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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.86.75801" ID-GBIF-Dataset="a816417f-204c-4403-9407-01cc6152be6f" ID-PMC="PMC8760234" ID-Pensoft-Pub="1314-4049-86-1" ID-Pensoft-UUID="A65492DEA8E55FC9B9050FC172C2748B" ID-PubMed="35095303" ModsDocID="1314-4049-86-1" checkinTime="1641658256192" checkinUser="pensoft" docAuthor="Dissanayake, Lakmali S., Marasinghe, Diana S., Samarakoon, Milan C., Maharachchikumbura, Sajeewa S. N., Mortimer, Peter E., Hyde, Kevin D., Kuo, Chang-Hsin &amp; Kang, Ji-Chuan" docDate="2022" docId="BCD7BA98E30956889ECD67D1321BA46A" docLanguage="en" docName="MycoKeys 86: 1-17" docOrigin="MycoKeys 86" docPubDate="2022-01-07" docSource="http://dx.doi.org/10.3897/mycokeys.86.75801" docTitle="Iodosphaeria thailandica L. S. Dissan., Marasinghe, &amp; K. D. Hyde, sp. nov." docType="treatment" docVersion="4" id="A65492DEA8E55FC9B9050FC172C2748B" lastPageNumber="1" masterDocId="A65492DEA8E55FC9B9050FC172C2748B" masterDocTitle="Three new species of Iodosphaeria (Xylariomycetidae): I. chiayiensis, I. jinghongensis and I. thailandica" masterLastPageNumber="17" masterPageNumber="1" pageNumber="1" updateTime="1668136780210" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Three new species of Iodosphaeria (Xylariomycetidae): I. chiayiensis, I. jinghongensis and I. thailandica</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dissanayake, Lakmali S.</mods:namePart>
<mods:affiliation>Engineering and Research Center for Southwest Bio-Pharmaceutical Resources of National Education Ministry of China, Guizhou University, Guiyang 550025, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marasinghe, Diana S.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Samarakoon, Milan C.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maharachchikumbura, Sajeewa S. N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9127-0783</mods:nameIdentifier>
<mods:affiliation>School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu 611731, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mortimer, Peter E.</mods:namePart>
<mods:affiliation>Centre for Mountain Futures (CMF), Kunming Institute of Botany, Honghe County 654400, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hyde, Kevin D.</mods:namePart>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Innovative Institute for Plant Health, Zhongkai University of Agriculture and Engineering, Guangzhou 510225, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kuo, Chang-Hsin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9011-6530</mods:nameIdentifier>
<mods:affiliation>Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kang, Ji-Chuan</mods:namePart>
<mods:affiliation>Engineering and Research Center for Southwest Bio-Pharmaceutical Resources of National Education Ministry of China, Guizhou University, Guiyang 550025, China</mods:affiliation>
<mods:nameIdentifier type="email">jckang@gzu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-01-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>86</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>17</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.86.75801</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.86.75801</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-86-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">A65492DEA8E55FC9B9050FC172C2748B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="191943042" LSID="urn:lsid:plazi:treatment:BCD7BA98E30956889ECD67D1321BA46A" httpUri="http://treatment.plazi.org/id/BCD7BA98E30956889ECD67D1321BA46A" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="BCD7BA98-E309-5688-9ECD-67D1321BA46A" authority="L. S. Dissan., Marasinghe, &amp; K. D. Hyde, sp. nov." authorityName="L. S. Dissan., Marasinghe, &amp; K. D. Hyde, sp. nov." class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">Iodosphaeria thailandica L.S. Dissan., Marasinghe, &amp; K.D. Hyde, sp. nov.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Iodosphaeria thailandica (MFLU 21 - 0041, holotype) a substrate b, c ascomata on the host surface d peridium e section of ascomata f appearance of setae (black arrow) on peridium g setae h paraphyses i, j asci k J + apical ring (in Melzer's reagent) l-q ascospores (p, q stained in Lactophenol Cotton Blue). Scale bars: 10 μm (d); 100 μm (e); 5 μm (f-h); 20 μm (i, j); 10 μm (k-q)." figureDoi="10.3897/mycokeys.86.75801.figure5" httpUri="https://binary.pensoft.net/fig/631572" pageId="0" pageNumber="1">Figure 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria thailandica</emphasis>
</taxonomicName>
refers to the country where the fungus was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype">
<paragraph pageId="0" pageNumber="1">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="1">MFLU 21-0041</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Saprobic</emphasis>
on dead leaves of unidentified host.
<emphasis bold="true" pageId="0" pageNumber="1">Sexual morph</emphasis>
:
<emphasis italics="true" pageId="0" pageNumber="1">Ascomata</emphasis>
250-285
<normalizedToken originalValue="×">x</normalizedToken>
250-295
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 267.3
<normalizedToken originalValue="×">x</normalizedToken>
272
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 10), globose to subglobose, superficial, black, solitary to gregarious, consisting of numerous long, flexuous setae.
<emphasis italics="true" pageId="0" pageNumber="1">Setae</emphasis>
4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, arising from cells at the peridium surface, dark brown to brown, unbranched, septate.
<emphasis italics="true" pageId="0" pageNumber="1">Ostiole</emphasis>
periphysate, apapillate.
<emphasis italics="true" pageId="0" pageNumber="1">Peridium</emphasis>
40-50
<normalizedToken originalValue="μm">μm</normalizedToken>
wide (x̅ = 44.6
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 10), comprising two layers of cells of
<emphasis italics="true" pageId="0" pageNumber="1">textura angularis</emphasis>
, outer layer of dark brown to black thick-walled cells and an inner layer of flattened, hyaline cells.
<emphasis italics="true" pageId="0" pageNumber="1">Paraphyses</emphasis>
5-8
<normalizedToken originalValue="μm">μm</normalizedToken>
wide, length as longer than asci, septate, hyaline, branched, embedded in a gelatinous matrix.
<emphasis italics="true" pageId="0" pageNumber="1">Asci</emphasis>
65-100
<normalizedToken originalValue="×">x</normalizedToken>
8-10
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 84.3
<normalizedToken originalValue="×">x</normalizedToken>
8.9
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 30), 8-spored, unitunicate, cylindrical, short pedicellate, apex rounded, with a J+ apical ring.
<emphasis italics="true" pageId="0" pageNumber="1">Ascospores</emphasis>
20-35
<normalizedToken originalValue="×">x</normalizedToken>
2-4
<normalizedToken originalValue="μm">μm</normalizedToken>
(x̅ = 29.1
<normalizedToken originalValue="×">x</normalizedToken>
3.2
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 30), overlapping uni-seriate, cylindrical to allantoid, aseptate, hyaline to pale brown, guttulate, slightly curved.
<emphasis bold="true" pageId="0" pageNumber="1">Asexual morph</emphasis>
: Undetermined.
</paragraph>
<caption doi="10.3897/mycokeys.86.75801.figure5" httpUri="https://binary.pensoft.net/fig/631572" pageId="0" pageNumber="1" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 5.</emphasis>
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria thailandica</emphasis>
</taxonomicName>
(MFLU 21-0041, holotype)
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
substrate
<emphasis bold="true" pageId="0" pageNumber="1">b, c</emphasis>
ascomata on the host surface
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
peridium
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
section of ascomata
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
appearance of setae (black arrow) on peridium
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
setae
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
paraphyses
<emphasis bold="true" pageId="0" pageNumber="1">i, j</emphasis>
asci
<emphasis bold="true" pageId="0" pageNumber="1">k</emphasis>
J+ apical ring (in
<normalizedToken originalValue="Melzers">Melzer's</normalizedToken>
reagent)
<emphasis bold="true" pageId="0" pageNumber="1">l-q</emphasis>
ascospores (
<emphasis bold="true" pageId="0" pageNumber="1">p, q</emphasis>
stained in Lactophenol Cotton Blue). Scale bars: 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
); 100
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
); 5
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">f-h</emphasis>
); 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">i, j</emphasis>
); 10
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="1">k-q</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2020-09-11" collectorName="D. S Marasinghe" country="Thailand" location="Mushroom Research Centre" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<collectingCountry name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:BCD7BA98E30956889ECD67D1321BA46A:17E404A948A225685E9E625B07340445" country="Thailand" name="Mushroom Research Centre" stateProvince="Chiang Mai">Mushroom Research Centre</location>
, on dead leaves of an undetermined species,
<collectingDate value="2020-09-11">11 September 2020</collectingDate>
,
<collectorName>D.S Marasinghe</collectorName>
, DMRC011 (MFLU 21-0041,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>holotype</typeStatus>
</emphasis>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria thailandica</emphasis>
</taxonomicName>
shares similar characteristics with
<taxonomicName lsidName="I. honghensis" pageId="0" pageNumber="1" rank="species" species="honghensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. honghensis</emphasis>
</taxonomicName>
in having globose to subglobose, superficial, solitary to gregarious ascomata, cylindrical, short pedicellate, J+, apical ring and cylindrical to allantoid asci with aseptate, guttulate ascospores (Marasinghe et al. 2019). However,
<taxonomicName lsidName="I. thailandica" pageId="0" pageNumber="1" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">I. thailandica</emphasis>
</taxonomicName>
differs from
<taxonomicName lsidName="I. honghensis" pageId="0" pageNumber="1" rank="species" species="honghensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. honghensis</emphasis>
</taxonomicName>
in having long, narrow (20-35
<normalizedToken originalValue="×">x</normalizedToken>
2-4
<normalizedToken originalValue="μm">μm</normalizedToken>
) and hyaline to pale brown ascospores versus short, broad (18.5-22.5
<normalizedToken originalValue="×">x</normalizedToken>
4.5-6.5
<normalizedToken originalValue="μm">μm</normalizedToken>
) and hyaline ascospores. In the phylogenetic analyses,
<taxonomicName lsidName="I. thailandica" pageId="0" pageNumber="1" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">I. thailandica</emphasis>
</taxonomicName>
is distinct from other species in the genus by 100 % ML and 1.00BYPP and sister to the
<taxonomicName lsidName="I. chiayiensis" pageId="0" pageNumber="1" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. chiayiensis</emphasis>
</taxonomicName>
.
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria thailandica</emphasis>
</taxonomicName>
has larger ascomata (250-285
<normalizedToken originalValue="×">x</normalizedToken>
250-295
<normalizedToken originalValue="μm">μm</normalizedToken>
), cylindrical to allantoid asci and hyaline to pale brown ascospores, while the ascomata of
<taxonomicName lsidName="I. chiayiensis" pageId="0" pageNumber="1" rank="species" species="chiayiensis">
<emphasis italics="true" pageId="0" pageNumber="1">I. chiayiensis</emphasis>
</taxonomicName>
are smaller (150-190
<normalizedToken originalValue="×">x</normalizedToken>
160-200
<normalizedToken originalValue="μm">μm</normalizedToken>
) and ascospores are hyaline and ellipsoidal to allantoid.
<taxonomicName class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria thailandica" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="species" species="thailandica">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria thailandica</emphasis>
</taxonomicName>
is the first report of
<taxonomicName authorityName="G.J.Samuels, E.Muller &amp; O.Petrini" authorityYear="1987" class="Sordariomycetes" family="Iodosphaeriaceae" genus="Iodosphaeria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Iodosphaeria" order="Xylariales" pageId="0" pageNumber="1" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Iodosphaeria</emphasis>
</taxonomicName>
from Thailand.
</paragraph>
</subSubSection>
</treatment>
</document>