treatments-xml/data/A2/47/CE/A247CE056C4531A19AD3C413B564AADC.xml
2024-06-21 12:46:14 +02:00

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<mods:title id="5A38B916078BC5950F2D838DB39D8111">A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae).</mods:title>
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<mods:namePart id="2EFFE0ADAC1EC4B4A25626DC9C9EBA29">Eguchi, K.</mods:namePart>
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<treatment id="A247CE056C4531A19AD3C413B564AADC" ID-DOI="http://doi.org/10.5281/zenodo.6235628" ID-GBIF-Taxon="100122315" ID-Zenodo-Dep="6235628" LSID="urn:lsid:plazi:treatment:A247CE056C4531A19AD3C413B564AADC" httpUri="http://treatment.plazi.org/id/A247CE056C4531A19AD3C413B564AADC" lastPageNumber="27" pageNumber="24">
<subSubSection id="6FF0AE7D10E17962EBAB5B606BB8E7E5" type="nomenclature">
<paragraph id="9F8AD6D3A6A4D402ABA9580517A667F2" pageNumber="24">
<taxonomicName id="A9ACD226E7637D7795DBEEDA4E35C27C" ID-CoL="4FYC2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">
<pageStartToken id="DDD4674696089B04718888058F5FC761" pageNumber="24">Pheidole</pageStartToken>
fervens F. Smith
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D0902B80D91E08C792FE9D88F3E859C4" type="description">
<paragraph id="DA33AA259B39D3286B7290FD478E015E" pageNumber="24">Figs. 5a-g</paragraph>
</subSubSection>
<subSubSection id="F31FB5C30E30414E14F7AD8C4C8DB746" type="reference_group">
<paragraph id="7AA8B80FAE24957D078BCB0E2573B32E" pageNumber="24">
<taxonomicName id="B58515886168E86A642F4767224E3E0F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">Pheidole fervens F. Smith</taxonomicName>
, 1858: 176. Ogata 1982: 197 (description of male), Eguchi 2001b: 53-55 (redescription of major &amp; minor), Eguchi 2004: 197-198 (taxonomic remarks). Syntypes: 1 major &amp; 2 minors, &quot;SING 33&quot; [Singapore],
<collectionCode id="10DBF9C89230C489831D4B176E28CF50" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
, examined.
</paragraph>
<paragraph id="76D721915F1D6646B20B8DFA59DCF95B" pageNumber="24">
<taxonomicName id="41DC77843F99C6364F89113F413BEAA9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142286" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole cavannae Emery" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="cavannae">Pheidole cavannae Emery</taxonomicName>
, 1887: 464. Emery 1914: 401 (as subspecies of
<taxonomicName id="9B4B9894B908D241B069839447898840" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33906" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="178" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole oceanica Mayr" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="oceanica">P. oceanica</taxonomicName>
), Wilson &amp; Taylor 1967: 45 (junior synonym offervens). Syntype(s): major, New Caledonia, not examined.
</paragraph>
<paragraph id="52D7952D01CB7AB87C464482684966AA" pageNumber="24">
<taxonomicName id="7B738405AF1E78606E173C0099622657" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142287" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana Mayr" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="javana">Pheidole javana Mayr</taxonomicName>
, 1867: 98. Wilson &amp; Taylor 1967: 45 (junior synonym of
<taxonomicName id="5AD1BBB1CABDF078A5DBFB38FB95F87C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
), Eguchi 2004: 197 (lectotype designation). Lectotype: major: &quot;Batavia&quot; [Jakarta, Java],
<collectionCode id="7876C7DEED6B5BADF852DB8656485164" collectionName="Austria, Wien, Naturhistorisches Museum Wien">NHMW</collectionCode>
, examined; paralectotypes: 2 majors, same data as lectotype,
<collectionCode id="D9616F4B3B465501E9D6109F5BA50D2D" collectionName="Austria, Wien, Naturhistorisches Museum Wien">NHMW</collectionCode>
, examined.
</paragraph>
<paragraph id="B5495F6C5EEE35AA413AE2FC720D8C05" pageNumber="24">
<taxonomicName id="326B52CAE080A2E55FC98768FFDD8AD4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142288" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole (Pheidole) oceanica var. nigriscapa Santschi" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="oceanica" subGenus="Pheidole" subGenus.completionEvidence="missing, matching species" subGenus.innerRound="1" subGenus.outerRound="1" variety="nigriscapa">Pheidole (Pheidole) oceanica var. nigriscapa Santschi</taxonomicName>
, 1928a: 48. Wilson &amp; Taylor 1967: 45 (junior synonym of
<taxonomicName id="F4EDC715756C72B8CB5148CF768BBD59" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Syntype: 1 minor, &quot;SAMOAN IS. Apia. 3.iii.24. P.A.Buxton. 627.&quot;,
<collectionCode id="8ED897E226178CAD9743F93D9A156EC0" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined; syntype(?): 1 major, &quot;Apia, Samoa oct 1911. (leg. H. Swole)&quot;,
<collectionCode id="1DA313276652AB7F99F0DF0BC1C0ADA9" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined.
</paragraph>
<paragraph id="A2525C6B7DAC24BC9AC9934D0F34C8A1" pageNumber="24">
<taxonomicName id="E12C33A0DC8B1B5AE6C68666D8E52013" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:232306" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole oceanica subsp. nigriscapa var. tahitiana Cheesman &amp; Crawley" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="oceanica" subGenus.completionEvidence="missing, matching species" subGenus.innerRound="1" subGenus.outerRound="1" subSpecies="nigriscapa" variety="tahitiana">Pheidole oceanica subsp. nigriscapa var. tahitiana Santschi</taxonomicName>
, 1928b: 516 (unavailable name). Wilson &amp; Taylor 1967: 45 (material referable to
<taxonomicName id="D5FF74B83B2AC23FF7D11FD8C4CE92AE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="867" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">P. fervens</taxonomicName>
). Material referable to this form: minor, Tahiti, not examined.
</paragraph>
<paragraph id="0A00E4F39F5A93C21C50AE49508C0936" pageNumber="24">
<taxonomicName id="583722D6A624A3154BBD9CB6C3BAE6D5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33581" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole amia Forel" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="amia">Pheidole amia Forel</taxonomicName>
, 1912a: 60-61. Bolton 1995: 316 (catalogue), Eguchi 2004: 197 (lectotype designation, junior synonym of
<taxonomicName id="ED87CDDCAE356CB59552A1DDC9B762D8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Lectotype: major, &quot;Takao Formosa Sauter XI 7-0&quot; [= Kaohsiung, Taiwan],
<collectionCode id="91423ADCC029CAB8D5D1DEE02FAC10E2" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined; paralectotypes: 3 minors, &quot;Takao Formosa (Sauter) 26&quot;,
<collectionCode id="83564A21519DF223A152221183F41C94" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined.
</paragraph>
<paragraph id="7D0E343440249E268E33C2FE93D1FDA8" pageNumber="24">
<taxonomicName id="A1656260A0FDCE564E12CCA70476A981" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142280" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana var. desucta Wheeler" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="javana" variety="desucta">Pheidole javana var. desucta Wheeler</taxonomicName>
, 1929a: 2. Eguchi 2001b: 53 (lectotype designation, junior synonym offervens). Lectotype: major, &quot;Back Liang, China&quot;,
<collectionCode id="69EEA7E342B3BCD041E06AF08468872D" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
co-type 20659, examined; paralectotypes: 1 major, 2 minors &amp; 3 queens, same data as lectotype,
<collectionCode id="DD4E70D9FD7DF15D7608D03D522EAADD" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
co-type 20659, examined.
</paragraph>
<paragraph id="9C4FBBD52F81967F33A80853155C1DD2" pageNumber="24">
<taxonomicName id="9282F33701E1757693DE6D5746E837BA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142281" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana var. dharmsalana Forel" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="javana" variety="dharmsalana">Pheidole javana var. dharmsalana Forel</taxonomicName>
, 1902: 184, 198. Eguchi 2004: 198 (lectotype designation, junior synonym of
<taxonomicName id="5777FB2853A23E3EC229B413A5DE4E82" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Lectotype: major, &quot;Dharmsala&quot; [India],
<collectionCode id="8D88095B67630077DAE4C468756041A1" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined; paralectotypes: 2 minors, same data as lectotype,
<collectionCode id="CB3A1B73B0EBFA4A92C7D2EF80FFA5D2" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined.
</paragraph>
<paragraph id="DC3121265F89223A148F70DD24629643" pageNumber="24">
<taxonomicName id="66B82F11824B10F3C1A68706D34A7038" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142282" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana var. dolenda Forel" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="javana" variety="dolenda">Pheidole javana var. dolenda Forel</taxonomicName>
, 1912a: 60. Eguchi 2004: 198 (lectotype designation, junior synonym of
<taxonomicName id="42D94F40E3033003C2F27128DEE0FE20" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Lectotype: major, &quot;Akau, Formosa&quot;,
<collectionCode id="1013C46CF528528D96A6AE5E755FD728" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined; paralectotypes: 1 major &amp; 3 minors, same data as lectotype,
<collectionCode id="D699D4E775841CEF0838DBC9190DF831" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
, examined.
</paragraph>
<paragraph id="C12D724797B9E0593F5CEE720B8CDB48" pageNumber="24">
<taxonomicName id="3C8A52B6DE9BCF5CD31EE1586A789290" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142285" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana st. soror Santschi" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="javana" variety="soror">Pheidole javana var. soror Santschi</taxonomicName>
, 1937b: 369. Eguchi 2004: 198 (lectotype designation, junior synonym offervens). Lectotype: major, &quot;Hokuto Formose K. Sato 731&quot; [= Taiwan],
<collectionCode id="09625106FC5BB06B0E0A4CB9FA65AB96" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined; paralectotypes: 4 minors, same data as lectotype,
<collectionCode id="0DF6286E578C7351D06C9B8DDD23E027" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined.
</paragraph>
<paragraph id="F43DE0FC11F24E98BD9DD431B062D991" pageNumber="24">
<taxonomicName id="E19A81C3AE2F49EBA170294ED1A08B58" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142457" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole nodus st. azumai Santschi" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="subSpecies" species="nodus" subSpecies="azumai">Pheidole nodus st. azumai Santschi</taxonomicName>
, 1941: 274. Eguchi 2004: 198 (lectotype designation, junior synonym of
<taxonomicName id="D536DDB0B13E5A603AA4C7EAC476EA64" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Lectotype: major, &quot;Japon. Osaka Tennooji 26II39 M. Azuma.&quot; [= Tennooji, Osaka, Japan],
<collectionCode id="79DB9023DCC475C5D77997B6F82C05A2" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined; paralectotypes: 2 minors, same data as holotype,
<collectionCode id="786E7E2BDF3641D4714BE28C69ADC512" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
, examined.
</paragraph>
<paragraph id="97FF718496D04463344E1820DEC5D453" pageNumber="24">
<taxonomicName id="3864A23F4464B984A05422E8B7BE7D9C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142526" class="Insecta" family="Formicidae" genus="Solenopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Solenopsis pungens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="pungens">Solenopsis pungens F. Smith</taxonomicName>
, 1861: 48-49. Donisthorpe 1932: 469 (combination in
<taxonomicName id="F46D44284AF7749CAFAB033394467A2C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24893" class="Insecta" family="Formicidae" genus="Pheidologeton" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidologeton Mayr" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="genus">Pheidologeton</taxonomicName>
), Bolton1995: 328 (combination in
<taxonomicName id="78B06CD4593C7169B112A57B48AFFD2E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
), Eguchi 2004: 198 (lectotype designation, junior synonym of
<taxonomicName id="DBE1E76D19D6FA03C1BA28E6E0827E0F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
). Lectotype: major, &quot;Menado&quot; [= Menado Sulawesi, Indonesia],
<collectionCode id="05C3CB2EC77A86CB14640AFCAB66074A" collectionName="United Kingdom, Oxford, University Museum of Natural History">OXUM</collectionCode>
, examined; paralectotypes: 1 major, same data as lectotype,
<collectionCode id="F1AAE99441DCBD52728CED91915C0EA7" collectionName="United Kingdom, Oxford, University Museum of Natural History">OXUM</collectionCode>
, examined; 1 minor, &quot;Men. 18.&quot; [= Menado Sulawesi, Indonesia],
<collectionCode id="D997693E80763331A4ADEBE42B4B1BEE" collectionName="United Kingdom, Oxford, University Museum of Natural History">OXUM</collectionCode>
, examined.
</paragraph>
<paragraph id="96E0228B8023D9D5AE95B3F35CE4C310" pageNumber="24">
Type material of
<taxonomicName id="C57292E2088638440CCFD8FC6CDBB6C0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142283" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="680" genus.bestMatchVote="2" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole javana var. pectinata Stitz" order="Hymenoptera" pageNumber="24" phylum="Arthropoda" rank="variety" species="javana" variety="pectinata">P. javana var. pectinata Stitz</taxonomicName>
, 1912: 504 (type locality: Seram I., Indonesia) has not yet been examined by me, and its status is still unsettled in the present study.
</paragraph>
</subSubSection>
<subSubSection id="76A37BB7E64FD7CF6023143DD86EF885" type="materials_examined">
<paragraph id="809B8239775088BF342568E05F466B3A" lastPageNumber="25" pageNumber="24">
Other material examined: Mainland Japan: Kagoshima: Toso, Kagoshima City [T. Akiyama's colony: 021105-1]. Ryukyus: Okinoerabu-jima I.: Azufu, [Litter shift-3 (A. Shimono)]; Iriomote-jima I. [K. Eguchi's colony 96-JPN-001, -003]. Taiwan: Nantou: Hori [S. Kubota's sample: 81-A-T16]; Kaohsiung: Meishankou [S. Kubota's sample: 80-E-K6]. S. China: Macau: Taipa I. [Eg99-MAC-01, -02], Hac-Sa, Coloane I. [K. Eguchi]; Hong Kong: Victoria Park, Hong Kong I. [K. Eguchi]. Vietnam: Thai Nguyen: My Yen Commune Forest (edge of secondary forest),
<geoCoordinate id="B75064BFA56263EDCDDC601C04AB4D09" direction="north" orientation="latitude" value="21.583334">21°35'N</geoCoordinate>
,
<geoCoordinate id="9C159AE5AD28AE572F7CD28F03095DD1" direction="east" orientation="longitude" value="105.6">105°36'E</geoCoordinate>
, Na Hau Village [Eg01-VN-143, -149, -150]; Ha Noi: urban
<pageStartToken id="50BE75574D9A03A6BE33F3C3EEC9B70D" pageNumber="25">area</pageStartToken>
[T.V. Bui's colony: BTV04-HN-01, -14, -15, -16]. Thailand: Bangkok: Campus of Kasetsart Univ. [Eg01- TH-583, -584], Bang Khean Distr. (residential area) [TH03-SKY-103]; Chanthaburi: Yay-am Dist. [Eg01-TH- 66B]. E. Malaysia: Sabah: Tawau Hills Park [Eg96-BOR-009, -021]; Sarawak: Niah N.P. [Sk. Yamane]. Indonesia: W. Sumatra: Maninjau, [SNS col.], Sako, nr. Tapan [SNS col.], Sukarami, Padang [M. Kawamura's colony: 9/28a (1999)], Ulu Gadut, nr. Padang [SNS col.]; W. Java: Bogor [M. Kawamura's colony: 153 (1998)]; E. Java: Air Panas Cargar, 1600 m alt., Bumiaji, Batu [JV02/03-SKY-71, -72, -73, -75, -80], Sumberbrantas, 1600 m alt., Tulungrejo, Batu [F. Yamane]. Philippines: Palawan: Pupok, Napsan, in a cottage [H. Fukuda]. Tonga: Eua Is.: Eua Forest Res. [J. Wetterer]. Samoa: Apia [H. Swole,
<collectionCode id="DB5A125B3D6C6CA4370E94F285869A21" collectionName="Switzerland, Basel, Naturhistorisches Museum">NHMB</collectionCode>
]. USA: California: Los Angeles[M.J. Martinez,
<collectionCode id="0779BC06034746ACDFE9D3F5E687BA22" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZC</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="721485298DD8210A217DAB6A319FC615" type="description">
<paragraph id="8CF839F328ACF553F77E190FC950389A" pageNumber="25">Worker measurements &amp; indices: Major (n=5). - HL 1.31-1.56 mm; HW 1.22-1.44 mm; CI 92-97; SL 0.84-0.95 mm; SI 66-70; FL 1.12-1.27 mm; FI 88-94.</paragraph>
<paragraph id="A6D7F942AAE85FA9E3CBCF52B550B212" pageNumber="25">Minor (n=5). - HL 0.65-0.73 mm; HW 0.54-0.60 mm; CI 81-84; SL 0.77-0.87 mm; SI 138-153; FL 0.81-0.93 mm; FI 149-163.</paragraph>
<paragraph id="44DB0BAFFCD1BBFD51B4A711E2586237" pageNumber="25">Worker description</paragraph>
<paragraph id="5ADDCD4751294D3F965546913C4B4624" pageNumber="25">Major. - Head in lateral view not or hardly impressed on vertex; posterior concavity of head in full-face view usually shallow; frons and anterior part of vertex rugose longitudinally; posterior part of vertex and dorsal face of vertexal lobe rugoso-reticulate, with interspaces weakly punctured; frontal carina conspicuous; antennal scrobe inconspicuous or shallowly impressed; clypeus without a median longitudinal carina; median and submedian processes of hypostoma absent or present but inconspicuous; lateral processes conspicuous; antenna with a 3-segmented club; maximal diameter of eye almost as long as or shorter than antennal segment X. Promesonotal dome in dorsal view largely smooth, or sparsely sculptured with weak transverse rugulae, in lateral view with a low to conspicuous mound on its posterior slope; humerus not produced; the dome much narrower at the humeri than at the bottom; mesopleuron, metapleuron and lateral face of propodeum well punctured, sometimes overlain with weak rugulae; propodeal spine narrowly based, usually slightly curved apically. Petiole longer than postpetiole (excluding helcium); postpetiole not massive. First gastral tergite smooth and shining entirely, or sometimes shagreened around its articulation with postpetiole.</paragraph>
<paragraph id="2F22BEFFAF61D381FB02B2AADFB43FD7" pageNumber="25">Minor. - Frons and vertex smooth and shining, or rarely shagreened or very weakly rugoso-punctate; area between antennal insertion and eye weakly rugoso-punctate; preoccipital carina conspicuous dorsally and laterally; median part of clypeus smooth, without a median longitudinal carina; antenna with a 3-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye much shorter than antennal segment X. Promesonotal dome largely smooth and shining, but sometimes weakly rugoso-punctate dorsolaterally, in lateral view with a low or inconspicuous mound on its gentle posterior slope; humerus of the dome in dorsal-oblique view not produced; mesopleuron, metapleuron and lateral face of propodeum well punctured; propodeal spine elongate-triangular, directing upward. Petiole (a little) longer than postpetiole (excluding helcium); postpetiole not massive.</paragraph>
</subSubSection>
<subSubSection id="F6BC14D3AB9B36167ECCBF22DF8F4C23" type="discussion">
<paragraph id="776A55A4698526983C1393415C725C18" pageNumber="25">
Recognition: This medium-sized species with general habitus is similar to
<taxonomicName id="3C23E1C14705E1EF9F31B79AEA0096DE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33623" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole binghamii Forel" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="binghamii">Pheidole binghamii Forel</taxonomicName>
,
<taxonomicName id="29237527CBBB7444DB3B61FF7FB70906" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244652" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="5538" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole elongicephala Eguchi" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="elongicephala">P. elongicephala</taxonomicName>
<taxonomicNameLabel id="7C95FA79BA5424EA49CEF4B6720A0649" rank="species">sp.n.</taxonomicNameLabel>
,
<taxonomicName id="88CE43A460B4EF3DA2F7E5D830D7C2C5" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33798" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="9196" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole indica Mayr" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="indica">P. indica Mayr</taxonomicName>
,
<taxonomicName id="B51F7D3C83F12D723A2E83E6C82FE8A2" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244655" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="11935" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole ochracea Eguchi" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="ochracea">P. ochracea</taxonomicName>
<taxonomicNameLabel id="923671908528DA5BAD614C96606F2F39" rank="species">sp.n.</taxonomicNameLabel>
and
<taxonomicName id="0F16F0699C1D0F23B31ACAD23AAC38A0" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="11926" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="plagiaria">P. plagiaria F. Smith</taxonomicName>
among Indo-Chinese species.
<taxonomicName id="1BA7F08744B7A1FCF4B030E5E67D3AB3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="163" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="fervens">P. fervens</taxonomicName>
is well separated from
<taxonomicName id="45CF3FCA3C4B0ABA219323E8AE0C3896" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33798" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="9226" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole indica Mayr" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="indica">P. indica</taxonomicName>
which has the following characteristics: eye relatively large (maximal diameter of eye much longer than antennal segment X in the major, and as long as or a little longer than X in the minor); in the major propodeal spine is relatively broadly based (see Eguchi 2004).
</paragraph>
<paragraph id="1E2CD714ABBDF271A8AB6D597580D198" pageNumber="25">
Posterior margin of head in full-face view is more deeply and narrowly concave in the major of
<taxonomicName id="715826DD5B7812F9C4C987E78AC3EAA4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33623" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole binghamii Forel" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="binghamii">P. binghamii,</taxonomicName>
<taxonomicName id="A2C7F17B0E314B6BC92B3C79228BB70D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244652" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="348" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole elongicephala Eguchi" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="elongicephala">P. elongicephala</taxonomicName>
and
<taxonomicName id="B63588205F880E95DB18C4637FCAE5DF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244655" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="12042" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole ochracea Eguchi" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="ochracea">P. ochracea</taxonomicName>
than in that of
<taxonomicName id="D4AB7F85C37A905E858D25CD0263F89D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="67" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="fervens">P. fervens</taxonomicName>
, and sculpture on dorsum of vertexal lobe is usually stronger in the major of the former three.
</paragraph>
<paragraph id="5A18315783A0571ED3FFBF569A390877" pageNumber="25">
The major of P
<taxonomicName id="B8789F47D74D15EFE13237E35B977EEC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="fervens">fervens</taxonomicName>
is separated from that of
<taxonomicName id="BD9A8CDB737BF4EB071F47A631E086E8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33933" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="12064" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole plagiaria Smith" order="Hymenoptera" pageNumber="25" phylum="Arthropoda" rank="species" species="plagiaria">P. plagiaria</taxonomicName>
which has the following characteristics: posterior margin of head in full-face view deeply concave; the rugulae running almost transversely along posterior margin of vertexal lobe.
</paragraph>
</subSubSection>
<subSubSection id="0870E4ED2F86AC611CF60F5AED9E3B56" type="multiple">
<caption id="BAF8BFDF9BCA1801702EF2B02AE0D81D">
<paragraph id="B5B79F86A70226F765AC2AF0CCEB72A0" pageNumber="26">
<pageStartToken id="E9CBB5C0AF8B4DFFEFF986D20C45FEA0" pageNumber="26">FIGURE</pageStartToken>
5a-d.
<taxonomicName id="8E8A4ACB9D75E06747950C87540CA4A1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="26" phylum="Arthropoda" rank="species" species="fervens">Pheidole fervens</taxonomicName>
, nontype major [BTV04-HN-14] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
</paragraph>
</caption>
<caption id="47C19847E264F50A925672F56BA4AEBE">
<paragraph id="8979ADED7C4CBC3D66DEA730A9C090EB" pageNumber="27">
<pageStartToken id="C65A34FC57FC6BB25924D3172DB1D1B7" pageNumber="27">FIGURE</pageStartToken>
5e-g,
<taxonomicName id="35A12759B0802549FD159A375C16D875" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33740" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole fervens Smith" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="fervens">Pheidole fervens</taxonomicName>
, nontype minor [BTV04-HN-14] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3DA0CF6648567DDF0C4281D41BD9E5E3" type="distribution">
<paragraph id="CFE5A588997489079BA9B92E9082BC43" pageNumber="27">
Distribution &amp; bionomics: Known from N. Vietnam, Manchurian subregion (southern part only), Oriental region, Austro-Malayan subregion, W. Pacific and West coast of N. America. In the Indo-Malayan subregion this species is one of the prevailing
<taxonomicName id="43E1E06159298EBD8E8BB6F34EDEBBE1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
species in semiurban and rural areas.
</paragraph>
</subSubSection>
</treatment>
</document>