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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.76790" ID-Pensoft-Pub="1314-2003-205-99" ID-Pensoft-UUID="4024A478048757B197E5CF8734331A9E" ModsDocID="1314-2003-205-99" checkinTime="1661230211668" checkinUser="pensoft" docAuthor="O'Donnell, Shawn A., Ringelberg, Jens J. &amp; Lewis, Gwilym P." docDate="2022" docId="6E465CCF9DEF53CBA3D5F320A3C4D078" docLanguage="en" docName="PhytoKeys 205: 99-145" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.76790" docTitle="Entada chrysostachys (Benth.) Drake, Hist. Phys. Madagascar 30: 51. 1902." docType="treatment" docVersion="1" id="4024A478048757B197E5CF8734331A9E" lastPageNumber="99" masterDocId="4024A478048757B197E5CF8734331A9E" masterDocTitle="Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)" masterLastPageNumber="145" masterPageNumber="99" pageNumber="99" updateTime="1661230211668" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Re-circumscription of the mimosoid genus Entada including new combinations for all species of the phylogenetically nested Elephantorrhiza (Leguminosae, Caesalpinioideae, mimosoid clade)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>O'Donnell, Shawn A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0731-7425</mods:nameIdentifier>
<mods:affiliation>Department of Geography and Environmental Sciences, Northumbria University, Newcastle upon Tyne, NE 1 8 ST, UK</mods:affiliation>
<mods:nameIdentifier type="email">shawn.odonnell@cantab.net</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ringelberg, Jens J.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0567-5210</mods:nameIdentifier>
<mods:affiliation>Department of Systematic and Evolutionary Botany, University of Zurich, 8008 Zurich, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lewis, Gwilym P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2599-4577</mods:nameIdentifier>
<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, TW 9 3 AE, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>205</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>99</mods:start>
<mods:end>145</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.205.76790</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.76790</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-99</mods:identifier>
<mods:identifier type="Pensoft-UUID">4024A478048757B197E5CF8734331A9E</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:6E465CCF9DEF53CBA3D5F320A3C4D078" httpUri="http://treatment.plazi.org/id/6E465CCF9DEF53CBA3D5F320A3C4D078" lastPageNumber="99" pageId="0" pageNumber="99">
<subSubSection pageId="0" pageNumber="99" type="nomenclature">
<paragraph pageId="0" pageNumber="99">
<taxonomicName LSID="6E465CCF-9DEF-53CB-A3D5-F320A3C4D078" authority="(Benth.) Drake, Hist. Phys. Madagascar 30: 51. 1902." authorityName="(Benth.) Drake, Hist. Phys. Madagascar 30: 51. 1902." class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada chrysostachys" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="chrysostachys">Entada chrysostachys (Benth.) Drake, Hist. Phys. Madagascar 30: 51. 1902.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="reference_group">
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada kirkii" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="kirkii">= Entada kirkii</taxonomicName>
Oliv., Fl. Trop. Afr. 2: 327. 1871.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada boiviniana" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="boiviniana">= Entada boiviniana</taxonomicName>
(Baill.) Drake, A. Grandidier, Hist. Phys. Madagascar 30: 51. 1902. (publ. 1903).
</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada grandidieri" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="grandidieri">= Entada grandidieri</taxonomicName>
(Baill.) Drake, A. Grandidier, Hist. Phys. Madagascar 30: 51. 1902. (publ. 1903).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="materials_examined">
<paragraph pageId="0" pageNumber="99">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="99">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Imamou, W" country="MADAGASCAR" location="Emirna Province" specimenCount="1" typeStatus="holotype">
<collectingCountry name="Madagascar">MADAGASCAR</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:6E465CCF9DEF53CBA3D5F320A3C4D078:8B0AA66C15907318C95E4FC1A671B416" country="MADAGASCAR" name="Emirna Province">Emirna Province</location>
[Imerina] and
<collectorName>
Imamou,
<emphasis italics="true" pageId="0" pageNumber="99">W. Bojer s.n.</emphasis>
</collectorName>
(
<typeStatus>holotype</typeStatus>
: K; isotypes: M [M0218663], P [P00367635 &amp; P00367637])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="basionym">
<paragraph pageId="0" pageNumber="99">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="99">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Adenanthera" kingdom="Plantae" lsidName="Adenanthera chrysostachys" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="chrysostachys">
<emphasis italics="true" pageId="0" pageNumber="99">Adenanthera chrysostachys</emphasis>
</taxonomicName>
Benth., J. Bot. (Hooker) 4: 343. 1841.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="description">
<paragraph pageId="0" pageNumber="99">Description.</paragraph>
<paragraph pageId="0" pageNumber="99">
Shrub or small tree to 10 m tall or liana to 12 m, stem to 20 cm thick, often twisted (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Entada chrysostachys habit, vegetative and reproductive structures. A small tree, Madagascar (photo: thierrycordenos CC BY-NC 4.0, Ueda (2021) observation 25108890) B leaf, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11) C axillary spiciform racemes of open, pedicellate flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and immature pods, Madagascar (photo: D Du Puy) E mature pods, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11)." figureDoi="10.3897/phytokeys.205.76790.figure10" httpUri="https://binary.pensoft.net/fig/732705" pageId="0" pageNumber="99">10A</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Leaves</emphasis>
: rachis 8-16 cm long, grooved above, glabrous; pinnae (2-)3-5(-8) pairs per leaf, (4.5-)5.2-8.9(-13) cm long, with 10-17(-21) pairs of leaflets; leaflets 13-19(-29)
<normalizedToken originalValue="×">x</normalizedToken>
(3-)4.9-5.5(-10) mm, oblong to obovate-oblong, apex rounded, base asymmetric rounded on proximal margin and cuneate on distal margin, mid-rib diagonal and raised above and below, lamina appressed-pubescent to glabrous (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Entada chrysostachys habit, vegetative and reproductive structures. A small tree, Madagascar (photo: thierrycordenos CC BY-NC 4.0, Ueda (2021) observation 25108890) B leaf, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11) C axillary spiciform racemes of open, pedicellate flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and immature pods, Madagascar (photo: D Du Puy) E mature pods, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11)." figureDoi="10.3897/phytokeys.205.76790.figure10" httpUri="https://binary.pensoft.net/fig/732705" pageId="0" pageNumber="99">10B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Inflorescence</emphasis>
: an axillary spiciform raceme, 4-12(-13.5) cm long, usually clustered, but sometimes solitary, rachis pubescent or glabrous (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Entada chrysostachys habit, vegetative and reproductive structures. A small tree, Madagascar (photo: thierrycordenos CC BY-NC 4.0, Ueda (2021) observation 25108890) B leaf, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11) C axillary spiciform racemes of open, pedicellate flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and immature pods, Madagascar (photo: D Du Puy) E mature pods, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11)." figureDoi="10.3897/phytokeys.205.76790.figure10" httpUri="https://binary.pensoft.net/fig/732705" pageId="0" pageNumber="99">10C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Flowers</emphasis>
: white to yellow, pedicels 1-1.5 mm long, with an unpleasant odour; calyx obconical, 1-1.5 mm long, glabrous to sparsely pubescent, distinctly toothed; petals 3-4
<normalizedToken originalValue="×">x</normalizedToken>
1-1.4 mm; stamen filaments 4-6 mm long.
<emphasis bold="true" pageId="0" pageNumber="99">Fruit</emphasis>
: a torulose, laterally compressed, slightly curved craspedium 20-45
<normalizedToken originalValue="×">x</normalizedToken>
5-10 cm, with transverse septa between seeds dividing the fruit into one-seeded segments which, upon ripening, fall from the persistent replum (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Entada chrysostachys habit, vegetative and reproductive structures. A small tree, Madagascar (photo: thierrycordenos CC BY-NC 4.0, Ueda (2021) observation 25108890) B leaf, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11) C axillary spiciform racemes of open, pedicellate flowers and closed flower buds, Madagascar (photo: D Du Puy) D leaves and immature pods, Madagascar (photo: D Du Puy) E mature pods, Mozambique (photo: O Maurin CC BY-NC-SA 3.0, iBOL (2016) record SAFH 1507 - 11)." figureDoi="10.3897/phytokeys.205.76790.figure10" httpUri="https://binary.pensoft.net/fig/732705" pageId="0" pageNumber="99">10D, E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="99">Seeds</emphasis>
: elliptic, 14-17.7
<normalizedToken originalValue="×">x</normalizedToken>
10.5-12.8
<normalizedToken originalValue="×">x</normalizedToken>
3.3-4 mm, pleurogram elliptic, open near hilum.
</paragraph>
<caption doi="10.3897/phytokeys.205.76790.figure10" httpUri="https://binary.pensoft.net/fig/732705" pageId="0" pageNumber="99" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="99">
<emphasis bold="true" pageId="0" pageNumber="99">Figure 10.</emphasis>
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Entada" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Entada chrysostachys" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="chrysostachys">
<emphasis italics="true" pageId="0" pageNumber="99">Entada chrysostachys</emphasis>
</taxonomicName>
habit, vegetative and reproductive structures.
<emphasis bold="true" pageId="0" pageNumber="99">A</emphasis>
small tree, Madagascar (photo: thierrycordenos
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC 4.0</emphasis>
,
<bibRefCitation author="Ueda, K" journalOrPublisher="Acta Phytotaxonomica et Geobotanica" pageId="0" pageNumber="99" publicationUrl="iNaturalist.org" refId="B59" refString="Ueda, K, 2021. iNaturalist Research-grade Observations. iNaturalist.org" title="iNaturalist Research-grade Observations." url="iNaturalist.org" year="2021">Ueda (2021)</bibRefCitation>
observation 25108890)
<emphasis bold="true" pageId="0" pageNumber="99">B</emphasis>
leaf, Mozambique (photo: O Maurin
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC-SA 3.0</emphasis>
,
<bibRefCitation DOI="https://doi.org/10.15468/inygc6" author="Hyde, MA" journalOrPublisher="Institut Francais de Pondichery, Pondichery" pageId="0" pageNumber="99" refId="B26" refString="2016. . https://doi.org/10.15468/inygc6" url="https://doi.org/10.15468/inygc6" year="2016">iBOL (2016)</bibRefCitation>
record SAFH1507-11)
<emphasis bold="true" pageId="0" pageNumber="99">C</emphasis>
axillary spiciform racemes of open, pedicellate flowers and closed flower buds, Madagascar (photo: D Du Puy)
<emphasis bold="true" pageId="0" pageNumber="99">D</emphasis>
leaves and immature pods, Madagascar (photo: D Du Puy)
<emphasis bold="true" pageId="0" pageNumber="99">E</emphasis>
mature pods, Mozambique (photo: O Maurin
<emphasis italics="true" pageId="0" pageNumber="99">CC BY-NC-SA 3.0</emphasis>
,
<bibRefCitation DOI="https://doi.org/10.15468/inygc6" author="Hyde, MA" journalOrPublisher="Institut Francais de Pondichery, Pondichery" pageId="0" pageNumber="99" refId="B26" refString="2016. . https://doi.org/10.15468/inygc6" url="https://doi.org/10.15468/inygc6" year="2016">iBOL (2016)</bibRefCitation>
record SAFH1507-11).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="distribution">
<paragraph pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="99">Madagascar, Comoro Islands, Mozambique, Zimbabwe, Zambia, Malawi, Tanzania.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="99" type="habitat">
<paragraph pageId="0" pageNumber="99">Habitat and ecology.</paragraph>
<paragraph pageId="0" pageNumber="99">
Disturbed forests and grassland; riparian thicket; woodland characterised by
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" lsidName="Brachystegia glaucescens" order="Fabales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="glaucescens">
<emphasis italics="true" pageId="0" pageNumber="99">Brachystegia glaucescens</emphasis>
</taxonomicName>
Hutch. &amp; Burtt Davy; and seasonally wet valley bottoms with
<taxonomicName class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" lsidName="Combretum" order="Myrtales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="99">Combretum</emphasis>
</taxonomicName>
; sandy soils and laterite.
</paragraph>
</subSubSection>
</treatment>
</document>