="[159,315,1744,1770]" captionStart-0="FIGURE 123" captionStart-1="FIGURE 124" captionStart-2="FIGURE 125" captionStart-3="FIGURE 126" captionStartId-0="212.[151,250,1869,1893]" captionStartId-1="213.[151,250,1869,1893]" captionStartId-2="214.[151,250,1748,1772]" captionStartId-3="215.[151,250,1869,1893]" captionTargetBox-0="[151,1435,181,1845]" captionTargetBox-1="[151,1435,181,1845]" captionTargetBox-2="[151,1435,181,1722]" captionTargetBox-3="[151,1435,181,1845]" captionTargetId-0="figure-26@212.[151,1435,181,1845]" captionTargetId-1="figure-19@213.[151,1435,181,1845]" captionTargetId-2="figure-26@214.[151,1435,181,1724]" captionTargetId-3="figure-19@215.[151,1435,181,1845]" captionTargetPageId-0="212" captionTargetPageId-1="213" captionTargetPageId-2="214" captionTargetPageId-3="215" captionText-0="FIGURE 123. Pamphilius ussuriensis, female, Russia – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) sawsheath laterally, (h) claw." captionText-1="FIGURE 124. Pamphilius ussuriensis, male, Russia – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." captionText-2="FIGURE 125. Pamphilius ussuriensis, male, Russia (holotype) – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head antenna dorsolaterally, (e) face." captionText-3="FIGURE 126. Pamphilius ussuriensis (specimen 86348 and holotype of P. zinovjevi), female, Russia – (a) head &amp; thorax dorsally (86348), (b) head &amp; thorax ventrolaterally (86348), (c) face (86348), (d) dorsally, (e) lateroventrally, (f) face." figureDoi-0="http://doi.org/10.5281/zenodo.6876799" figureDoi-1="http://doi.org/10.5281/zenodo.6876801" figureDoi-2="http://doi.org/10.5281/zenodo.6876803" figureDoi-3="http://doi.org/10.5281/zenodo.6876805" httpUri-0="https://zenodo.org/record/6876799/files/figure.png" httpUri-1="https://zenodo.org/record/6876801/files/figure.png" httpUri-2="https://zenodo.org/record/6876803/files/figure.png" httpUri-3="https://zenodo.org/record/6876805/files/figure.png" pageId="81" pageNumber="82">Figs 123–126</figureCitation>
) (https://doi.org/10.6084/m9.figshare.11405316)
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<emphasis id="41E1EAF5F221AC56FF67F887FE2EAC2F" box="[151,385,1812,1836]" italics="true" pageId="81" pageNumber="82">Pamphilius ussuriensis</emphasis>
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Taeger
<emphasis id="41E1EAF5F221AC56FCABF8A7FC22AC48" box="[859,909,1843,1867]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
, 2010: 91
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<emphasis id="41E1EAF5F221AC56FF67F8E3FEC1AC8B" box="[151,366,1904,1928]" italics="true" pageId="81" pageNumber="82">Pamphilius zinovjevi</emphasis>
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, 114;
<treatmentCitation id="F23410F6F221AC56FADFF8E3FE91ACA4" author="Shinohara, A." page="431" pageId="81" pageNumber="82" year="2002">
<bibRefCitation id="17044B16F221AC56FADFF8E3FE91ACA4" author="Shinohara, A." pageId="81" pageNumber="82" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara, 2002b: 431</bibRefCitation>
</treatmentCitation>
; Shinohara, 2004: 265;
<treatmentCitation id="F23410F6F221AC56FDC2F81CFCC9ACA4" author="Shinohara, A. &amp; Lelej, A. S." box="[562,870,1935,1959]" page="933" pageId="81" pageNumber="82" year="2007">
<bibRefCitation id="17044B16F221AC56FDC2F81CFCC9ACA4" author="Shinohara, A. &amp; Lelej, A. S." box="[562,870,1935,1959]" pageId="81" pageNumber="82" pagination="922 - 942" refId="ref78590" refString="Shinohara, A. &amp; Lelej, A. S. (2007) 2. Sem. Pamphiliidae - Pautinnye Pilil'shchiki, ili Pilil'shchiki-tkachi. In: Lelej, A. S. (Ed.), Setchatokryloobraznye, Skorpionnitsy, Pereponchatokrylye. Ch. 5. Opredelitel' Nasekomykh Dal'nego Vostoka Rossii. Tom IV. Dal'nauka, Vladivostok, pp. 922 - 942." type="book chapter" year="2007">Shinohara &amp; Lelej, 2007: 933</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F23410F6F221AC56FC80F803FBF5ACA4" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[880,1114,1935,1960]" page="92" pageId="81" pageNumber="82" year="2010">
<bibRefCitation id="17044B16F221AC56FC80F803FBF5ACA4" author="Taeger, A. &amp; Blank, S. M. &amp; Liston, A. D." box="[880,1114,1935,1960]" pageId="81" pageNumber="82" pagination="1 - 1064" refId="ref80219" refString="Taeger, A., Blank, S. M. &amp; Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
Taeger
<emphasis id="41E1EAF5F221AC56FC4CF803FC5FACA4" box="[956,1008,1935,1959]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
, 2010: 92
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F23410F6F221AC56FB95F81CFA37ACA4" author="Sundukov, Yu. N. &amp; Lelej, A. S." box="[1125,1432,1935,1959]" page="110" pageId="81" pageNumber="82" year="2012">
<bibRefCitation id="17044B16F221AC56FB95F81CFA37ACA4" author="Sundukov, Yu. N. &amp; Lelej, A. S." box="[1125,1432,1935,1959]" pageId="81" pageNumber="82" pagination="62 - 119" refId="ref79983" refString="Sundukov, Yu. N. &amp; Lelej, A. S. (2012) Podotryad Symphyta - Sidyachebryukhie. In: Lelej, A. S. (Ed), Annotated Catalogue of the Insects of Russian Far East. Vol. I. Hymenoptera. Dalnauka, Vladivostok, pp. 62 - 119. [in Russian]" type="book chapter" year="2012">Sundukov &amp; Lelej, 2012: 110</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F23410F6F221AC56FF37F83DFE0DACC5" author="Sundukov, Yu. N." box="[199,418,1966,1990]" page="106" pageId="81" pageNumber="82" year="2017">
<bibRefCitation id="17044B16F221AC56FF37F83DFE0DACC5" author="Sundukov, Yu. N." box="[199,418,1966,1990]" pageId="81" pageNumber="82" pagination="20 - 117" refId="ref79858" refString="Sundukov, Yu. N. (2017) Suborder Symphyta - Sawflies and wood wasps. In: Lelej, A. S., Proshchalykin, M. Yu. &amp; Loktionov, V. M. (Eds.), Annotated catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences, Supplement 6, pp. 20 - 117." type="journal article" year="2017">Sundukov, 2017: 106</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F23410F6F221AC56FE5CF83DFDDBACC5" author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." box="[428,628,1966,1990]" page="12" pageId="81" pageNumber="82" year="2019">
<bibRefCitation id="17044B16F221AC56FE5CF83DFDDBACC5" author="Lee, J. - W. &amp; Choi, J. - K. &amp; Park, B." box="[428,628,1966,1990]" pageId="81" pageNumber="82" pagination="1 - 96" refId="ref74711" refString="Lee, J. - W., Choi, J. - K. &amp; Park, B. (2019) Synoptic list of Symphyta (Hymenoptera) in Korea. Journal of Species Research, 8, 1 - 96." type="journal article" year="2019">
Lee
<emphasis id="41E1EAF5F221AC56FE29F83CFDA3ACC5" box="[473,524,1966,1990]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
, 2019: 12
</bibRefCitation>
</treatmentCitation>
.
<emphasis id="41E1EAF5F221AC56FD8EF83EFD70ACC5" bold="true" box="[638,735,1965,1990]" pageId="81" pageNumber="82">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF222AC55FF67FF05FAD1AA67" pageId="82" pageNumber="83" type="materials_examined">
<paragraph id="732A36E7F222AC55FF67FF05FAD1AA67" blockId="82.[151,1437,150,861]" pageId="82" pageNumber="83">
<emphasis id="41E1EAF5F222AC55FF67FF05FE2FABB3" bold="true" box="[151,384,150,176]" pageId="82" pageNumber="83">Material examined.</emphasis>
Ten specimens, including the
<typeStatus id="AC2E8845F222AC55FD22FF05FC90ABB3" box="[722,831,150,176]" pageId="82" pageNumber="83" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="B4954D64F222AC55FC93FF04FC55ABB3" authorityName="Shinohara" authorityYear="1988" box="[867,1018,151,176]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FC93FF04FC55ABB3" box="[867,1018,151,176]" italics="true" pageId="82" pageNumber="83">P. ussuriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F222AC55FBC0FF04FB01ABB3" authorityName="Shinohara" authorityYear="1988" box="[1072,1198,151,176]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FBC0FF04FB01ABB3" box="[1072,1198,151,176]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
, all from the Russian Far East and South Korea (
<bibRefCitation id="17044B16F222AC55FE23FF29FD37ABD7" author="Shinohara, A." box="[467,664,186,212]" pageId="82" pageNumber="83" pagination="307 - 320" refId="ref76348" refString="Shinohara, A. (1988 a) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Five new species and additional records from the Far East. Kontyu, Tokyo, 56, 307 - 320." type="journal article" year="1988">
Shinohara 1988
<bibRefCitation id="17044B16F222AC55FD7BFF29FD37ABD7" author="Shinohara, A." box="[651,664,186,212]" pageId="82" pageNumber="83" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">a</bibRefCitation>
</bibRefCitation>
, 2001; present work).
<emphasis id="41E1EAF5F222AC55FC6EFF29FB39ABD7" bold="true" box="[926,1174,186,212]" pageId="82" pageNumber="83">New collection data:</emphasis>
<materialsCitation id="C3FD3CBAF222AC55FB50FF29FD32AA1F" ID-GBIF-Occurrence="3866537389" collectionCode="V" collectorName="A. Taeger" elevation="250" latitude="44.46" location="Sikhote-Alin Reserve" longLatPrecision="682" longitude="134.13" municipality="Upper River Dzhigitovka" pageId="82" pageNumber="83" specimenCount="1" specimenCount-male="1">
<emphasis id="41E1EAF5F222AC55FB50FF29FF7EABFB" bold="true" pageId="82" pageNumber="83">
<collectingCountry id="0B827677F222AC55FB50FF29FAA4ABD7" box="[1184,1291,186,212]" name="Russia" pageId="82" pageNumber="83">RUSSIA</collectingCountry>
:
<collectingRegion id="B151F805F222AC55FAEAFF29FF7EABFB" country="Russia" name="Primorskiy" pageId="82" pageNumber="83">Primorskij Kraj</collectingRegion>
</emphasis>
:
<specimenCount id="6593FD6EF222AC55FF10FF4DFEACABFB" box="[224,259,222,248]" pageId="82" pageNumber="83" type="female">1♀</specimenCount>
(
<figureCitation id="EBAE2A62F222AC55FEE3FF4DFEDBABFB" box="[275,372,222,248]" captionStart="FIGURE 123" captionStartId="212.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-26@212.[151,1435,181,1845]" captionTargetPageId="212" captionText="FIGURE 123. Pamphilius ussuriensis, female, Russia – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) sawsheath laterally, (h) claw." figureDoi="http://doi.org/10.5281/zenodo.6876799" httpUri="https://zenodo.org/record/6876799/files/figure.png" pageId="82" pageNumber="83">Fig. 123</figureCitation>
, DEI-GISHym 12972), Samarka, Gordeyevskaya Mtn.,
<quantity id="B46D9B02F222AC55FC1DFF4DFB9BABFB" box="[1005,1076,222,248]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="82" pageNumber="83" unit="km" value="70.0">70 km</quantity>
N Chuguyevka,
<quantity id="B46D9B02F222AC55FB03FF4DFA92ABFA" box="[1267,1341,222,249]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="82" pageNumber="83" unit="m" value="250.0">
<elevation id="F8B8D1D4F222AC55FB03FF4DFA92ABFA" box="[1267,1341,222,249]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="82" pageNumber="83" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<geoCoordinate id="16A15020F222AC55FAB9FF4DFA33ABFB" box="[1353,1436,222,248]" degrees="44.46" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="555" value="44.46">44.46N</geoCoordinate>
<geoCoordinate id="16A15020F222AC55FF67FE91FF57AA1F" box="[151,248,258,284]" degrees="134.13" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="555" value="134.13">134.13E</geoCoordinate>
,
<date id="072B1027F222AC55FEF6FE91FEEAAA1E" box="[262,325,258,285]" pageId="82" pageNumber="83" value="1993-05-29">
29.
<collectionCode id="1584AE22F222AC55FEC1FE90FEEAAA1E" box="[305,325,259,285]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="82" pageNumber="83" type="Museum">V</collectionCode>
</date>
. 1993, leg.
<collectorName id="DE605331F222AC55FE3BFE91FD94AA1E" box="[459,571,258,285]" pageId="82" pageNumber="83">A. Taeger</collectorName>
(
<collectionCode id="1584AE22F222AC55FDBDFE91FD3EAA1F" box="[589,657,258,284]" pageId="82" pageNumber="83">SDEI</collectionCode>
);
</materialsCitation>
<materialsCitation id="C3FD3CBAF222AC55FD56FE91FC67AA43" ID-GBIF-Occurrence="3866537332" collectionCode="V" collectorName="A. Taeger" elevation="250" latitude="44.46" location="Sikhote-Alin Reserve" longLatPrecision="682" longitude="134.13" municipality="Upper River Dzhigitovka" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="6593FD6EF222AC55FD56FE91FD66AA1F" box="[678,713,258,284]" pageId="82" pageNumber="83" type="male">1♂</specimenCount>
(
<figureCitation id="EBAE2A62F222AC55FD2BFE91FC91AA1F" box="[731,830,258,284]" captionStart="FIGURE 124" captionStartId="213.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-19@213.[151,1435,181,1845]" captionTargetPageId="213" captionText="FIGURE 124. Pamphilius ussuriensis, male, Russia – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head &amp; thorax ventrolaterally, (e) face, (f) antenna, (g) genitalia dorsally, (h) genitalia ventrally, (i) claw." figureDoi="http://doi.org/10.5281/zenodo.6876801" httpUri="https://zenodo.org/record/6876801/files/figure.png" pageId="82" pageNumber="83">Fig. 124</figureCitation>
, DEI-GISHym 21853),
<location id="764A603CF222AC55FBA2FE91FAE6AA1E" LSID="urn:lsid:plazi:treatment:FB3C87F1F221AC55FF67F938FAE0A85E:764A603CF222AC55FBA2FE91FAE6AA1E" box="[1106,1353,258,285]" municipality="Upper River Dzhigitovka" name="Sikhote-Alin Reserve" pageId="82" pageNumber="83">Sikhote-Alin Reserve</location>
,
<collectingMunicipality id="934EAC9DF222AC55FAA6FE90FEC3AA43" pageId="82" pageNumber="83">Upper River Dzhigitovka</collectingMunicipality>
,
<geoCoordinate id="16A15020F222AC55FE8AFEB5FE49AA43" box="[378,486,294,320]" degrees="44.970" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="55" value="44.97">44.970°N</geoCoordinate>
<geoCoordinate id="16A15020F222AC55FE00FEB5FDC5AA43" box="[496,618,294,320]" degrees="136.080" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="55" value="136.08">136.080°E</geoCoordinate>
,
<date id="072B1027F222AC55FD87FEB5FD18AA42" box="[631,695,294,321]" pageId="82" pageNumber="83" value="2015-05-27">
27.
<collectionCode id="1584AE22F222AC55FD53FEB4FD18AA42" box="[675,695,295,321]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="82" pageNumber="83" type="Museum">V</collectionCode>
</date>
. 2015,
<collectorName id="DE605331F222AC55FCFDFEB5FCC8AA43" box="[781,871,294,320]" pageId="82" pageNumber="83">Sergeev</collectorName>
(
<collectionCode id="1584AE22F222AC55FC88FEB5FC13AA43" box="[888,956,294,320]" pageId="82" pageNumber="83">SDEI</collectionCode>
);
</materialsCitation>
<materialsCitation id="C3FD3CBAF222AC55FC22FEB5FAD5AA67" ID-GBIF-Occurrence="3866537338" collectionCode="V" collectorName="A. Taeger" elevation="250" latitude="44.46" location="Sikhote-Alin Reserve" longLatPrecision="682" longitude="134.13" municipality="Upper River Dzhigitovka" pageId="82" pageNumber="83" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="6593FD6EF222AC55FC22FEB5FC5AAA43" box="[978,1013,294,320]" pageId="82" pageNumber="83" type="female">1♀</specimenCount>
(
<figureCitation id="EBAE2A62F222AC55FBF7FEB5FB3EAA43" box="[1031,1169,294,320]" captionStart="FIGURE 126" captionStartId="215.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-19@215.[151,1435,181,1845]" captionTargetPageId="215" captionText="FIGURE 126. Pamphilius ussuriensis (specimen 86348 and holotype of P. zinovjevi), female, Russia – (a) head &amp; thorax dorsally (86348), (b) head &amp; thorax ventrolaterally (86348), (c) face (86348), (d) dorsally, (e) lateroventrally, (f) face." figureDoi="http://doi.org/10.5281/zenodo.6876805" httpUri="https://zenodo.org/record/6876805/files/figure.png" pageId="82" pageNumber="83">Fig. 126a–c</figureCitation>
, DEI-GISHym 86348),
<location id="764A603CF222AC55FF67FED9FEBDAA67" LSID="urn:lsid:plazi:treatment:FB3C87F1F221AC55FF67F938FAE0A85E:764A603CF222AC55FF67FED9FEBDAA67" box="[151,274,330,356]" name="Rettihovka" pageId="82" pageNumber="83">Rettihovka</location>
<quantity id="B46D9B02F222AC55FEE9FED9FEFDAA67" box="[281,338,330,356]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="82" pageNumber="83" unit="km" value="9.0">9 km</quantity>
E,
<quantity id="B46D9B02F222AC55FE88FED9FE14AA67" box="[376,443,330,356]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="82" pageNumber="83" unit="m" value="170.0">
<elevation id="F8B8D1D4F222AC55FE88FED9FE14AA67" box="[376,443,330,356]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="82" pageNumber="83" unit="m" value="170.0">170m</elevation>
</quantity>
,
<geoCoordinate id="16A15020F222AC55FE36FED9FD9CAA67" box="[454,563,330,356]" degrees="44.178" direction="north" orientation="latitude" pageId="82" pageNumber="83" precision="55" value="44.178">44.178°N</geoCoordinate>
<geoCoordinate id="16A15020F222AC55FDC9FED9FD1CAA67" box="[569,691,330,356]" degrees="132.877" direction="east" orientation="longitude" pageId="82" pageNumber="83" precision="55" value="132.877">132.877°E</geoCoordinate>
,
<date id="072B1027F222AC55FD4EFED9FD53AA66" box="[702,764,330,357]" pageId="82" pageNumber="83" value="2016-05-29">
29.
<collectionCode id="1584AE22F222AC55FD18FED8FD53AA66" box="[744,764,331,357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="82" pageNumber="83" type="Museum">V</collectionCode>
</date>
. 2016, K.
<collectorName id="DE605331F222AC55FC9FFED8FC6FAA66" box="[879,960,331,357]" pageId="82" pageNumber="83">Kramp</collectorName>
,
<collectorName id="DE605331F222AC55FC3AFED8FB9EAA66" box="[970,1073,331,357]" pageId="82" pageNumber="83">M. Prous</collectorName>
&amp;
<collectorName id="DE605331F222AC55FBA3FED9FB6CAA66" box="[1107,1219,330,357]" pageId="82" pageNumber="83">A. Taeger</collectorName>
, RU024 (
<collectionCode id="1584AE22F222AC55FADEFED9FADDAA67" box="[1326,1394,330,356]" pageId="82" pageNumber="83">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF222AC55FF37FEFDFC54AA8A" box="[199,1019,366,393]" pageId="82" pageNumber="83" type="distribution">
<paragraph id="732A36E7F222AC55FF37FEFDFC54AA8A" blockId="82.[151,1437,150,861]" box="[199,1019,366,393]" pageId="82" pageNumber="83">
<emphasis id="41E1EAF5F222AC55FF37FEFDFEF6AA8B" bold="true" box="[199,345,366,392]" pageId="82" pageNumber="83">Distribution</emphasis>
.
<collectingCountry id="0B827677F222AC55FE97FEFDFE1DAA8B" box="[359,434,366,392]" name="Russia" pageId="82" pageNumber="83">Russia</collectingCountry>
(Amurskaja Oblast',
<collectingRegion id="B151F805F222AC55FD50FEFDFCFBAA8B" box="[672,852,366,392]" country="Russia" name="Primorskiy" pageId="82" pageNumber="83">Primorskij Kraj</collectingRegion>
),
<collectingCountry id="0B827677F222AC55FC97FEFDFC58AA8A" box="[871,1015,366,393]" name="South Korea" pageId="82" pageNumber="83">South Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF222AC55FF37FE01FE66AAAF" box="[199,457,402,429]" pageId="82" pageNumber="83" type="biology_ecology">
<paragraph id="732A36E7F222AC55FF37FE01FE66AAAF" blockId="82.[151,1437,150,861]" box="[199,457,402,429]" pageId="82" pageNumber="83">
<emphasis id="41E1EAF5F222AC55FF37FE01FEEBAAAF" bold="true" box="[199,324,402,428]" pageId="82" pageNumber="83">Host plant</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B8F656CF222AC55FF37FE25FAE0A85E" pageId="82" pageNumber="83" type="discussion">
<paragraph id="732A36E7F222AC55FF37FE25FD1CA9F2" blockId="82.[151,1437,150,861]" pageId="82" pageNumber="83">
<emphasis id="41E1EAF5F222AC55FF37FE25FE9BAAD3" bold="true" box="[199,308,438,464]" pageId="82" pageNumber="83">Remarks</emphasis>
.
<taxonomicName id="B4954D64F222AC55FEB2FE25FDEAAAD3" authorityName="Shinohara" authorityYear="1988" box="[322,581,438,464]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FEB2FE25FDEAAAD3" box="[322,581,438,464]" italics="true" pageId="82" pageNumber="83">Pamphilius ussuriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F222AC55FD8DFE24FD52AAD3" authorityName="Shinohara" authorityYear="1988" box="[637,765,439,464]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FD8DFE24FD52AAD3" box="[637,765,439,464]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
were described from the
<typeStatus id="AC2E8845F222AC55FBEBFE25FB27AAD3" box="[1051,1160,438,464]" pageId="82" pageNumber="83" type="holotype">holotypes</typeStatus>
only. The
<typeStatus id="AC2E8845F222AC55FAF0FE25FACDAAD3" box="[1280,1378,438,464]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B4954D64F222AC55FA78FE24FEBCAAF7" authorityName="Shinohara" authorityYear="1988" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FA78FE24FEBCAAF7" italics="true" pageId="82" pageNumber="83">P. ussuriensis</emphasis>
</taxonomicName>
is a rather poorly preserved (discoloured) male specimen (
<figureCitation id="EBAE2A62F222AC55FC55FE49FBABAAF6" box="[933,1028,474,501]" captionStart="FIGURE 125" captionStartId="214.[151,250,1748,1772]" captionTargetBox="[151,1435,181,1722]" captionTargetId="figure-26@214.[151,1435,181,1724]" captionTargetPageId="214" captionText="FIGURE 125. Pamphilius ussuriensis, male, Russia (holotype) – (a) dorsally, (b) lateroventrally, (c) head &amp; thorax dorsally, (d) head antenna dorsolaterally, (e) face." figureDoi="http://doi.org/10.5281/zenodo.6876803" httpUri="https://zenodo.org/record/6876803/files/figure.png" pageId="82" pageNumber="83">Fig. 125</figureCitation>
) and that of
<taxonomicName id="B4954D64F222AC55FB61FE48FABEAAF7" authorityName="Shinohara" authorityYear="1988" box="[1169,1297,475,500]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FB61FE48FABEAAF7" box="[1169,1297,475,500]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
is an exceptionally small female specimen (
<figureCitation id="EBAE2A62F222AC55FDF0FE6DFD29A91B" box="[512,646,510,536]" captionStart="FIGURE 126" captionStartId="215.[151,250,1869,1893]" captionTargetBox="[151,1435,181,1845]" captionTargetId="figure-19@215.[151,1435,181,1845]" captionTargetPageId="215" captionText="FIGURE 126. Pamphilius ussuriensis (specimen 86348 and holotype of P. zinovjevi), female, Russia – (a) head &amp; thorax dorsally (86348), (b) head &amp; thorax ventrolaterally (86348), (c) face (86348), (d) dorsally, (e) lateroventrally, (f) face." figureDoi="http://doi.org/10.5281/zenodo.6876805" httpUri="https://zenodo.org/record/6876805/files/figure.png" pageId="82" pageNumber="83">Fig. 126d–f</figureCitation>
;
<bibRefCitation id="17044B16F222AC55FD60FE6DFCFFA91B" author="Shinohara, A." box="[656,848,510,536]" pageId="82" pageNumber="83" pagination="307 - 320" refId="ref76348" refString="Shinohara, A. (1988 a) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Five new species and additional records from the Far East. Kontyu, Tokyo, 56, 307 - 320." type="journal article" year="1988">Shinohara 1988a</bibRefCitation>
).
<bibRefCitation id="17044B16F222AC55FC93FE6DFB8AA91B" author="Shinohara, A." box="[867,1061,510,536]" pageId="82" pageNumber="83" pagination="99 - 116" refId="ref76721" refString="Shinohara, A. (2001) The group of Pamphilius sylvaticus (Hymenoptera, Pamphiliidae): Two new species, new collection records, and a key to Palearctic species. Japanese Journal of Systematic Entomology, 7, 99 - 116." type="journal article" year="2001">Shinohara (2001)</bibRefCitation>
examined five additional females of
<taxonomicName id="B4954D64F222AC55FF46FDB0FE98A93F" authorityName="Shinohara" authorityYear="1988" box="[182,311,547,572]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FF46FDB0FE98A93F" box="[182,311,547,572]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
from Primorskij Kraj and
<collectingCountry id="0B827677F222AC55FD95FDB1FD5DA93E" box="[613,754,546,573]" name="South Korea" pageId="82" pageNumber="83">South Korea</collectingCountry>
and discussed the variation among them.
<taxonomicName id="B4954D64F222AC55FB3FFDB1FF78A963" authorityName="Shinohara" authorityYear="1988" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FB3FFDB1FF78A963" italics="true" pageId="82" pageNumber="83">Pamphilius ussuriensis</emphasis>
</taxonomicName>
was only known from the
<typeStatus id="AC2E8845F222AC55FE0FFDD5FDCEA963" box="[511,609,582,608]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
male until now. The COI sequences of the newly obtained specimens of
<taxonomicName id="B4954D64F222AC55FA78FDD4FEBCA987" authorityName="Shinohara" authorityYear="1988" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FA78FDD4FEBCA987" italics="true" pageId="82" pageNumber="83">P. ussuriensis</emphasis>
</taxonomicName>
(a male, DEI-GISHym 21853, and a female, DEI-GISHym 86348, initially determined as
<taxonomicName id="B4954D64F222AC55FAE3FDF8FA3CA987" authorityName="Shinohara" authorityYear="1988" box="[1299,1427,619,644]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FAE3FDF8FA3CA987" box="[1299,1427,619,644]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
) were retrieved as monophyletic with 100% UFBoot support (
<figureCitation id="EBAE2A62F222AC55FCABFD1DFC6FA9AB" box="[859,960,654,680]" captionStart="FIGURE 146" captionStartId="233.[151,250,1448,1472]" captionTargetBox="[157,1428,188,1418]" captionTargetId="figure-19@233.[151,1436,181,1424]" captionTargetPageId="233" captionText="FIGURE 146. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius sylvaticus group, part 2." figureDoi="http://doi.org/10.5281/zenodo.6876853" httpUri="https://zenodo.org/record/6876853/files/figure.png" pageId="82" pageNumber="83">Fig. 146</figureCitation>
). Though the
<specimenCount id="6593FD6EF222AC55FB93FD1CFAA4A9AB" box="[1123,1291,654,681]" pageId="82" pageNumber="83" type="generic">two specimens</specimenCount>
diverged by 1.8% in COI, it now seems reasonable to regard them as different sexes of the same species and here we propose to treat
<taxonomicName id="B4954D64F222AC55FF20FD44FEE0A9F3" authorityName="Shinohara" authorityYear="1988" box="[208,335,727,752]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="zinovjevi">
<emphasis id="41E1EAF5F222AC55FF20FD44FEE0A9F3" box="[208,335,727,752]" italics="true" pageId="82" pageNumber="83">P. zinovjevi</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName id="B4954D64F222AC55FDE4FD44FD04A9F3" authorityName="Shinohara" authorityYear="1988" box="[532,683,727,752]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FDE4FD44FD04A9F3" box="[532,683,727,752]" italics="true" pageId="82" pageNumber="83">P. ussuriensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="732A36E7F222AC55FF37FD69FAE0A85E" blockId="82.[151,1437,150,861]" pageId="82" pageNumber="83">
This species belongs to the
<taxonomicName id="B4954D64F222AC55FE07FD68FD2FA817" baseAuthorityName="Linne" baseAuthorityYear="1758" box="[503,640,762,788]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="sylvaticus">
<emphasis id="41E1EAF5F222AC55FE07FD68FD2FA817" box="[503,640,762,788]" italics="true" pageId="82" pageNumber="83">P. sylvaticus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="17044B16F222AC55FD25FD69FC36A817" author="Shinohara, A." box="[725,921,762,788]" pageId="82" pageNumber="83" pagination="359 - 438" refId="ref76809" refString="Shinohara, A. (2002 b) Systematics of the leaf-rolling or webspinning sawfly subfamily Pamphiliinae: a preliminary overview. In: Viitasaari, M. (Ed.), Sawflies (Hymenoptera, Symphyta) I. Tremex Press, Helsinki, pp. 359 - 438." type="book chapter" year="2002">Shinohara 2002b</bibRefCitation>
). Diverging by a minimum of 4.8% in the COI sequences, the nearest neighbour was
<taxonomicName id="B4954D64F222AC55FDCCFC8CFD7AA83B" authorityName="Shinohara &amp; Kramp &amp; Taeger" authorityYear="2022" box="[572,725,798,824]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="graciloides">
<emphasis id="41E1EAF5F222AC55FDCCFC8CFD7AA83B" box="[572,725,798,824]" italics="true" pageId="82" pageNumber="83">P. graciloides</emphasis>
</taxonomicName>
, according to our molecular analysis. In the COI tree (
<figureCitation id="EBAE2A62F222AC55FADFFC8DFA20A83B" box="[1327,1423,798,824]" captionStart="FIGURE 146" captionStartId="233.[151,250,1448,1472]" captionTargetBox="[157,1428,188,1418]" captionTargetId="figure-19@233.[151,1436,181,1424]" captionTargetPageId="233" captionText="FIGURE 146. Condensed ML tree based on CO1 gene sequences, with details of Pamphilius sylvaticus group, part 2." figureDoi="http://doi.org/10.5281/zenodo.6876853" httpUri="https://zenodo.org/record/6876853/files/figure.png" pageId="82" pageNumber="83">Fig. 146</figureCitation>
),
<taxonomicName id="B4954D64F222AC55FF67FCD0FE80A85F" authorityName="Shinohara" authorityYear="1988" box="[151,303,835,860]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="ussuriensis">
<emphasis id="41E1EAF5F222AC55FF67FCD0FE80A85F" box="[151,303,835,860]" italics="true" pageId="82" pageNumber="83">P. ussuriensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B4954D64F222AC55FE96FCD0FD6BA85F" authority="(Eversmann, 1847)" baseAuthorityName="Eversmann" baseAuthorityYear="1847" box="[358,708,834,860]" class="Insecta" family="Pamphiliidae" genus="Pamphilius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="jucundus">
<emphasis id="41E1EAF5F222AC55FE96FCD0FE49A85F" box="[358,486,834,860]" italics="true" pageId="82" pageNumber="83">P. jucundus</emphasis>
(
<bibRefCitation id="17044B16F222AC55FE05FCD1FD14A85F" author="Eversmann, E." box="[501,699,834,860]" pageId="82" pageNumber="83" pagination="3 - 68" refId="ref73264" refString="Eversmann, E. (1847) Fauna hymenopterologica volgo-uralensis exhibens Hymenopterorum species quas in provinciis Volgam fluvium inter et montes Uralenses sitis observavit et nunc descripsit. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 20, 3 - 68." type="journal article" year="1847">Eversmann, 1847</bibRefCitation>
)
</taxonomicName>
formed a monophyletic group with 99% UFBoot support.
</paragraph>
</subSubSection>
</treatment>
</document>
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