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476 lines
44 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-GBIF-Taxon="182223668" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" LSID="urn:lsid:zoobank.org:act:F59062DE-95C1-48B7-A947-CAFE2EA6F98C" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="B1E57655968261FDF43BFDDB169B96E9" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax laterobustus Liebherr 2018, sp. n." docType="treatment" docUuid="F59062DE-95C1-48B7-A947-CAFE2EA6F98C" docUuidSource="ZooBank" docVersion="3" id="77D7AEC9294554E5BBDCABA37901022A" lastPageId="16" lastPageNumber="17" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageId="15" pageNumber="16" updateTime="1643465225333" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-01-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>65</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
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<mods:identifier type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
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<mods:identifier type="Pensoft-UUID">77D7AEC9294554E5BBDCABA37901022A</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1166397</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182223668" LSID="urn:lsid:zoobank.org:act:F59062DE-95C1-48B7-A947-CAFE2EA6F98C" httpUri="http://treatment.plazi.org/id/B1E57655968261FDF43BFDDB169B96E9" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection pageId="15" pageNumber="16" type="nomenclature">
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<paragraph pageId="15" pageNumber="16">
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<pageBreakToken pageId="15" pageNumber="16" start="start">1</pageBreakToken>
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.
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<taxonomicName LSID="http://zoobank.org/F59062DE-95C1-48B7-A947-CAFE2EA6F98C" authority="Liebherr" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax laterobustus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="laterobustus" status="sp. n.">Mecyclothorax laterobustus Liebherr</taxonomicName>
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<taxonomicNameLabel pageId="15" pageNumber="16">sp. n.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paired left (above) and right (below) parameres of Mecyclothorax spp. (ectal view): A, M. blackburni; B, M. storeyi; C, M. amplipennis; D, M. montivagus; E, M. laterobustus. As the aedeagal assembly of M. storeyi males is inverted relative to other Mecyclothorax species (Moore 1984), the configuration of the upper, or right paramere of M. storeyi should be compared to the ventral, or left parameres of the other species, and vice versa." figureDoi="10.3897/dez.65.21000.figure5" httpUri="https://binary.pensoft.net/fig/179961" pageId="15" pageNumber="16">Figures 5E</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="15" pageNumber="16">, 9A</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F11" captionText="Figure 10. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. laterorobustus, right view, dorsal view in situ (Ningua); C, M. laterosinuatus, right view (Col d'Amieu); D, M. laterosinuatus, right view (Touho TV tower); E, M. laterosinuatus ring, dorsal view; F, M. laterorectus, right view (Mt. Panie); G-H, M. laterorectus, right vew, dorsal view in situ (Mandjelia); I-J, M. fleutiauxi, right view, dorsal view in situ (Mt. Do); K-L, M. fleutiauxi, right view, sac inverted and everted (Me Maoya); M, M. fleutiauxi, right view (Ningua); N-O, M. jeanneli, right view, dorsal view in situ (Mt. Humboldt, 1350 m); P, M. jeanneli, right view (Mt. Humboldt, 1300 m); Q, M. jeanneli, right view, sac everted (Mt. Dzumac); R, M. jeanneli, right view (Riviere Bleue). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure10" httpUri="https://binary.pensoft.net/fig/179966" pageId="15" pageNumber="16">, 10A-B</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F13" captionText="Figure 12. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli; F, M. plurisetosus; G, M. megalovatulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure12" httpUri="https://binary.pensoft.net/fig/179968" pageId="15" pageNumber="16">, 12A</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F14" captionText="Figure 13. Left female gonocoxa, ventral view, Mecyclothorax (Phacothorax) spp.: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli; F, M. plurisetosus; G, M. megalovatulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure13" httpUri="https://binary.pensoft.net/fig/179969" pageId="15" pageNumber="16">, 13A</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F15" captionText="Figure 14. Geographical distributions of Mecyclothorax (Phacothorax) spp." figureDoi="10.3897/dez.65.21000.figure14" httpUri="https://binary.pensoft.net/fig/179970" pageId="15" pageNumber="16">, 14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="16" type="diagnosis">
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<paragraph pageId="15" pageNumber="16">Diagnosis.</paragraph>
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<paragraph pageId="15" pageNumber="16">
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These beetles are very robust, with a broad pronotum and broadly convex elytra (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="15" pageNumber="16">9A</figureCitation>
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). The pronotal base is broad, with MPW/BPW = 1.30-1.32, and the hind angles are projected, with the pronotal lateral margin concave just before the angle. Only the posterior supraorbital seta is present, and the pronotum is glabrous. Elytral striae 1-5 are well developed basally, with striae 3-5 deeper than those of individuals of the similar appearing
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<taxonomicName genus="M." lsidName="M. laterosinuatus" pageId="15" pageNumber="16" rank="species" species="laterosinuatus">
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<emphasis italics="true" pageId="15" pageNumber="16">M. laterosinuatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. laterorectus" pageId="15" pageNumber="16" rank="species" species="laterorectus">
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<emphasis italics="true" pageId="15" pageNumber="16">M. laterorectus</emphasis>
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</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="15" pageNumber="16">9B-C</figureCitation>
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). Standardized body length 4.1-4.4 mm. Chaetotaxy -/+//-/-//+/2/+/+.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
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<paragraph pageId="15" pageNumber="16">Description</paragraph>
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<paragraph pageId="15" pageNumber="16">
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(n = 5). Head capsule broad, eyes moderately convex, ocular lobe meeting gena at very obtuse angle; 16 ommatidia along horizontal diameter of eye; ocular ratio 1.33-1.42, EyL/EyD = 2.34-3.0; frontal grooves deep, arcuately convergent and deepest just posterad clypeus, briefly and shallowly extended onto clypeus; mandibles moderately elongate, mandibular ratio 1.67; ligular anterior margin rounded to ligular seta, concave between setae, the two setae separated by two setal diameters; paraglossae thin, extended as far beyond ligular margin as their basal length to margin; antennae moderately elongate, antennomere 9 length 2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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maximal breadth; antennomere 3 glabrous except for apical ring of setae. Pronotum moderately constricted basally, with obtuse-rounded hind angles and lateral margins briefly sinuate anterad angles (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="15" pageNumber="16">9A</figureCitation>
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); MPW/BPW = 1.30-1.32, MPW/PL = 1.19-1.22; front angles protruded anteriorly, but anterior margin narrowly approaching head capsule, APW/BPW = 0.69-0.71; basal margin broadly convex, beaded from just mesad deep, longitudinal laterobasal depression to and around obtuse-rounded hind angle; median longitudinal impression deep and narrow on disc, broader and shallower approaching base, absent anterad very shallow anterior transverse impression; proepisternum separated from prosternum by a very shallow groove anteriorly, but with a deep, slightly punctate groove ventrally; prosternal process deeply and narrowly depressed between procoxae, that depression extended 1/3 distance toward anterior prothoracic margin. Elytra broadly ovoid, humeri broad, humeral angle obtusely rounded laterad pronotal hind angles; MEW/EL = 0.87-0.93; basal groove evenly arcuate from scutellum to humeral angles, with depression at bases of sutural and elytral striae 4-5; sutural stria deep throughout length, stria 2 obsolete though traceable, stria 3 somewhat deeper, striae 4-5 deep in basal half, striae 6-7 obsolete; only sutural stria and stria 8 evident apically, elytra appressed and conjoined apically, sutural intervals narrower and upraised at apex. Pterothoracic mesepisternal anterior furrow with 6-7 deep punctures; mesosternal-mesepisternal suture complete (as in Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Thoracic structures of Mecyclothorax spp. A-B. Lateral view of elytral epipleuron and meso-and metapleurites: A, M. laterorectus; B, M. fleutiauxi. C-D. Metathoracic flight wing: C, M. punctipennis, showing fully developed flight wing folded under elytron; D, M. lissus, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include: mms, mesepisternal-mesosternal suture; msp, mesepimeron; mst, mesepisternum; mtp, metepimeron; mtt, metepisternum." figureDoi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="15" pageNumber="16">3A</figureCitation>
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); metepisternum maximum width/lateral length = 1.0; metepisternal-metepimeral suture incomplete, shallower and incomplete laterally. Abdomen with deep crescent-shaped depression along suture between first and second ventrite, second ventrite depressed within crescent; suture between second and third ventrites reduced though traceable laterally; ventrites 2-6 with broad, shallow, linear plaques near lateral margin. Microsculpture of frons a shallow transverse mesh, sculpticells twice as broad as long, these mixed with isodiametric sculpticells on vertex; pronotal disc and base covered with elongate transverse mesh plus transverse lines, surface iridescent; elytral iridescent, disc with loose elongate transverse mesh, apex with very elongate transverse mesh, sculpticell breadth 3-4
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<normalizedToken originalValue="×">x</normalizedToken>
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length.
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</paragraph>
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<caption ID-Zenodo-Dep="3488485" doi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="15" pageNumber="16" start="Figure 3" startId="F3">
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<paragraph pageId="15" pageNumber="16">
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<emphasis bold="true" pageId="15" pageNumber="16">Figure 3.</emphasis>
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Thoracic structures of
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<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
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</taxonomicName>
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spp.
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<emphasis bold="true" pageId="15" pageNumber="16">A-B</emphasis>
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. Lateral view of elytral epipleuron and meso-and metapleurites:
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<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. laterorectus" pageId="15" pageNumber="16" rank="species" species="laterorectus">
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<emphasis italics="true" pageId="15" pageNumber="16">M. laterorectus</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">B</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. fleutiauxi" pageId="15" pageNumber="16" rank="species" species="fleutiauxi">
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<emphasis italics="true" pageId="15" pageNumber="16">M. fleutiauxi</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="15" pageNumber="16">C-D</emphasis>
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. Metathoracic flight wing:
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<emphasis bold="true" pageId="15" pageNumber="16">C</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. punctipennis" pageId="15" pageNumber="16" rank="species" species="punctipennis">
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<emphasis italics="true" pageId="15" pageNumber="16">M. punctipennis</emphasis>
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</taxonomicName>
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, showing fully developed flight wing folded under elytron;
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<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. lissus" pageId="15" pageNumber="16" rank="species" species="lissus">
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<emphasis italics="true" pageId="15" pageNumber="16">M. lissus</emphasis>
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</taxonomicName>
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, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include:
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<emphasis bold="true" pageId="15" pageNumber="16">mms</emphasis>
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, mesepisternal-mesosternal suture;
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<emphasis bold="true" pageId="15" pageNumber="16">msp</emphasis>
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, mesepimeron;
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<emphasis bold="true" pageId="15" pageNumber="16">mst</emphasis>
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, mesepisternum;
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<emphasis bold="true" pageId="15" pageNumber="16">mtp</emphasis>
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, metepimeron;
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<emphasis bold="true" pageId="15" pageNumber="16">mtt</emphasis>
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, metepisternum.
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="3488487" doi="10.3897/dez.65.21000.figure4" httpUri="https://binary.pensoft.net/fig/179960" pageId="15" pageNumber="16" start="Figure 4" startId="F4">
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<paragraph pageId="15" pageNumber="16">
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<emphasis bold="true" pageId="15" pageNumber="16">Figure 4.</emphasis>
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Male aedeagal median lobe and parameres of
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<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
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</taxonomicName>
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spp., right lateral view, internal sac everted in
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<emphasis bold="true" pageId="15" pageNumber="16">C-G</emphasis>
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:
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<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. blackburni" pageId="15" pageNumber="16" rank="species" species="blackburni">
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<emphasis italics="true" pageId="15" pageNumber="16">M. blackburni</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">B-C</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. peryphoides" pageId="15" pageNumber="16" rank="species" species="peryphoides">
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<emphasis italics="true" pageId="15" pageNumber="16">M. peryphoides</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. moorei" pageId="15" pageNumber="16" rank="species" species="moorei">
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<emphasis italics="true" pageId="15" pageNumber="16">M. moorei</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">E</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. lewisensis" pageId="15" pageNumber="16" rank="species" species="lewisensis">
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<emphasis italics="true" pageId="15" pageNumber="16">M. lewisensis</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">F</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. punctipennis" pageId="15" pageNumber="16" rank="species" species="punctipennis">
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<emphasis italics="true" pageId="15" pageNumber="16">M. punctipennis</emphasis>
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</taxonomicName>
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;
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<emphasis bold="true" pageId="15" pageNumber="16">G</emphasis>
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,
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<taxonomicName genus="M." lsidName="M. montivagus" pageId="15" pageNumber="16" rank="species" species="montivagus">
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<emphasis italics="true" pageId="15" pageNumber="16">M. montivagus</emphasis>
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</taxonomicName>
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. See Table
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<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key to abbreviations for structures of male genitalia, and female genitalia and reproductive tracts." httpUri="http://table.plazi.org/id/E2C2FA730E55AFD2D7E83E55AE47B542" pageId="15" pageNumber="16" tableUuid="E2C2FA730E55AFD2D7E83E55AE47B542">2</tableCitation>
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for abbreviations.
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</paragraph>
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</caption>
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<caption ID-Zenodo-Dep="3488489" doi="10.3897/dez.65.21000.figure5" httpUri="https://binary.pensoft.net/fig/179961" pageId="15" pageNumber="16" start="Figure 5" startId="F5">
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<paragraph pageId="15" pageNumber="16">
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<emphasis bold="true" pageId="15" pageNumber="16">Figure 5.</emphasis>
|
||
Paired left (above) and right (below) parameres of
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
spp. (ectal view):
|
||
<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. blackburni" pageId="15" pageNumber="16" rank="species" species="blackburni">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. blackburni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">B</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. storeyi" pageId="15" pageNumber="16" rank="species" species="storeyi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. storeyi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">C</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. amplipennis" pageId="15" pageNumber="16" rank="species" species="amplipennis">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. amplipennis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. montivagus" pageId="15" pageNumber="16" rank="species" species="montivagus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. montivagus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">E</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. laterobustus" pageId="15" pageNumber="16" rank="species" species="laterobustus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. laterobustus</emphasis>
|
||
</taxonomicName>
|
||
. As the aedeagal assembly of
|
||
<taxonomicName genus="M." lsidName="M. storeyi" pageId="15" pageNumber="16" rank="species" species="storeyi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. storeyi</emphasis>
|
||
</taxonomicName>
|
||
males is inverted relative to other
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" author="Moore, BP" journalOrPublisher="Journal of the Australian Entomological Society" pageId="52" pageNumber="53" pagination="161 - 166" refId="B77" refString="Moore, BP, 1984. Taxonomic notes on some Australasian Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae) and descriptions of new species. Journal of the Australian Entomological Society 23: 161 - 166, DOI: https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" title="Taxonomic notes on some Australasian Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae) and descriptions of new species." url="https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" volume="23" year="1984">Moore 1984</bibRefCitation>
|
||
), the configuration of the upper, or right paramere of
|
||
<taxonomicName genus="M." lsidName="M. storeyi" pageId="15" pageNumber="16" rank="species" species="storeyi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. storeyi</emphasis>
|
||
</taxonomicName>
|
||
should be compared to the ventral, or left parameres of the other species, and vice versa.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Zenodo-Dep="3488492" doi="10.3897/dez.65.21000.figure6" httpUri="https://binary.pensoft.net/fig/179962" pageId="15" pageNumber="16" start="Figure 6" startId="F6">
|
||
<paragraph pageId="15" pageNumber="16">
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Figure 6.</emphasis>
|
||
Female reproductive tract, gonocoxites and associated laterotergites,
|
||
<taxonomicName authorityName="Erichson" authorityYear="1842" class="Insecta" family="Carabidae" genus="Amblytelus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Amblytelus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Amblytelus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
spp:
|
||
<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
|
||
,
|
||
<taxonomicName genus="A." lsidName="A. matthewsi" pageId="15" pageNumber="16" rank="species" species="matthewsi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">A. matthewsi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">B</emphasis>
|
||
,
|
||
<taxonomicName genus="A." lsidName="A. brevis" pageId="15" pageNumber="16" rank="species" species="brevis">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">A. brevis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">C</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. blackburni" pageId="15" pageNumber="16" rank="species" species="blackburni">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. blackburni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. storeyi" pageId="15" pageNumber="16" rank="species" species="storeyi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. storeyi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">E</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. minutus" pageId="15" pageNumber="16" rank="species" species="minutus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. minutus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">F</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. rectangulus" pageId="15" pageNumber="16" rank="species" species="rectangulus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. rectangulus</emphasis>
|
||
</taxonomicName>
|
||
. See Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key to abbreviations for structures of male genitalia, and female genitalia and reproductive tracts." httpUri="http://table.plazi.org/id/E2C2FA730E55AFD2D7E83E55AE47B542" pageId="15" pageNumber="16" tableUuid="E2C2FA730E55AFD2D7E83E55AE47B542">2</tableCitation>
|
||
for abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Zenodo-Dep="3488495" doi="10.3897/dez.65.21000.figure7" httpUri="https://binary.pensoft.net/fig/179963" pageId="15" pageNumber="16" start="Figure 7" startId="F7">
|
||
<paragraph pageId="15" pageNumber="16">
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Figure 7.</emphasis>
|
||
Strict consensus cladogram of 16 equally most-parsimonious trees derived from cladistic analysis of 65 moriomorphine taxa, including the secondary outgroup
|
||
<taxonomicName baseAuthorityName="Sloane" baseAuthorityYear="1915" class="Insecta" family="Carabidae" genus="Neonomius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Neonomius laevicollis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="laevicollis">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Neonomius laevicollis</emphasis>
|
||
</taxonomicName>
|
||
, six primary outgroup taxa in
|
||
<taxonomicName authorityName="Erichson" authorityYear="1842" class="Insecta" family="Carabidae" genus="Amblytelus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Amblytelus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Amblytelus</emphasis>
|
||
</taxonomicName>
|
||
and allied genera, and 58 ingroup terminals; 57
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
species plus two genitalic forms of
|
||
<taxonomicName genus="M." lsidName="M. andersoni" pageId="15" pageNumber="16" rank="species" species="andersoni">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M. andersoni</emphasis>
|
||
</taxonomicName>
|
||
. Species terminals are labeled with species name and three-letter abbreviation of relevant generic or subgeneric name. Full listing of included taxa along with subgeneric assignments is presented in Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key to abbreviations for structures of male genitalia, and female genitalia and reproductive tracts." httpUri="http://table.plazi.org/id/E2C2FA730E55AFD2D7E83E55AE47B542" pageId="15" pageNumber="16" tableUuid="E2C2FA730E55AFD2D7E83E55AE47B542">2</tableCitation>
|
||
. Cladistic analysis is based on 137 characters (see character descriptions for nature of each character Areas occupied by the included taxa indicated by abbreviations following species epithets:
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Bo</emphasis>
|
||
, Borneo;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">EOZ</emphasis>
|
||
, eastern Australia, i.e. east of the Nullarbor Plain;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">FP</emphasis>
|
||
, French Polynesia, Tahiti;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">HI</emphasis>
|
||
, Hawaiian Islands, Maui;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Jv</emphasis>
|
||
, Java;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">LH</emphasis>
|
||
, Lord Howe Island;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">NC</emphasis>
|
||
, New Caledonia;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Nf</emphasis>
|
||
, Norfolk Island;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">NNZ</emphasis>
|
||
, North Island of New Zealand;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">NZ</emphasis>
|
||
, generally distributed across New Zealand;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">OZ</emphasis>
|
||
, generally distributed across Australia;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">PNG</emphasis>
|
||
, Papua New Guinea;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">QOZ</emphasis>
|
||
, restricted to Queensland, Australia;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">SNZ</emphasis>
|
||
, South Island of New Zealand plus Chatham Islands;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">SP&A</emphasis>
|
||
, St. Paul and Amsterdam Islands;
|
||
<emphasis bold="true" pageId="15" pageNumber="16">WOZ</emphasis>
|
||
, western Australia, i.e. west of the Nullarbor Plain.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-Zenodo-Dep="3488497" doi="10.3897/dez.65.21000.figure8" httpUri="https://binary.pensoft.net/fig/179964" pageId="15" pageNumber="16" start="Figure 8" startId="F9">
|
||
<paragraph pageId="15" pageNumber="16">
|
||
<emphasis bold="true" pageId="15" pageNumber="16">Figure 8.</emphasis>
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
spp., dorsal view, with subgeneric assignments:
|
||
<emphasis bold="true" pageId="15" pageNumber="16">A</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M." pageId="15" pageNumber="16" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M.</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Eucyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eucyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Eucyclothorax</emphasis>
|
||
</taxonomicName>
|
||
, subgen. n.)
|
||
<taxonomicName baseAuthorityName="Sloane" baseAuthorityYear="1898" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax blackburni" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="blackburni">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax blackburni</emphasis>
|
||
</taxonomicName>
|
||
(type species);
|
||
<emphasis bold="true" pageId="15" pageNumber="16">B</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M." pageId="15" pageNumber="16" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M.</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Qecyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Qecyclothorax" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Qecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
, subgen. n.)
|
||
<taxonomicName authorityName="Moore" authorityYear="1984" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax storeyi" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="storeyi">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Mecyclothorax storeyi</emphasis>
|
||
</taxonomicName>
|
||
(type species);
|
||
<emphasis bold="true" pageId="15" pageNumber="16">C</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M." pageId="15" pageNumber="16" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">M.</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName authorityName="Liebherr & Marris" authorityYear="2009" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meonochilus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Meonochilus</emphasis>
|
||
</taxonomicName>
|
||
, status n.)
|
||
<taxonomicName baseAuthorityName="Broun" baseAuthorityYear="1912" class="Insecta" family="Carabidae" genus="Meonochilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Meonochilus amplipennis" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="amplipennis">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Meonochilus amplipennis</emphasis>
|
||
</taxonomicName>
|
||
(type species);
|
||
<emphasis bold="true" pageId="15" pageNumber="16">D</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. (Mecyclothorax) montivagus" pageId="15" pageNumber="16" rank="species" species="montivagus" subGenus="Mecyclothorax">M. (Mecyclothorax) montivagus</taxonomicName>
|
||
(type species);
|
||
<emphasis bold="true" pageId="15" pageNumber="16">E</emphasis>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. (Mecyclothorax) lissus" pageId="15" pageNumber="16" rank="species" species="lissus" subGenus="Mecyclothorax">M. (Mecyclothorax) lissus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="15" pageNumber="16">
|
||
<emphasis italics="true" pageId="15" pageNumber="16">Male genitalia</emphasis>
|
||
(n = 1). Antecostal margin of male mediotergite IX distally angulate, little extended (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F11" captionText="Figure 10. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. laterorobustus, right view, dorsal view in situ (Ningua); C, M. laterosinuatus, right view (Col d'Amieu); D, M. laterosinuatus, right view (Touho TV tower); E, M. laterosinuatus ring, dorsal view; F, M. laterorectus, right view (Mt. Panie); G-H, M. laterorectus, right vew, dorsal view in situ (Mandjelia); I-J, M. fleutiauxi, right view, dorsal view in situ (Mt. Do); K-L, M. fleutiauxi, right view, sac inverted and everted (Me Maoya); M, M. fleutiauxi, right view (Ningua); N-O, M. jeanneli, right view, dorsal view in situ (Mt. Humboldt, 1350 m); P, M. jeanneli, right view (Mt. Humboldt, 1300 m); Q, M. jeanneli, right view, sac everted (Mt. Dzumac); R, M. jeanneli, right view (Riviere Bleue). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure10" httpUri="https://binary.pensoft.net/fig/179966" pageId="15" pageNumber="16">10B</figureCitation>
|
||
); right paramere very elongate, moderately narrow, with a single seta on dorsal margin distant from two apical setae (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paired left (above) and right (below) parameres of Mecyclothorax spp. (ectal view): A, M. blackburni; B, M. storeyi; C, M. amplipennis; D, M. montivagus; E, M. laterobustus. As the aedeagal assembly of M. storeyi males is inverted relative to other Mecyclothorax species (Moore 1984), the configuration of the upper, or right paramere of M. storeyi should be compared to the ventral, or left parameres of the other species, and vice versa." figureDoi="10.3897/dez.65.21000.figure5" httpUri="https://binary.pensoft.net/fig/179961" pageId="15" pageNumber="16">5E</figureCitation>
|
||
); left paramere narrow basally, evenly narrowed to a whip-shaped apex, two longer setae at apex; aedeagal median lobe shaft robust, broad dorsoventrally (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F11" captionText="Figure 10. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. laterorobustus, right view, dorsal view in situ (Ningua); C, M. laterosinuatus, right view (Col d'Amieu); D, M. laterosinuatus, right view (Touho TV tower); E, M. laterosinuatus ring, dorsal view; F, M. laterorectus, right view (Mt. Panie); G-H, M. laterorectus, right vew, dorsal view in situ (Mandjelia); I-J, M. fleutiauxi, right view, dorsal view in situ (Mt. Do); K-L, M. fleutiauxi, right view, sac inverted and everted (Me Maoya); M, M. fleutiauxi, right view (Ningua); N-O, M. jeanneli, right view, dorsal view in situ (Mt. Humboldt, 1350 m); P, M. jeanneli, right view (Mt. Humboldt, 1300 m); Q, M. jeanneli, right view, sac everted (Mt. Dzumac); R, M. jeanneli, right view (Riviere Bleue). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure10" httpUri="https://binary.pensoft.net/fig/179966" pageId="15" pageNumber="16">10A</figureCitation>
|
||
), median lobe apex broad, sides subparallel, with tip broadly rounded; aedeagal internal sac with flagellum, flagellar sheath, and sclerotized dorsal plate.
|
||
</paragraph>
|
||
<paragraph pageId="16" pageNumber="17">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">
|
||
<pageBreakToken pageId="16" pageNumber="17" start="start">Female</pageBreakToken>
|
||
reproductive tract
|
||
</emphasis>
|
||
(n = 1). Bursa copulatrix elongate, length more than twice circumference, surface translucent, membranous (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F13" captionText="Figure 12. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli; F, M. plurisetosus; G, M. megalovatulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure12" httpUri="https://binary.pensoft.net/fig/179968" pageId="16" pageNumber="17">12A</figureCitation>
|
||
); spermathecal duct entering near bursa-common oviduct juncture with duct oriented toward right side of bursa, duct longer than spermathecal reservoir; a rounded laminar helminthoid sclerite present at base of spermathecal duct; spermatheca fusiform on narrow duct, spermathecal gland duct entering at base of spermathecal reservoir; basal gonocoxite 1 with apical fringe of two setae near medial margin, a series of small setae lining medial margin (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F14" captionText="Figure 13. Left female gonocoxa, ventral view, Mecyclothorax (Phacothorax) spp.: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli; F, M. plurisetosus; G, M. megalovatulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure13" httpUri="https://binary.pensoft.net/fig/179969" pageId="16" pageNumber="17">13A</figureCitation>
|
||
); gonocoxite 2 moderately broad basally, basal width about half medial length; two gracile lateral ensiform setae of moderate length present.
|
||
</paragraph>
|
||
<paragraph pageId="16" pageNumber="17">
|
||
<emphasis italics="true" pageId="16" pageNumber="17">Types</emphasis>
|
||
- Holotype male (MNHN): NEW CALEDONIA / Ningua Res.camp / 12-13Nov 2001 / G.B.Monteith // QM Berlesate 1039 / 21°45
|
||
<normalizedToken originalValue="’Sx166°09’">'Sx
|
||
<geoCoordinate degrees="166" direction="east" minutes="09" orientation="longitude" precision="925" value="166.15">166°09'E</geoCoordinate>
|
||
</normalizedToken>
|
||
/ Rainforest, 1100m / Sieved litter // QUEENSLAND / MUSEUM LOAN / Date: Nov. 2003 / No. LEN-1686 (green label) // HOLOTYPE /
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax laterobustus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="laterobustus">Mecyclothorax laterobustus</taxonomicName>
|
||
/ J.K.Liebherr 2017 (black-bordered red label).
|
||
</paragraph>
|
||
<paragraph pageId="16" pageNumber="17">
|
||
Paratypes (10 specimens). NEW CALEDONIA: Ningua Reserve, camp, 1100 m el.,
|
||
<geoCoordinate degrees="21" direction="south" minutes="45" orientation="latitude" precision="925" value="-21.75">21°45'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="09" orientation="longitude" precision="925" value="166.15">166°09'E</geoCoordinate>
|
||
, rainforest, sieved litter, 12-13-xi-2001, lot 1039, Monteith (QMB, 4), near summit, rainforest, 1300 m el.,
|
||
<geoCoordinate degrees="21" direction="south" minutes="45" orientation="latitude" precision="925" value="-21.75">21°45'S</geoCoordinate>
|
||
<geoCoordinate degrees="166" direction="east" minutes="09" orientation="longitude" precision="925" value="166.15">166°09'E</geoCoordinate>
|
||
, berlesate, sieved litter, 13-xi-2001, lot 1052, Monteith (QMB, 3); Plateau de Dogny, rainforest, 910 m el.,
|
||
<geoCoordinate degrees="21" direction="south" minutes="37" orientation="latitude" precision="925" value="-21.616667">21°37'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="165" direction="east" minutes="53" orientation="longitude" precision="925" value="165.88333">165°53'E</geoCoordinate>
|
||
, berlesate sieved litter, 16-xi-2002, lot 1085, Burwell (QMB, 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph pageId="16" pageNumber="17">Etymology.</paragraph>
|
||
<paragraph pageId="16" pageNumber="17">
|
||
The very broad and robust body form of these beetles (Fig.
|
||
<figureCitation captionStart="Figure 9" captionStartId="F10" captionText="Figure 9. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. laterobustus; B, M. laterosinuatus; C, M. laterorectus; D, M. fleutiauxi; E, M. jeanneli." figureDoi="10.3897/dez.65.21000.figure9" httpUri="https://binary.pensoft.net/fig/179965" pageId="16" pageNumber="17">9A</figureCitation>
|
||
) suggests the compound, adjectival species epithet laterorobustus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph pageId="16" pageNumber="17">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="16" pageNumber="17">
|
||
This species is known from two localities, Ningua Reserve and Plateau de Dogny (Fig.
|
||
<figureCitation captionStart="Figure 14" captionStartId="F15" captionText="Figure 14. Geographical distributions of Mecyclothorax (Phacothorax) spp." figureDoi="10.3897/dez.65.21000.figure14" httpUri="https://binary.pensoft.net/fig/179970" pageId="16" pageNumber="17">14</figureCitation>
|
||
), and specimens have been collected from 900-1300 m elevation. The three collecting events are all based on recovering specimens from sieved litter, consistent with an interpretation that these beetles occupy ground-level microhabitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |